Humulus, hop, is a small genus of flowering plants in the family Cannabaceae. The hop is native to temperate regions of the Northern Hemisphere. Hops are the female flowers of the hop species. Although referred to in American literature as the hops "vine", it is technically a bine. In British literature the term “vine” is reserved for the grape genus Vitis. Humulus is described as a twining perennial herbaceous plant which sends up new shoots in early spring and dies back to the cold-hardy rhizome in autumn. Hop shoots grow rapidly, at the peak of growth can grow 20 to 50 centimetres per week. Hop bines climb by wrapping clockwise around anything within reach, individual bines grow between 2 to 15 metres depending on what is available to grow on; the leaves are opposite, with a 7 to 12 cm leafstalk and a heart-shaped, fan-lobed blade 12 to 25 cm long and broad. When the hop bines run out of material to climb, horizontal shoots sprout between the leaves of the main stem to form a network of stems wound round each other.
Male and female flowers of the hop plant develop on separate plants. Female plants, which produce the hop flowers used in brewing beer, are propagated vegetatively and grown in the absence of male plants; this prevents pollination and the development of viable seeds, which are sometimes considered undesirable for brewing beer owing to the potential for off-flavors arising from the introduction of fatty acids from the seeds. There are three species, one with five varieties: Humulus japonicus. Asian hop. Leaves with 5–7 lobes. Eastern Asia. Humulus yunnanensis Asian hop from China. Humulus lupulus. Common hop. Leaves with 3–5 lobes. Europe, western Asia, North America. Humulus lupulus var. lupulus. Europe, western Asia. Humulus lupulus var. cordifolius. Eastern Asia. Humulus lupulus var. lupuloides. Eastern North America. Humulus lupulus var. neomexicanus. Western North America. Humulus lupulus var. pubescens. Midwest and eastern North America. Brewers' hops are specific cultivars, propagated by asexual reproduction, see the article, "List of hop varieties".
Hops sometimes added post-ferment. In pharmacy lupulus is the designation of hop; the dried catkins referred to as hop cones, of the female plant of H. lupulus are used to prepare infusion of hop, tincture of hop, extract of hop. The characteristic bitterness imparted by the addition of hops to the brewing process is due to the presence of the bitter acids, which are prenylated acylphloroglucinol derivatives. Bitter acids are divided into the alpha-acids, with humulone the major compound, the beta-acids, with lupulone the major compound; these hop acids are vinylogous acids, with acidic ring enols in conjugation with ring and substituent carbonyl groups. Plants in the genus Humulus produce terpenophenolic metabolites. Hops contain xanthohumol, a prenylated chalcone, other compounds under preliminary research for their potential health properties. Jeanine S. DeNoma: Humulus Genetic Resources Hops varieties research Plants for a Future: Humulus lupulus
Wikispecies is a wiki-based online project supported by the Wikimedia Foundation. Its aim is to create a comprehensive free content catalogue of all species. Jimmy Wales stated that editors are not required to fax in their degrees, but that submissions will have to pass muster with a technical audience. Wikispecies is available under the GNU Free Documentation License and CC BY-SA 3.0. Started in September 2004, with biologists across the world invited to contribute, the project had grown a framework encompassing the Linnaean taxonomy with links to Wikipedia articles on individual species by April 2005. Benedikt Mandl co-ordinated the efforts of several people who are interested in getting involved with the project and contacted potential supporters in early summer 2004. Databases were evaluated and the administrators contacted, some of them have agreed on providing their data for Wikispecies. Mandl defined two major tasks: Figure out how the contents of the data base would need to be presented—by asking experts, potential non-professional users and comparing that with existing databases Figure out how to do the software, which hardware is required and how to cover the costs—by asking experts, looking for fellow volunteers and potential sponsorsAdvantages and disadvantages were discussed by the wikimedia-I mailing list.
The board of directors of the Wikimedia Foundation voted by 4 to 0 in favor of the establishment of a Wikispecies. The project is hosted at species.wikimedia.org. It was merged to a sister project of Wikimedia Foundation on September 14, 2004. On October 10, 2006, the project exceeded 75,000 articles. On May 20, 2007, the project exceeded 100,000 articles with a total of 5,495 registered users. On September 8, 2008, the project exceeded 150,000 articles with a total of 9,224 registered users. On October 23, 2011, the project reached 300,000 articles. On June 16, 2014, the project reached 400,000 articles. On January 7, 2017, the project reached 500,000 articles. On October 30, 2018, the project reached 600,000 articles, a total of 1.12 million pages. Wikispecies comprises taxon pages, additionally pages about synonyms, taxon authorities, taxonomical publications, institutions or repositories holding type specimen. Wikispecies asks users to use images from Wikimedia Commons. Wikispecies does not allow the use of content.
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Ferdinand von Mueller
Baron Sir Ferdinand Jacob Heinrich von Mueller, was a German-Australian physician and most notably, a botanist. He was appointed government botanist for the colony of Victoria by Governor Charles La Trobe in 1853, director of the Royal Botanic Gardens, Melbourne, he founded the National Herbarium of Victoria. He named many Australian plants. Mueller was born in the Grand Duchy of Mecklenburg-Schwerin. After the early death of his parents and Louisa, his grandparents gave him a good education in Tönning, Schleswig. Apprenticed to a chemist at the age of 15, he passed his pharmaceutical examinations and studied botany under Professor Ernst Ferdinand Nolte at Kiel University. In 1847, he received his degree of Doctor of Philosophy from Kiel for a thesis on the plants of the southern regions of Schleswig. Mueller's sister Bertha had been advised to seek a warmer climate for her health, the great botanist Ludwig Preiss, who had returned from Perth, recommended Australia, so in 1847, Mueller and his two surviving sisters sailed from Bremen.
While still on the ship, he fished his first plants out of the water to analyse them. He arrived at Adelaide on 18 December 1847 and found employment as a chemist with Moritz J. Heuzenroeder, in Rundle Street, he was an inveterate explorer, walking alone to Mount Brown during his first year. Shortly afterwards, he obtained 20 acres of land not far from Adelaide in the Bugle Ranges, had a cottage built there, he moved there with his sister Clara, intending to start a farm, but after a few months, he returned to his former employment. Mueller thought to open a chemist's shop in the gold diggings, so in 1851, he moved to Melbourne, capital of the new colony of Victoria, he had contributed a few papers on botanical subjects to German periodicals, in 1852, sent a paper to the Linnean Society of London on "The Flora of South Australia", thus beginning to be well known in botanical circles. Mueller was appointed government botanist for Victoria by Governor Charles La Trobe in 1853, a post, newly created for him.
He examined its flora the Alpine vegetation of Australia, unknown. He explored the Buffalo Ranges went to the upper reaches of the Goulburn River and across Gippsland to the coast; the neighbourhoods of Port Albert and Wilsons Promontory were explored, the journey of some 1,500 miles was completed along the coast to Melbourne. In the same year, he established the National Herbarium of Victoria, which can still be visited today, it has many plants from Australia and abroad. His large private library was transferred to the government of Victoria in 1865 and is incorporated into the library of the herbarium in Melbourne; as a phytographic naturalist, he joined the expedition sent out under Augustus Gregory by the Duke of Newcastle, Secretary of State for the colonies. He explored the Victoria River and other portions of North Australia, was one of the four who reached Termination Lake in 1856, accompanied Gregory's expedition overland to Moreton Bay. Mueller, for his part, found nearly 800 species in Australia new to science.
He published in this year his Definitions of Hitherto Undescribed Australian Plants. From 1854 to 1872, Mueller was a member of the Victorian Institute for the Advancement of Science, which became the Philosophical Institute of Victoria, he was president of the Philosophical Institute in 1859 when it received a royal charter and became the Royal Society of Victoria. He was an active member of the society's "Exploration Committee" which established the Burke and Wills expedition of 1860. Mueller promoted the exploration of Australia, as one of only two members of the Exploration Committee with any experience of exploration, he made several speeches to the society on the topic, he did not favour the selection of Burke as leader, but due to factionalism in the committee, he had little say in the establishment, provisioning, or composition of the exploration party. From 1857 to 1873, he was director of the Royal Botanic Gardens and not only introduced many plants into Victoria, but made the excellent qualities of the blue gum known all over the world, succeeded in introducing it into the south of Europe and South Africa and the extratropical portions of South America.
Mueller was decorated by many foreign countries, including Germany, Spain and Portugal. He was appointed a fellow of the Royal Society in 1861, knighted as Knight Commander of the Order of St Michael and St George in 1879. A list of his'Orders, offices and sundry honours' has been assembled. Many of his decorations were received in return for supplying zoological specimens to royal museums, he was the benefactor of the discoverer of Lake Amadeus and Kata Tjuta. Giles had wanted to name these Lake Mueller and Mt Ferdinand, but Mueller prevailed upon Giles to name them Lake Amadeus, after King Amadeus of Spain, Mt Olga, after Queen Olga of Württemberg. In 1871, King Karl of Württemberg gave him the hereditary title of Freiherr, to mark his distinction in'natural sciences and in particular for the natural history collections and institutions of Our Kingdom' He was known as Baron Sir Ferdinand von Mueller. By 1873, influential Melburnians were critical of Mueller's scientific and educational approach with the Royal Botanic Gardens.
Development of the gardens with an eye to aesthetics was sought. Mueller was dismissed from his position as director of the Botanic Gardens on 31 May 1873. He
Western Australia is a state occupying the entire western third of Australia. It is bounded by the Indian Ocean to the north and west, the Southern Ocean to the south, the Northern Territory to the north-east, South Australia to the south-east. Western Australia is Australia's largest state, with a total land area of 2,529,875 square kilometres, the second-largest country subdivision in the world, surpassed only by Russia's Sakha Republic; the state has about 2.6 million inhabitants – around 11 percent of the national total – of whom the vast majority live in the south-west corner, 79 per cent of the population living in the Perth area, leaving the remainder of the state sparsely populated. The first European visitor to Western Australia was the Dutch explorer Dirk Hartog, who visited the Western Australian coast in 1616; the first European settlement of Western Australia occurred following the landing by Major Edmund Lockyer on 26 December 1826 of an expedition on behalf of the New South Wales colonial government.
He established a convict-supported military garrison at King George III Sound, at present-day Albany, on 21 January 1827 formally took possession of the western third of the continent for the British Crown. This was followed by the establishment of the Swan River Colony in 1829, including the site of the present-day capital, Perth. York was the first inland settlement in Western Australia. Situated 97 kilometres east of Perth, it was settled on 16 September 1831. Western Australia achieved responsible government in 1890 and federated with the other British colonies in Australia in 1901. Today, its economy relies on mining, agriculture and tourism; the state produces 46 per cent of Australia's exports. Western Australia is the second-largest iron ore producer in the world. Western Australia is bounded to the east by longitude 129°E, the meridian 129 degrees east of Greenwich, which defines the border with South Australia and the Northern Territory, bounded by the Indian Ocean to the west and north.
The International Hydrographic Organization designates the body of water south of the continent as part of the Indian Ocean. The total length of the state's eastern border is 1,862 km. There are 20,781 km including 7,892 km of island coastline; the total land area occupied by the state is 2.5 million km2. The bulk of Western Australia consists of the old Yilgarn craton and Pilbara craton which merged with the Deccan Plateau of India and the Karoo and Zimbabwe cratons of Southern Africa, in the Archean Eon to form Ur, one of the oldest supercontinents on Earth. In May 2017, evidence of the earliest known life on land may have been found in 3.48-billion-year-old geyserite and other related mineral deposits uncovered in the Pilbara craton. Because the only mountain-building since has been of the Stirling Range with the rifting from Antarctica, the land is eroded and ancient, with no part of the state above 1,245 metres AHD. Most of the state is a low plateau with an average elevation of about 400 metres low relief, no surface runoff.
This descends sharply to the coastal plains, in some cases forming a sharp escarpment. The extreme age of the landscape has meant that the soils are remarkably infertile and laterised. Soils derived from granitic bedrock contain an order of magnitude less available phosphorus and only half as much nitrogen as soils in comparable climates in other continents. Soils derived from extensive sandplains or ironstone are less fertile, nearly devoid of soluble phosphate and deficient in zinc, copper and sometimes potassium and calcium; the infertility of most of the soils has required heavy application by farmers of fertilizers. These have resulted in damage to bacterial populations; the grazing and use of hoofed mammals and heavy machinery through the years have resulted in compaction of soils and great damage to the fragile soils. Large-scale land clearing for agriculture has damaged habitats for native fauna; as a result, the South West region of the state has a higher concentration of rare, threatened or endangered flora and fauna than many areas of Australia, making it one of the world's biodiversity "hot spots".
Large areas of the state's wheatbelt region have problems with dryland salinity and the loss of fresh water. The southwest coastal area has a Mediterranean climate, it was heavily forested, including large stands of karri, one of the tallest trees in the world. This agricultural region is one of the nine most bio-diverse terrestrial habitats, with a higher proportion of endemic species than most other equivalent regions. Thanks to the offshore Leeuwin Current, the area is one of the top six regions for marine biodiversity and contains the most southerly coral reefs in the world. Average annual rainfall varies from 300 millimetres at the edge of the Wheatbelt region to 1,400 millimetres in the wettest areas near Northcliffe, but from November to March, evaporation exceeds rainfall, it is very dry. Plants are adapted to this as well as the extreme poverty of all soils; the central two-thirds of the state is sparsely inhabited. The only significant economic activity is mining. Annual rainfall averages less than 300 millimetres, most of which occurs in sporadic torrential falls related to cyclone events in summer.
An exception to this is
The eudicots, Eudicotidae or eudicotyledons are a clade of flowering plants, called tricolpates or non-magnoliid dicots by previous authors. The botanical terms were introduced in 1991 by evolutionary botanist James A. Doyle and paleobotanist Carol L. Hotton to emphasize the evolutionary divergence of tricolpate dicots from earlier, less specialized, dicots; the close relationships among flowering plants with tricolpate pollen grains was seen in morphological studies of shared derived characters. These plants have a distinct trait in their pollen grains of exhibiting three colpi or grooves paralleling the polar axis. Molecular evidence confirmed the genetic basis for the evolutionary relationships among flowering plants with tricolpate pollen grains and dicotyledonous traits; the term means "true dicotyledons", as it contains the majority of plants that have been considered dicots and have characteristics of the dicots. The term "eudicots" has subsequently been adopted in botany to refer to one of the two largest clades of angiosperms, monocots being the other.
The remaining angiosperms include magnoliids and what are sometimes referred to as basal angiosperms or paleodicots, but these terms have not been or adopted, as they do not refer to a monophyletic group. The other name for the eudicots is tricolpates, a name which refers to the grooved structure of the pollen. Members of the group have tricolpate pollen; these pollens have three or more pores set in furrows called colpi. In contrast, most of the other seed plants produce monosulcate pollen, with a single pore set in a differently oriented groove called the sulcus; the name "tricolpates" is preferred by some botanists to avoid confusion with the dicots, a nonmonophyletic group. Numerous familiar plants are eudicots, including many common food plants and ornamentals; some common and familiar eudicots include members of the sunflower family such as the common dandelion, the forget-me-not and other members of its family, buttercup and macadamia. Most leafy trees of midlatitudes belong to eudicots, with notable exceptions being magnolias and tulip trees which belong to magnoliids, Ginkgo biloba, not an angiosperm.
The name "eudicots" is used in the APG system, of 1998, APG II system, of 2003, for classification of angiosperms. It is applied to a monophyletic group, which includes most of the dicots. "Tricolpate" is a synonym for the "Eudicot" monophyletic group, the "true dicotyledons". The number of pollen grain furrows or pores helps classify the flowering plants, with eudicots having three colpi, other groups having one sulcus. Pollen apertures are any modification of the wall of the pollen grain; these modifications include thinning and pores, they serve as an exit for the pollen contents and allow shrinking and swelling of the grain caused by changes in moisture content. The elongated apertures/ furrows in the pollen grain are called colpi, along with pores, are a chief criterion for identifying the pollen classes; the eudicots can be divided into two groups: the basal eudicots and the core eudicots. Basal eudicot is an informal name for a paraphyletic group; the core eudicots are a monophyletic group.
A 2010 study suggested the core eudicots can be divided into two clades, Gunnerales and a clade called "Pentapetalae", comprising all the remaining core eudicots. The Pentapetalae can be divided into three clades: Dilleniales superrosids consisting of Saxifragales and rosids superasterids consisting of Santalales, Berberidopsidales and asteridsThis division of the eudicots is shown in the following cladogram: The following is a more detailed breakdown according to APG IV, showing within each clade and orders: clade Eudicots order Ranunculales order Proteales order Trochodendrales order Buxales clade Core eudicots order Gunnerales order Dilleniales clade Superrosids order Saxifragales clade Rosids order Vitales clade Fabids order Fabales order Rosales order Fagales order Cucurbitales order Oxalidales order Malpighiales order Celastrales order Zygophyllales clade Malvids order Geraniales order Myrtales order Crossosomatales order Picramniales order Malvales order Brassicales order Huerteales order Sapindales clade Superasterids order Berberidopsidales order Santalales order Caryophyllales clade Asterids order Cornales order Ericales clade Campanulids order Aquifoliales order Asterales order Escalloniales order Bruniales order Apiales order Dipsacales order Paracryphiales clade Lamiids order Solanales order Lamiales order Vahliales order Gentianales order Boraginales order Garryales order Metteniusales order Icacinales Eudicots at the Encyclopedia of Life Eudicots, Tree of Life Web Project Dicots Plant Life Forms
The Fabaceae or Leguminosae known as the legume, pea, or bean family, are a large and economically important family of flowering plants. It includes trees and perennial or annual herbaceous plants, which are recognized by their fruit and their compound, stipulate leaves. Many legumes have characteristic fruits; the family is distributed, is the third-largest land plant family in terms of number of species, behind only the Orchidaceae and Asteraceae, with about 751 genera and about 19,000 known species. The five largest of the genera are Astragalus, Indigofera and Mimosa, which constitute about a quarter of all legume species; the ca. 19,000 known legume species amount to about 7% of flowering plant species. Fabaceae is the most common family found in tropical rainforests and in dry forests in the Americas and Africa. Recent molecular and morphological evidence supports the fact that the Fabaceae is a single monophyletic family; this conclusion has been supported not only by the degree of interrelation shown by different groups within the family compared with that found among the Leguminosae and their closest relations, but by all the recent phylogenetic studies based on DNA sequences.
These studies confirm that the Fabaceae are a monophyletic group, related to the Polygalaceae and Quillajaceae families and that they belong to the order Fabales. Along with the cereals, some fruits and tropical roots, a number of Leguminosae have been a staple human food for millennia and their use is related to human evolution; the Fabaceae family includes a number of important agricultural and food plants, including Glycine max, Pisum sativum, Cicer arietinum, Medicago sativa, Arachis hypogaea, Ceratonia siliqua, Glycyrrhiza glabra. A number of species are weedy pests in different parts of the world, including: Cytisus scoparius, Robinia pseudoacacia, Ulex europaeus, Pueraria lobata, a number of Lupinus species; the name'Fabaceae' comes from the defunct genus Faba, now included in Vicia. The term "faba" comes from Latin, appears to mean "bean". Leguminosae is an older name still considered valid, refers to the fruit of these plants, which are called legumes. Fabaceae range in habit from giant trees to small annual herbs, with the majority being herbaceous perennials.
Plants have indeterminate inflorescences. The flowers have a short hypanthium and a single carpel with a short gynophore, after fertilization produce fruits that are legumes; the Leguminosae have a wide variety of growth forms, including trees, herbaceous plants, vines or lianas. The herbaceous plants can be annuals, biennials, or perennials, without basal or terminal leaf aggregations. Many Legumes have tendrils, they are epiphytes, or vines. The latter support themselves by means of shoots that twist around a support or through cauline or foliar tendrils. Plants can be mesophytes, or xerophytes; the leaves are alternate and compound. Most they are even- or odd-pinnately compound trifoliate and palmately compound, in the Mimosoideae and the Caesalpinioideae bipinnate, they always have stipules, which can be rather inconspicuous. Leaf margins are entire or serrate. Both the leaves and the leaflets have wrinkled pulvini to permit nastic movements. In some species, leaflets have evolved into tendrils.
Many species have leaves with structures that attract ants that protect the plant from herbivore insects. Extrafloral nectaries are common among the Mimosoideae and the Caesalpinioideae, are found in some Faboideae. In some Acacia, the modified hollow stipules are known as domatia. Many Fabaceae host bacteria in their roots within structures called root nodules; these bacteria, known as rhizobia, have the ability to take nitrogen gas out of the air and convert it to a form of nitrogen, usable to the host plant. This process is called nitrogen fixation; the legume, acting as a host, rhizobia, acting as a provider of usable nitrate, form a symbiotic relationship. The flowers have five fused sepals and five free petals, they are hermaphrodite, have a short hypanthium cup shaped. There are ten stamens and one elongated superior ovary, with a curved style, they are arranged in indeterminate inflorescences. Fabaceae are entomophilous plants, the flowers are showy to attract pollinators. In the Caesalpinioideae, the flowers are zygomorphic, as in Cercis, or nearly symmetrical with five equal petals in Bauhinia.
The upper petal is the innermost one, unlike in the Faboideae. Some species, like some in the genus Senna, have asymmetric flowers, with one of the lower petals larger than the opposing one, the style bent to one side; the calyx, corolla, or stamens can be showy in this group. In the Mimosoideae, the flowers are actinomorphic and arranged in globose inflorescences; the petals are small and the stamens, which can be more than just 10, have long, coloured filaments, which are the showiest part of the flower. All of the flowers in an inflorescence open at once. In the Faboideae, the flowers are zygom
The flowering plants known as angiosperms, Angiospermae or Magnoliophyta, are the most diverse group of land plants, with 64 orders, 416 families 13,164 known genera and c. 369,000 known species. Like gymnosperms, angiosperms are seed-producing plants. However, they are distinguished from gymnosperms by characteristics including flowers, endosperm within the seeds, the production of fruits that contain the seeds. Etymologically, angiosperm means a plant; the term comes from the Greek words sperma. The ancestors of flowering plants diverged from gymnosperms in the Triassic Period, 245 to 202 million years ago, the first flowering plants are known from 160 mya, they diversified extensively during the Early Cretaceous, became widespread by 120 mya, replaced conifers as the dominant trees from 100 to 60 mya. Angiosperms differ from other seed plants in several ways, described in the table below; these distinguishing characteristics taken together have made the angiosperms the most diverse and numerous land plants and the most commercially important group to humans.
Angiosperm stems are made up of seven layers. The amount and complexity of tissue-formation in flowering plants exceeds that of gymnosperms; the vascular bundles of the stem are arranged such that the phloem form concentric rings. In the dicotyledons, the bundles in the young stem are arranged in an open ring, separating a central pith from an outer cortex. In each bundle, separating the xylem and phloem, is a layer of meristem or active formative tissue known as cambium. By the formation of a layer of cambium between the bundles, a complete ring is formed, a regular periodical increase in thickness results from the development of xylem on the inside and phloem on the outside; the soft phloem becomes crushed, but the hard wood persists and forms the bulk of the stem and branches of the woody perennial. Owing to differences in the character of the elements produced at the beginning and end of the season, the wood is marked out in transverse section into concentric rings, one for each season of growth, called annual rings.
Among the monocotyledons, the bundles are more numerous in the young stem and are scattered through the ground tissue. They once formed the stem increases in diameter only in exceptional cases; the characteristic feature of angiosperms is the flower. Flowers show remarkable variation in form and elaboration, provide the most trustworthy external characteristics for establishing relationships among angiosperm species; the function of the flower is to ensure fertilization of the ovule and development of fruit containing seeds. The floral apparatus may arise terminally from the axil of a leaf; as in violets, a flower arises singly in the axil of an ordinary foliage-leaf. More the flower-bearing portion of the plant is distinguished from the foliage-bearing or vegetative portion, forms a more or less elaborate branch-system called an inflorescence. There are two kinds of reproductive cells produced by flowers. Microspores, which will divide to become pollen grains, are the "male" cells and are borne in the stamens.
The "female" cells called megaspores, which will divide to become the egg cell, are contained in the ovule and enclosed in the carpel. The flower may consist only of these parts, as in willow, where each flower comprises only a few stamens or two carpels. Other structures are present and serve to protect the sporophylls and to form an envelope attractive to pollinators; the individual members of these surrounding structures are known as petals. The outer series is green and leaf-like, functions to protect the rest of the flower the bud; the inner series is, in general, white or brightly colored, is more delicate in structure. It functions to attract bird pollinators. Attraction is effected by color and nectar, which may be secreted in some part of the flower; the characteristics that attract pollinators account for the popularity of flowers and flowering plants among humans. While the majority of flowers are perfect or hermaphrodite, flowering plants have developed numerous morphological and physiological mechanisms to reduce or prevent self-fertilization.
Heteromorphic flowers have short carpels and long stamens, or vice versa, so animal pollinators cannot transfer pollen to the pistil. Homomorphic flowers may employ a biochemical mechanism called self-incompatibility to discriminate between self and non-self pollen grains. In other species, the male and female parts are morphologically separated, developing on different flowers; the botanical term "Angiosperm", from the Ancient Greek αγγείον, angeíon and σπέρμα, was coined in the form Angiospermae by Paul Hermann in 1690, as the name of one of his primary divisions of the plant kingdom. This included flowering plants possessing seeds enclosed in capsules, distinguished from his Gymnospermae, or flowering plants with achenial or schizo-carpic fruits, the whole fruit or each of its pieces being here regarded as a seed and naked; the term and its antonym were maintained by Carl Linnaeus with the same sense, but with restricted application, in the names of the orders of his class Didynamia. Its use with any