The Mimosoideae are trees, herbs and shrubs that grow in tropical and subtropical climates. They comprise a clade placed at the subfamily or family level in the flowering plant family Fabaceae. In previous classifications, Mimosoideae refers to what was considered the tribe Mimoseae. Characteristics include flowers in radial symmetry with petals that are valvate in bud, have numerous showy, prominent stamens. Mimosoideae comprise 2,500 species; some classification systems, for example the Cronquist system, treat the Fabaceae in a narrow sense, raising the Mimisoideae to the rank of family as Mimosaceae. The Angiosperm Phylogeny Group treats Fabaceae in the broad sense; the Mimosoideae were subdivided into four tribes. However, modern molecular phylogenetics has shown. Several informal subgroups have been proposed, but not yet described formally as tribes. Additionally, the genus Acacia was segregated into five genera; the following fossil wood morphogenera have been described: Modern molecular phylogenetics suggests the following relationships: Acacieae is a wide-ranging, polyphyletic tribe of legumes in the Mimosoideae, native to the tropics and warm-temperate regions.
It includes six genera and some 1,450 species. Subdivision – 5 or 6 genera Acacia Mill. -- type genus Vachellia Arn. Senegalia Rafinesque Acaciella Britton & Rose Mariosousa Seigler & Ebinger Racosperma Martius In Bentham's 1842 circumscription of the subfamily Mimosoideae, Acacieae was one of its three constituent tribes, the others being Ingeae Benth. & Hook.f. and Mimoseae Bornn. His Acacieae tribe of 1842 included many genera that were subsequently assigned to tribe Ingeae Benth. In 1875, Bentham narrowed his definition of Acacieae so as to include only Acacia Mill; the only morphological character of Acacieae used to distinguish it from the Ingeae is the presence of free stamens. In the Ingeae they are fused in the form of a tube, whereas in the Acacieae only a few species have the stamens fused at the base. Several characters of the foliage, seed pods and stipules are shared by the two tribes; the flower morphology of Acacia s.l. has characteristics in common with the genera Leucaena and Mimosa and Enterolobium and Lysiloma.
The tribal position of monotypic genus Faidherbia A. Chevalier is equivocal, it was included in the Acacieae by Vassal and Maslin et al. but Lewis & Rico Arce placed it in tribe Ingeae following Polhill and Luckow et al.. In the latter case, tribe Acacieae may conform to genus Acacia s.l. pending the latter's relationship to other mimosoid genera. Faidherbia is troublesome as its stamens are shortly united at their base and its pollen is similar to some taxa in the Ingeae, they are shrubs or lianas, which may be armed or unarmed. Where they have spines, these are modified stipules. In some, prickles epidermis; the leaves are modified to vertically oriented phyllodes. A few have cladodes rather than leaves. Extrafloral nectaries may be present on the petiole and rachis, the pinnule tips may carry protein-lipid Beltian bodies; the leaflets are opposite, are carried on shortly stalks or are sessile. The heartwood is red and hard, the sap of various species hardens into gum; the inflorescences are dense pedunculate heads or spikes borne in axillary clusters, or are aggregated in terminal panicles.
The tetra - or pentamerous flowers are male and bisexual. Sepals are valvate; the reduced petals are valvate, or absent. The flowers have numerous exserted stamens, their filaments are sometimes connate at their base. Male flowers of some neotropical species have a reduced staminal tube. Flowers are yellow or cream-coloured, but may be white, red, or purple; the ovary is stipitate, with many ovules or ovules arranged in two rows. The ovary is attached by a filiform style to a capitate stigma; the legume's endocarp is attached to the exocarp, but is otherwise variable, may be dehiscent or indehiscent. Seeds are elliptic to oblong and flattened to varying degrees. Seeds have a hard black-brown testa with a pleurogram, visible as a closed or closed O-shaped line; some phyllodinous species have a colourful elaiosome on the seed. Data related to Acacieae at Wikispecies Media related to Mimosoideae at Wikimedia Commons
The Fabaceae or Leguminosae known as the legume, pea, or bean family, are a large and economically important family of flowering plants. It includes trees and perennial or annual herbaceous plants, which are recognized by their fruit and their compound, stipulate leaves. Many legumes have characteristic fruits; the family is distributed, is the third-largest land plant family in terms of number of species, behind only the Orchidaceae and Asteraceae, with about 751 genera and about 19,000 known species. The five largest of the genera are Astragalus, Indigofera and Mimosa, which constitute about a quarter of all legume species; the ca. 19,000 known legume species amount to about 7% of flowering plant species. Fabaceae is the most common family found in tropical rainforests and in dry forests in the Americas and Africa. Recent molecular and morphological evidence supports the fact that the Fabaceae is a single monophyletic family; this conclusion has been supported not only by the degree of interrelation shown by different groups within the family compared with that found among the Leguminosae and their closest relations, but by all the recent phylogenetic studies based on DNA sequences.
These studies confirm that the Fabaceae are a monophyletic group, related to the Polygalaceae and Quillajaceae families and that they belong to the order Fabales. Along with the cereals, some fruits and tropical roots, a number of Leguminosae have been a staple human food for millennia and their use is related to human evolution; the Fabaceae family includes a number of important agricultural and food plants, including Glycine max, Pisum sativum, Cicer arietinum, Medicago sativa, Arachis hypogaea, Ceratonia siliqua, Glycyrrhiza glabra. A number of species are weedy pests in different parts of the world, including: Cytisus scoparius, Robinia pseudoacacia, Ulex europaeus, Pueraria lobata, a number of Lupinus species; the name'Fabaceae' comes from the defunct genus Faba, now included in Vicia. The term "faba" comes from Latin, appears to mean "bean". Leguminosae is an older name still considered valid, refers to the fruit of these plants, which are called legumes. Fabaceae range in habit from giant trees to small annual herbs, with the majority being herbaceous perennials.
Plants have indeterminate inflorescences. The flowers have a short hypanthium and a single carpel with a short gynophore, after fertilization produce fruits that are legumes; the Leguminosae have a wide variety of growth forms, including trees, herbaceous plants, vines or lianas. The herbaceous plants can be annuals, biennials, or perennials, without basal or terminal leaf aggregations. Many Legumes have tendrils, they are epiphytes, or vines. The latter support themselves by means of shoots that twist around a support or through cauline or foliar tendrils. Plants can be mesophytes, or xerophytes; the leaves are alternate and compound. Most they are even- or odd-pinnately compound trifoliate and palmately compound, in the Mimosoideae and the Caesalpinioideae bipinnate, they always have stipules, which can be rather inconspicuous. Leaf margins are entire or serrate. Both the leaves and the leaflets have wrinkled pulvini to permit nastic movements. In some species, leaflets have evolved into tendrils.
Many species have leaves with structures that attract ants that protect the plant from herbivore insects. Extrafloral nectaries are common among the Mimosoideae and the Caesalpinioideae, are found in some Faboideae. In some Acacia, the modified hollow stipules are known as domatia. Many Fabaceae host bacteria in their roots within structures called root nodules; these bacteria, known as rhizobia, have the ability to take nitrogen gas out of the air and convert it to a form of nitrogen, usable to the host plant. This process is called nitrogen fixation; the legume, acting as a host, rhizobia, acting as a provider of usable nitrate, form a symbiotic relationship. The flowers have five fused sepals and five free petals, they are hermaphrodite, have a short hypanthium cup shaped. There are ten stamens and one elongated superior ovary, with a curved style, they are arranged in indeterminate inflorescences. Fabaceae are entomophilous plants, the flowers are showy to attract pollinators. In the Caesalpinioideae, the flowers are zygomorphic, as in Cercis, or nearly symmetrical with five equal petals in Bauhinia.
The upper petal is the innermost one, unlike in the Faboideae. Some species, like some in the genus Senna, have asymmetric flowers, with one of the lower petals larger than the opposing one, the style bent to one side; the calyx, corolla, or stamens can be showy in this group. In the Mimosoideae, the flowers are actinomorphic and arranged in globose inflorescences; the petals are small and the stamens, which can be more than just 10, have long, coloured filaments, which are the showiest part of the flower. All of the flowers in an inflorescence open at once. In the Faboideae, the flowers are zygom
The eudicots, Eudicotidae or eudicotyledons are a clade of flowering plants, called tricolpates or non-magnoliid dicots by previous authors. The botanical terms were introduced in 1991 by evolutionary botanist James A. Doyle and paleobotanist Carol L. Hotton to emphasize the evolutionary divergence of tricolpate dicots from earlier, less specialized, dicots; the close relationships among flowering plants with tricolpate pollen grains was seen in morphological studies of shared derived characters. These plants have a distinct trait in their pollen grains of exhibiting three colpi or grooves paralleling the polar axis. Molecular evidence confirmed the genetic basis for the evolutionary relationships among flowering plants with tricolpate pollen grains and dicotyledonous traits; the term means "true dicotyledons", as it contains the majority of plants that have been considered dicots and have characteristics of the dicots. The term "eudicots" has subsequently been adopted in botany to refer to one of the two largest clades of angiosperms, monocots being the other.
The remaining angiosperms include magnoliids and what are sometimes referred to as basal angiosperms or paleodicots, but these terms have not been or adopted, as they do not refer to a monophyletic group. The other name for the eudicots is tricolpates, a name which refers to the grooved structure of the pollen. Members of the group have tricolpate pollen; these pollens have three or more pores set in furrows called colpi. In contrast, most of the other seed plants produce monosulcate pollen, with a single pore set in a differently oriented groove called the sulcus; the name "tricolpates" is preferred by some botanists to avoid confusion with the dicots, a nonmonophyletic group. Numerous familiar plants are eudicots, including many common food plants and ornamentals; some common and familiar eudicots include members of the sunflower family such as the common dandelion, the forget-me-not and other members of its family, buttercup and macadamia. Most leafy trees of midlatitudes belong to eudicots, with notable exceptions being magnolias and tulip trees which belong to magnoliids, Ginkgo biloba, not an angiosperm.
The name "eudicots" is used in the APG system, of 1998, APG II system, of 2003, for classification of angiosperms. It is applied to a monophyletic group, which includes most of the dicots. "Tricolpate" is a synonym for the "Eudicot" monophyletic group, the "true dicotyledons". The number of pollen grain furrows or pores helps classify the flowering plants, with eudicots having three colpi, other groups having one sulcus. Pollen apertures are any modification of the wall of the pollen grain; these modifications include thinning and pores, they serve as an exit for the pollen contents and allow shrinking and swelling of the grain caused by changes in moisture content. The elongated apertures/ furrows in the pollen grain are called colpi, along with pores, are a chief criterion for identifying the pollen classes; the eudicots can be divided into two groups: the basal eudicots and the core eudicots. Basal eudicot is an informal name for a paraphyletic group; the core eudicots are a monophyletic group.
A 2010 study suggested the core eudicots can be divided into two clades, Gunnerales and a clade called "Pentapetalae", comprising all the remaining core eudicots. The Pentapetalae can be divided into three clades: Dilleniales superrosids consisting of Saxifragales and rosids superasterids consisting of Santalales, Berberidopsidales and asteridsThis division of the eudicots is shown in the following cladogram: The following is a more detailed breakdown according to APG IV, showing within each clade and orders: clade Eudicots order Ranunculales order Proteales order Trochodendrales order Buxales clade Core eudicots order Gunnerales order Dilleniales clade Superrosids order Saxifragales clade Rosids order Vitales clade Fabids order Fabales order Rosales order Fagales order Cucurbitales order Oxalidales order Malpighiales order Celastrales order Zygophyllales clade Malvids order Geraniales order Myrtales order Crossosomatales order Picramniales order Malvales order Brassicales order Huerteales order Sapindales clade Superasterids order Berberidopsidales order Santalales order Caryophyllales clade Asterids order Cornales order Ericales clade Campanulids order Aquifoliales order Asterales order Escalloniales order Bruniales order Apiales order Dipsacales order Paracryphiales clade Lamiids order Solanales order Lamiales order Vahliales order Gentianales order Boraginales order Garryales order Metteniusales order Icacinales Eudicots at the Encyclopedia of Life Eudicots, Tree of Life Web Project Dicots Plant Life Forms
The rosids are members of a large clade of flowering plants, containing about 70,000 species, more than a quarter of all angiosperms. The clade is divided into 16 to 20 orders, depending upon circumscription and classification; these orders, in turn, together comprise about 140 families. Fossil rosids are known from the Cretaceous period. Molecular clock estimates indicate that the rosids originated in the Aptian or Albian stages of the Cretaceous, between 125 and 99.6 million years ago. The name is based upon the name "Rosidae", understood to be a subclass. In 1967, Armen Takhtajan showed that the correct basis for the name "Rosidae" is a description of a group of plants published in 1830 by Friedrich Gottlieb Bartling; the clade was renamed "Rosidae" and has been variously delimited by different authors. The name "rosids" is informal and not assumed to have any particular taxonomic rank like the names authorized by the ICBN; the rosids are monophyletic based upon evidence found by molecular phylogenetic analysis.
Three different definitions of the rosids were used. Some authors included the orders Vitales in the rosids. Others excluded both of these orders; the circumscription used in this article is that of the APG IV classification, which includes Vitales, but excludes Saxifragales. The rosids and Saxifragales form the superrosids clade; this is one of three groups that compose the Pentapetalae, the others being Dilleniales and the superasterids. The rosids consist of two groups: the eurosids; the eurosids, in turn, are divided into two groups: malvids. The rosids consist of 17 orders. In addition to Vitales, there are 8 orders in malvids; some of the orders have only been recognized. These are Vitales, Crossosomatales and Huerteales; the phylogeny of Rosids shown below is adapted from the Angiosperm Phylogeny Group website. The nitrogen-fixing clade contains a high number of actinorhizal plants. Not all plants in this clade are actinorhizal, however. Media related to Rosids at Wikimedia Commons
Natural Resources Conservation Service
Natural Resources Conservation Service known as the Soil Conservation Service, is an agency of the United States Department of Agriculture that provides technical assistance to farmers and other private landowners and managers. Its name was changed in 1994 during the presidency of Bill Clinton to reflect its broader mission, it is a small agency comprising about 12,000 employees. Its mission is to improve and conserve natural resources on private lands through a cooperative partnership with state and local agencies. While its primary focus has been agricultural lands, it has made many technical contributions to soil surveying and water quality improvement. One example is the Conservation Effects Assessment Project, set up to quantify the benefits of agricultural conservation efforts promoted and supported by programs in the Farm Security and Rural Investment Act of 2002. NRCS is the leading agency in this project; the agency was founded through the efforts of Hugh Hammond Bennett, a soil conservation pioneer who worked for the Department of Agriculture from 1903 to 1952.
Bennett's motivation was based on his knowledge of the detrimental effects of soil erosion and the impacts on U. S lands. On September 13, 1933, the Soil Erosion Service was formed in the Department of the Interior, with Bennett as chief; the service was transferred to the Department of Agriculture on March 23, 1935, was shortly thereafter combined with other USDA units to form the Soil Conservation Service by the Soil Conservation and Domestic Allotment Act of 1935. The Soil Conservation Service was in charge of 500 Civilian Conservation Corps camps between 1933 and 1942; the primary purpose of these camps was erosion control. Hugh Bennett continued as chief, a position he held until his retirement in 1952. On October 20, 1994, the agency was renamed to the Natural Resources Conservation Service as part of the Federal Crop Insurance Reform and Department of Agriculture Reorganization Act of 1994. NRCS offers financial assistance to farmers and ranchers; the financial assistance is authorized by the Farm Bill, a law, renewed every five years.
The 2014 Farm Bill consolidated 23 programs into 15. NRCS offers these services to private land owners, conservation districts and other types of organizations. NRCS collects and shares information on the nation's soil, water and plants; the Conservation Title of the Farm Bill provides the funding to agricultural producers, a conservation plan must be included. All of these programs are voluntary; the main programs include: The purpose of EQIP is to provide assistance to landowners to help them improve their soil and related natural resources, including grazing lands and wildlife habitat. Conservation Stewardship Program CSP is targeted to a producers who maintain a higher level of environmental stewardship. Regional Conservation Partnership Program RCPP consolidated four programs from the prior 2008 Farm Bill, it aims at more watershed scale projects, rather than individual farms and ranches. Agricultural Conservation Easement Program ACEP was another consolidation effort of the 2014 Farm Bill, which includes the former Grasslands Reserve Program and Ranch Lands Protection Program, Wetlands Reserve Program.
ACEP includes technical and financial help to maintain or improve land for agriculture or environmental benefits. Landowners volunteer to protect forests in 30 or 10 year contracts; this program hands assisting funds to participants. The objectives of HFRP are to: Promote the recovery of endangered and threatened species under the Endangered Species Act Improve plant and animal biodiversity Enhance carbon sequestration Serves 10 states in the Midwest United States in helping to reduce Nitrate levels in soil due to runoff from fertilized farmland; the project began in 2010 and focused on the Mississippi Basin area. The main goal of the project is to implement better methods of managing water drainage from agricultural uses, in place of letting the water drain as it had done in the past. In October 2011, The National "Managing Water, Harvesting Results" Summit was held to promote the drainage techniques used in hopes of people adopting them nationwide. Includes water supply forecasts and the Surface Water Supply Index for Alaska and other Western states.
NRCS agents collect data from snowpack and mountain sites to predict spring runoff and summer streamflow amounts. These predictions are used in decision making for agriculture, wildlife management and development, several other areas; these predictions are available within the first 5 days of each month from January to June. Is a blanket program which involves conservation efforts on soil and water conservation, as well as management of agricultural wastes and general longterm sustainability. NRCS and related agencies work with landowners, communities, or developers to protect the environment. Serve to guide people to comply with acts such as the Highly Erodible Land and Conservation Compliance Provisions acts; the CTA can cover projects by state and federal governments. Is a program to assist gulf bordering states improve water quality and use sustainable methods of farming and other industry; the program will deliver up to 50 million dollars over 2011-2013 to apply these sustainable methods, as well as wildlife habitat management systems that do not hinder agricultural productivity, prevent future over use of water resources to protect native endangered spe
Flora of Australia
The flora of Australia comprises a vast assemblage of plant species estimated to over 20,000 vascular and 14,000 non-vascular plants, 250,000 species of fungi and over 3,000 lichens. The flora has strong affinities with the flora of Gondwana, below the family level has a endemic angiosperm flora whose diversity was shaped by the effects of continental drift and climate change since the Cretaceous. Prominent features of the Australian flora are adaptations to aridity and fire which include scleromorphy and serotiny; these adaptations are common in species from the large and well-known families Proteaceae and Fabaceae. The arrival of humans around 50,000 years ago and settlement by Europeans from 1788, has had a significant impact on the flora; the use of fire-stick farming by Aboriginal people led to significant changes in the distribution of plant species over time, the large-scale modification or destruction of vegetation for agriculture and urban development since 1788 has altered the composition of most terrestrial ecosystems, leading to the extinction of 61 plant species and endangering over 1000 more.
Austrial major commonwealth foundations Australia was part of the southern supercontinent Gondwana, which included South America, Africa and Antarctica. Most of the modern Australian flora had their origin in Gondwana during the Cretaceous when Australia was covered in subtropical rainforest. Australian ferns and gymnosperm bear strong resemblance to their Gondwanan ancestors, prominent members of the early Gondwanan angiosperm flora such as the Nothofagus and Proteaceae were present in Australia. Gondwana began to break up 140 million years ago; as Australia drifted and global climate change had a significant and lasting effect: a circumpolar oceanic current developed, atmospheric circulation increased as Australia moved away from Antarctica, precipitation fell, there was a slow warming of the continent and arid conditions started to develop. These conditions of geographic isolation and aridity led to the development of a more complex flora. From 25-10 MYA pollen records suggest the rapid radiation of species like Eucalyptus, Allocasuarina and the pea-flowered legumes, the development of open forest.
Collision with the Eurasian Plate led to additional South-east Asian and cosmopolitan elements entering the flora like the Lepidium and Chenopodioideae. The development of aridity and the old and nutrient poor soils of the continent led to some unique adaptations in the Australian flora and evolutionary radiation of genera – like Acacia and Eucalyptus – that adapted to those conditions. Hard leaves with a thick outer layer, a condition known as scleromorphy, C4 and CAM carbon fixation which reduce water loss during photosynthesis are two common adaptations in Australian arid-adapted dicot and monocot species respectively. Rising aridity increased the frequency of fires in Australia. Fire is thought to have played a role in the development and distribution of fire-adapted species from the Late Pleistocene. An increase in charcoal in sediment around 38,000 years ago coincides with dates for the inhabitation of Australia by the Indigenous Australians and suggests that man-made fires, from practices like fire-stick farming, have played an important role in the establishment and maintenance of sclerophyll forest on the east coast of Australia.
Adaptations to fire include lignotubers and epicormic buds in Eucalyptus and Banksia species that allow fast regeneration following fire. Some genera exhibit serotiny, the release of seed only in response to heat and/or smoke. Xanthorrhoea grass trees and some species of orchids only flower after fire. In biogeography and zoogeography, Australia alone is sometimes considered a realm, while some authors unite the area with other regions to form the Australasian realm. In phytogeography, the area is considered a floristic kingdom, with the following endemic families, according to Takhtajan: Platyzomataceae, Austrobaileyaceae, Gyrostemonaceae, Davidsoniaceae, Eremosynaceae, Emblingiaceae, Tremandraceae, Brunoniaceae, Doryanthaceae and Xanthorrhoeaceae, it is the center of origin of Eupomatiaceae, Epacridaceae, Stackhousiaceae and Goodeniaceae. Other families with high occurrences are Poaceae, Asteraceae, Euphorbiaceae, Rutaceae and Proteaceae. Australia's terrestrial flora can be collected into characteristic vegetation groups.
The most important determinant is rainfall, followed by temperature which affects water availability. Several schemes of varying complexity have been created, the most recent scheme developed by the Natural Heritage Trust divides Australia's terrestrial flora into 30 Major Vegetation Groups, 67 Major Vegetation Subgroups. According to the scheme the most common vegetation types are those that are adapted to arid conditions where the area has not been reduced by human activities such as land clearing for agriculture; the dominant vegetation type in Australia is the hummock grasslands that occur extensively in arid Western Australia, South Australia and the Northern Territory. It accounts for 23% of the native vegetation, the predominant species of which are from the genus Triodia. Zygochloa occurs in inland sandy areas like the Simpson Desert. A furt
Wikispecies is a wiki-based online project supported by the Wikimedia Foundation. Its aim is to create a comprehensive free content catalogue of all species. Jimmy Wales stated that editors are not required to fax in their degrees, but that submissions will have to pass muster with a technical audience. Wikispecies is available under the GNU Free Documentation License and CC BY-SA 3.0. Started in September 2004, with biologists across the world invited to contribute, the project had grown a framework encompassing the Linnaean taxonomy with links to Wikipedia articles on individual species by April 2005. Benedikt Mandl co-ordinated the efforts of several people who are interested in getting involved with the project and contacted potential supporters in early summer 2004. Databases were evaluated and the administrators contacted, some of them have agreed on providing their data for Wikispecies. Mandl defined two major tasks: Figure out how the contents of the data base would need to be presented—by asking experts, potential non-professional users and comparing that with existing databases Figure out how to do the software, which hardware is required and how to cover the costs—by asking experts, looking for fellow volunteers and potential sponsorsAdvantages and disadvantages were discussed by the wikimedia-I mailing list.
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