The Orchidaceae are a diverse and widespread family of flowering plants, with blooms that are colourful and fragrant known as the orchid family. Along with the Asteraceae, they are one of the two largest families of flowering plants; the Orchidaceae have about 28,000 accepted species, distributed in about 763 genera. The determination of which family is larger is still under debate, because verified data on the members of such enormous families are continually in flux. Regardless, the number of orchid species nearly equals the number of bony fishes and is more than twice the number of bird species, about four times the number of mammal species; the family encompasses about 6–11% of all seed plants. The largest genera are Bulbophyllum, Epidendrum and Pleurothallis, it includes Vanilla–the genus of the vanilla plant, the type genus Orchis, many cultivated plants such as Phalaenopsis and Cattleya. Moreover, since the introduction of tropical species into cultivation in the 19th century, horticulturists have produced more than 100,000 hybrids and cultivars.
Orchids are distinguished from other plants, as they share some evident, shared derived characteristics, or synapomorphies. Among these are: bilateral symmetry of the flower, many resupinate flowers, a nearly always modified petal, fused stamens and carpels, small seeds. All orchids are perennial herbs, they can grow according to two patterns: Monopodial: The stem grows from a single bud, leaves are added from the apex each year and the stem grows longer accordingly. The stem of orchids with a monopodial growth can reach several metres in length, as in Vanda and Vanilla. Sympodial: Sympodial orchids have a front and a back; the plant produces a series of adjacent shoots, which grow to a certain size and stop growing and are replaced. Sympodial orchids grow laterally following the surface of their support; the growth continues by development of new leads, with their own leaves and roots, sprouting from or next to those of the previous year, as in Cattleya. While a new lead is developing, the rhizome may start its growth again from a so-called'eye', an undeveloped bud, thereby branching.
Sympodial orchids may have visible pseudobulbs joined by a rhizome, which creeps along the top or just beneath the soil. Terrestrial orchids may form corms or tubers; the root caps of terrestrial orchids are white. Some sympodial terrestrial orchids, such as Orchis and Ophrys, have two subterranean tuberous roots. One is used as a food reserve for wintry periods, provides for the development of the other one, from which visible growth develops. In warm and humid climates, many terrestrial orchids do not need pseudobulbs. Epiphytic orchids, those that grow upon a support, have modified aerial roots that can sometimes be a few meters long. In the older parts of the roots, a modified spongy epidermis, called velamen, has the function of absorbing humidity, it can have a silvery-grey, white or brown appearance. In some orchids, the velamen includes spongy and fibrous bodies near the passage cells, called tilosomes; the cells of the root epidermis grow at a right angle to the axis of the root to allow them to get a firm grasp on their support.
Nutrients for epiphytic orchids come from mineral dust, organic detritus, animal droppings and other substances collecting among on their supporting surfaces. The base of the stem of sympodial epiphytes, or in some species the entire stem, may be thickened to form a pseudobulb that contains nutrients and water for drier periods; the pseudobulb has a smooth surface with lengthwise grooves, can have different shapes conical or oblong. Its size is variable; some Dendrobium species have long, canelike pseudobulbs with short, rounded leaves over the whole length. With ageing, the pseudobulb becomes dormant. At this stage, it is called a backbulb. Backbulbs still hold nutrition for the plant, but a pseudobulb takes over, exploiting the last reserves accumulated in the backbulb, which dies off, too. A pseudobulb lives for about five years. Orchids without noticeable pseudobulbs are said to have growths, an individual component of a sympodial plant. Like most monocots, orchids have simple leaves with parallel veins, although some Vanilloideae have reticulate venation.
Leaves may be ovate, lanceolate, or orbiculate, variable in size on the individual plant. Their characteristics are diagnostic, they are alternate on the stem folded lengthwise along the centre, have no stipules. Orchid leaves have siliceous bodies called stegmata in the vascular bundle sheaths and are fibrous; the structure of the leaves corresponds to the specific habitat of the plant. Species that bask in sunlight, or grow on sites which can be very dry, have thick, leathery leaves and the laminae are covered by a waxy cuticle to retain their necessary water supply. Shade-loving species, on the other hand, have thin leaves; the leaves of most orchids are perennial, that is, they live for several years, while others those with plicate leaves as in Catasetum, shed them annually and de
Barbosella cogniauxiana is a species of orchid. It is native to the Atlantic Forest in southern Brazil. Flora of Atlantic Forest List of plants of Atlantic Forest vegetation of Brazil Media related to Barbosella cogniauxiana at Wikimedia Commons Data related to Barbosella cogniauxiana at Wikispecies
In biology, a species is the basic unit of classification and a taxonomic rank of an organism, as well as a unit of biodiversity. A species is defined as the largest group of organisms in which any two individuals of the appropriate sexes or mating types can produce fertile offspring by sexual reproduction. Other ways of defining species include their karyotype, DNA sequence, behaviour or ecological niche. In addition, paleontologists use the concept of the chronospecies since fossil reproduction cannot be examined. While these definitions may seem adequate, when looked at more they represent problematic species concepts. For example, the boundaries between related species become unclear with hybridisation, in a species complex of hundreds of similar microspecies, in a ring species. Among organisms that reproduce only asexually, the concept of a reproductive species breaks down, each clone is a microspecies. All species are given a two-part name, a "binomial"; the first part of a binomial is the genus.
The second part is called the specific epithet. For example, Boa constrictor is one of four species of the genus Boa. None of these is satisfactory definitions, but scientists and conservationists need a species definition which allows them to work, regardless of the theoretical difficulties. If species were fixed and distinct from one another, there would be no problem, but evolutionary processes cause species to change continually, to grade into one another. Species were seen from the time of Aristotle until the 18th century as fixed kinds that could be arranged in a hierarchy, the great chain of being. In the 19th century, biologists grasped. Charles Darwin's 1859 book The Origin of Species explained how species could arise by natural selection; that understanding was extended in the 20th century through genetics and population ecology. Genetic variability arises from mutations and recombination, while organisms themselves are mobile, leading to geographical isolation and genetic drift with varying selection pressures.
Genes can sometimes be exchanged between species by horizontal gene transfer. Viruses are a special case, driven by a balance of mutation and selection, can be treated as quasispecies. Biologists and taxonomists have made many attempts to define species, beginning from morphology and moving towards genetics. Early taxonomists such as Linnaeus had no option but to describe what they saw: this was formalised as the typological or morphological species concept. Ernst Mayr emphasised reproductive isolation, but this, like other species concepts, is hard or impossible to test. Biologists have tried to refine Mayr's definition with the recognition and cohesion concepts, among others. Many of the concepts are quite similar or overlap, so they are not easy to count: the biologist R. L. Mayden recorded about 24 concepts, the philosopher of science John Wilkins counted 26. Wilkins further grouped the species concepts into seven basic kinds of concepts: agamospecies for asexual organisms biospecies for reproductively isolated sexual organisms ecospecies based on ecological niches evolutionary species based on lineage genetic species based on gene pool morphospecies based on form or phenotype and taxonomic species, a species as determined by a taxonomist.
A typological species is a group of organisms in which individuals conform to certain fixed properties, so that pre-literate people recognise the same taxon as do modern taxonomists. The clusters of variations or phenotypes within specimens would differentiate the species; this method was used as a "classical" method of determining species, such as with Linnaeus early in evolutionary theory. However, different phenotypes are not different species. Species named in this manner are called morphospecies. In the 1970s, Robert R. Sokal, Theodore J. Crovello and Peter Sneath proposed a variation on this, a phenetic species, defined as a set of organisms with a similar phenotype to each other, but a different phenotype from other sets of organisms, it differs from the morphological species concept in including a numerical measure of distance or similarity to cluster entities based on multivariate comparisons of a reasonably large number of phenotypic traits. A mate-recognition species is a group of sexually reproducing organisms that recognize one another as potential mates.
Expanding on this to allow for post-mating isolation, a cohesion species is the most inclusive population of individuals having the potential for phenotypic cohesion through intrinsic cohesion mechanisms. A further development of the recognition concept is provided by the biosemiotic concept of species. In microbiology, genes can move even between distantly related bacteria extending to the whole bacterial domain; as a rule of thumb, microbiologists have assumed that kinds of Bacteria or Archaea with 16S ribosomal RNA gene sequences more similar than 97% to each other need to be checked by DNA-DNA hybridisation to decide if they belong to the same species or not. This concept was narrowed in 2006 to a similarity of 98.7%. DNA-DNA hybri
The tribe Epidendreae of the Orchidaceae comprises six subtribes: Bletiinae sensu MMIV, which contains only the genera Basiphyllaea and Hexalectris Chysinae Coeliinae Laeliinae Pleurothallidinae Ponerinae
Wikispecies is a wiki-based online project supported by the Wikimedia Foundation. Its aim is to create a comprehensive free content catalogue of all species. Jimmy Wales stated that editors are not required to fax in their degrees, but that submissions will have to pass muster with a technical audience. Wikispecies is available under the GNU Free Documentation License and CC BY-SA 3.0. Started in September 2004, with biologists across the world invited to contribute, the project had grown a framework encompassing the Linnaean taxonomy with links to Wikipedia articles on individual species by April 2005. Benedikt Mandl co-ordinated the efforts of several people who are interested in getting involved with the project and contacted potential supporters in early summer 2004. Databases were evaluated and the administrators contacted, some of them have agreed on providing their data for Wikispecies. Mandl defined two major tasks: Figure out how the contents of the data base would need to be presented—by asking experts, potential non-professional users and comparing that with existing databases Figure out how to do the software, which hardware is required and how to cover the costs—by asking experts, looking for fellow volunteers and potential sponsorsAdvantages and disadvantages were discussed by the wikimedia-I mailing list.
The board of directors of the Wikimedia Foundation voted by 4 to 0 in favor of the establishment of a Wikispecies. The project is hosted at species.wikimedia.org. It was merged to a sister project of Wikimedia Foundation on September 14, 2004. On October 10, 2006, the project exceeded 75,000 articles. On May 20, 2007, the project exceeded 100,000 articles with a total of 5,495 registered users. On September 8, 2008, the project exceeded 150,000 articles with a total of 9,224 registered users. On October 23, 2011, the project reached 300,000 articles. On June 16, 2014, the project reached 400,000 articles. On January 7, 2017, the project reached 500,000 articles. On October 30, 2018, the project reached 600,000 articles, a total of 1.12 million pages. Wikispecies comprises taxon pages, additionally pages about synonyms, taxon authorities, taxonomical publications, institutions or repositories holding type specimen. Wikispecies asks users to use images from Wikimedia Commons. Wikispecies does not allow the use of content.
All Species Foundation Catalogue of Life Encyclopedia of Life Tree of Life Web Project List of online encyclopedias The Plant List Wikispecies, The free species directory that anyone can edit Species Community Portal The Wikispecies Charter, written by Wales
Brazil the Federative Republic of Brazil, is the largest country in both South America and Latin America. At 8.5 million square kilometers and with over 208 million people, Brazil is the world's fifth-largest country by area and the fifth most populous. Its capital is Brasília, its most populated city is São Paulo; the federation is composed of the union of the 26 states, the Federal District, the 5,570 municipalities. It is the largest country to have Portuguese as an official language and the only one in the Americas. Bounded by the Atlantic Ocean on the east, Brazil has a coastline of 7,491 kilometers, it borders all other South American countries except Ecuador and Chile and covers 47.3% of the continent's land area. Its Amazon River basin includes a vast tropical forest, home to diverse wildlife, a variety of ecological systems, extensive natural resources spanning numerous protected habitats; this unique environmental heritage makes Brazil one of 17 megadiverse countries, is the subject of significant global interest and debate regarding deforestation and environmental protection.
Brazil was inhabited by numerous tribal nations prior to the landing in 1500 of explorer Pedro Álvares Cabral, who claimed the area for the Portuguese Empire. Brazil remained a Portuguese colony until 1808, when the capital of the empire was transferred from Lisbon to Rio de Janeiro. In 1815, the colony was elevated to the rank of kingdom upon the formation of the United Kingdom of Portugal and the Algarves. Independence was achieved in 1822 with the creation of the Empire of Brazil, a unitary state governed under a constitutional monarchy and a parliamentary system; the ratification of the first constitution in 1824 led to the formation of a bicameral legislature, now called the National Congress. The country became a presidential republic in 1889 following a military coup d'état. An authoritarian military junta came to power in 1964 and ruled until 1985, after which civilian governance resumed. Brazil's current constitution, formulated in 1988, defines it as a democratic federal republic. Due to its rich culture and history, the country ranks thirteenth in the world by number of UNESCO World Heritage Sites.
Brazil is considered an advanced emerging economy. It has the ninth largest GDP in the world by nominal, eight and PPP measures, it is one of the world's major breadbaskets, being the largest producer of coffee for the last 150 years. It is classified as an upper-middle income economy by the World Bank and a newly industrialized country, with the largest share of global wealth in Latin America. Brazil is a regional power and sometimes considered a great or a middle power in international affairs. On account of its international recognition and influence, the country is subsequently classified as an emerging power and a potential superpower by several analysts. Brazil is a founding member of the United Nations, the G20, BRICS, Union of South American Nations, Organization of American States, Organization of Ibero-American States and the Community of Portuguese Language Countries, it is that the word "Brazil" comes from the Portuguese word for brazilwood, a tree that once grew plentifully along the Brazilian coast.
In Portuguese, brazilwood is called pau-brasil, with the word brasil given the etymology "red like an ember", formed from brasa and the suffix -il. As brazilwood produces a deep red dye, it was valued by the European textile industry and was the earliest commercially exploited product from Brazil. Throughout the 16th century, massive amounts of brazilwood were harvested by indigenous peoples along the Brazilian coast, who sold the timber to European traders in return for assorted European consumer goods; the official Portuguese name of the land, in original Portuguese records, was the "Land of the Holy Cross", but European sailors and merchants called it the "Land of Brazil" because of the brazilwood trade. The popular appellation eclipsed and supplanted the official Portuguese name; some early sailors called it the "Land of Parrots". In the Guarani language, an official language of Paraguay, Brazil is called "Pindorama"; this was the name the indigenous population gave to the region, meaning "land of the palm trees".
Some of the earliest human remains found in the Americas, Luzia Woman, were found in the area of Pedro Leopoldo, Minas Gerais and provide evidence of human habitation going back at least 11,000 years. The earliest pottery found in the Western Hemisphere was excavated in the Amazon basin of Brazil and radiocarbon dated to 8,000 years ago; the pottery was found near Santarém and provides evidence that the tropical forest region supported a complex prehistoric culture. The Marajoara culture flourished on Marajó in the Amazon delta from 800 CE to 1400 CE, developing sophisticated pottery, social stratification, large populations, mound building, complex social formations such as chiefdoms. Around the time of the Portuguese arrival, the territory of current day Brazil had an estimated indigenous population of 7 million people semi-nomadic who subsisted on hunting, fishing and migrant agriculture; the indigenous population of Brazil comprised several large indigenous ethnic groups. The Tupí people were subdivided into the Tupiniquins and Tupinambás, there were many subdivisions of the other gro
João Barbosa Rodrigues
João Barbosa Rodrigues was considered one of Brazil's greatest botanists, known for his work on orchids and palms. For nearly two decades he was director of the Botanic Garden of Rio de Janeiro. Something of a polymath, he was a prolific botanical artist who made contributions to his country's ethnography, linguistics and literature. Rodrigues was born on June 22, 1842, in São Gonçalo do Sapucaí, was raised in Campanha, in the state of Minas Gerais, before the family returned to Rio in 1858, his father was a Portuguese merchant, his mother was a Brazilian of Indian descent. He had Arthur and sisters Maria and Olympia, he showed early ability as a writer and he was always interested in natural science in collecting insects and plants. However, he went to the Central School of Engineering in Rio de Janeiro, where he got a solid classical education. Only after graduating in 1869 did he begin to pursue his scientific interests. Rodrigues became a teacher of drawing at the Colegio Pedro II who specialized in botany, under the supervision of Francisco Freire Allemão e Cysneiro.
He began his botanical expeditions in 1868 and was commissioned by the Brazilian government in 1871 to explore the Amazon basin and study palms, in part due to sponsorship by Guilherme Schüch, Baron of Capanema. This expedition lasted more than three years, during which time which he drew not only plants but animals and Indian artefacts. By the early 1890s, he had created nearly 900 color plates of Brazilian orchids; because of difficulty financing a publication with illustrations, Rodrigues published descriptions of more than 540 new orchids and 28 new genera without any images in his important two-volume work Genera et species orchidearum novarum. He allowed another botanist, Alfred Cogniaux, to publish black-and-white versions of some 260 of his drawings in Cogniaux's Flora Brasiliensis. Nearly one-third of the species included in Flora Brasiliensis were described by Rodrigues. Cogniaux gave the British botanical artist Harriet Anne Hooker Thiselton-Dyer access to the Rodrigues drawings in his possession, she copied some 550 of them for the Kew Gardens collection before they were returned to Brazil.
The original drawings disappeared sometime after Rodrigues's death and are now lost, making Thiselton-Dyer's copies a uniquely valuable resource on Rodrigues's work. In 1875, Rodrigues published Enumeratio Palmarum Novarum. In subsequent years, an English botanist who had met Rodrigues in Brazil, James William Helenus Trail, published some of the same species and claimed precedence over Rodrigues. Despite this dispute over primacy, which lasted for many years, in 1903, Rodrigues published a classic work on palms, Setum Palmarum Brasiliensis; this was a survey of over 380 species of palms. Financed by the Brazilian government, it was printed in Belgium and lavishly illustrated with some 170 of Rodrigues's watercolors. After the Amazon expedition of the 1970s, Rodrigues became discouraged over the difficulty of funding another expedition and detoured into business, running a chemical factory. Not long afterwards, he was tasked to organize and direct a brand-new Botanic Museum of the Amazon in Manaus, which opened in 1883 under the sponsorship of Princess Isabel.
He founded Vellosia, to showcase the museum's work. Despite its royal patronage, the museum struggled financially and closed in 1890 when Rodrigues moved on to become director of the Botanic Garden of Rio de Janeiro, a position he would hold until his death. At the height of its development, the Botanic Museum of the Amazon had held some 10,000 botanical specimens, nearly all of which were lost on the museum's closure. During his directorship of the Rio de Janeiro botanic garden, Rodrigues constructed a greenhouse and library, diverted the Monkey River and landscaped new areas of the garden, planted an arboretum increased the size of the live plant collection, took steps towards organizing a herbarium, he started a journal to publish the botanic garden's scientific work, the Contributions du Jardin Botanique du Rio de Janeiro. Rodrigues died on March 6, 1909. Rodrigues soon lost his wife to tuberculosis, he married again a few years and once again was widowed, this time being left with a small daughter.
His third marriage, to Constança Pacca, lasted many years and produced seven more daughters and six sons. He named the orchid genus Constantia after her; the genera Barbosa, Barbosella and Brodriguesia are named after Rodrigues. The Brazilian botanical journal Rodriguésia is named after him, as is the Barbosa Rodrigues herbarium in Itajaí. On the centenary of his birth, the João Barbosa Rodrigues Botany Museum was named in his honor. In 1996, his Iconograhie des orchidées du Brésil, which he began working on in the 1860s, was published for the first time