Binomial nomenclature called binominal nomenclature or binary nomenclature, is a formal system of naming species of living things by giving each a name composed of two parts, both of which use Latin grammatical forms, although they can be based on words from other languages. Such a name is called a binomen, binominal name or a scientific name; the first part of the name – the generic name – identifies the genus to which the species belongs, while the second part – the specific name or specific epithet – identifies the species within the genus. For example, humans belong within this genus to the species Homo sapiens. Tyrannosaurus rex is the most known binomial; the formal introduction of this system of naming species is credited to Carl Linnaeus beginning with his work Species Plantarum in 1753. But Gaspard Bauhin, in as early as 1623, had introduced in his book Pinax theatri botanici many names of genera that were adopted by Linnaeus; the application of binomial nomenclature is now governed by various internationally agreed codes of rules, of which the two most important are the International Code of Zoological Nomenclature for animals and the International Code of Nomenclature for algae and plants.
Although the general principles underlying binomial nomenclature are common to these two codes, there are some differences, both in the terminology they use and in their precise rules. In modern usage, the first letter of the first part of the name, the genus, is always capitalized in writing, while that of the second part is not when derived from a proper noun such as the name of a person or place. Both parts are italicized when a binomial name occurs in normal text, thus the binomial name of the annual phlox is now written as Phlox drummondii. In scientific works, the authority for a binomial name is given, at least when it is first mentioned, the date of publication may be specified. In zoology "Patella vulgata Linnaeus, 1758"; the name "Linnaeus" tells the reader who it was that first published a description and name for this species of limpet. "Passer domesticus". The original name given by Linnaeus was Fringilla domestica; the ICZN does not require that the name of the person who changed the genus be given, nor the date on which the change was made, although nomenclatorial catalogs include such information.
In botany "Amaranthus retroflexus L." – "L." is the standard abbreviation used in botany for "Linnaeus". "Hyacinthoides italica Rothm. – Linnaeus first named this bluebell species Scilla italica. The name is composed of two word-forming elements: "bi", a Latin prefix for two, "-nomial", relating to a term or terms; the word "binomium" was used in Medieval Latin to mean a two-term expression in mathematics. Prior to the adoption of the modern binomial system of naming species, a scientific name consisted of a generic name combined with a specific name, from one to several words long. Together they formed a system of polynomial nomenclature; these names had two separate functions. First, to designate or label the species, second, to be a diagnosis or description. In a simple genus, containing only two species, it was easy to tell them apart with a one-word genus and a one-word specific name; such "polynomial names" may sometimes look like binomials, but are different. For example, Gerard's herbal describes various kinds of spiderwort: "The first is called Phalangium ramosum, Branched Spiderwort.
The other... is aptly termed Phalangium Ephemerum Virginianum, Soon-Fading Spiderwort of Virginia". The Latin phrases are short descriptions, rather than identifying labels; the Bauhins, in particular Caspar Bauhin, took some important steps towards the binomial system, by pruning the Latin descriptions, in many cases to two words. The adoption by biologists of a system of binomial nomenclature is due to Swedish botanist and physician Carl von Linné, more known by his Latinized name Carl Linnaeus, it was in his 1753 Species Plantarum that he first began using a one-word "trivial name" together with a generic name in a system of binomial nomenclature. This trivial name is what is now known as specific name; the Bauhins' genus names were retained in many of these, but the descriptive part was reduced to a single word. Linnaeus's trivial names introduced an important new idea, namely that the function of a name could be to give a species a unique label; this meant. Thus Gerard's Phalangium ephemerum virginianum became Tradescantia virgi
Cypriniformes is an order of ray-finned fish, including the carps, minnows and relatives. This order contains 11-12 families, over 400 genera, more than 4,250 species, with new species being described every few months or so, new genera being recognized frequently, they are most diverse in southeastern Asia, are absent from Australia and South America. Their closest living relatives are the Gymnotiformes and the Siluriformes. Like other orders of the Ostariophysi, fishes of cypriniformes possess a Weberian apparatus, they differ from most of their relatives in having only a dorsal fin on their back. Further differences are the Cypriniformes' unique kinethmoid, a small median bone in the snout, the lack of teeth in the mouth. Instead, they have convergent structures called pharyngeal teeth in the throat. While other groups of fish, such as cichlids possess pharyngeal teeth, the cypriniformes' teeth grind against a chewing pad on the base of the skull, instead of an upper pharyngeal jaw; the most notable family placed here is Cyprinidae.
This is one of the largest families of fish, is distributed across Africa and North America. Most species are freshwater inhabitants, but a considerable number are found in brackish water, such as roach and bream. At least one species is found in the Pacific redfin, Tribolodon brandtii. Brackish water and marine cyprinids are invariably anadromous, swimming upstream into rivers to spawn. Sometimes separated as family Psilorhynchidae, they seem to be specially-adapted fishes of Cyprinidae. Balitoridae and Gyrinocheilidae are families of mountain stream fishes feeding on algae and small invertebrates, they are found only in subtropical Asia. While the former are a speciose group, the latter contain only a handful of species; the suckers are found in temperate North eastern Asia. These large fishes are similar to carps in ecology. Members of Cobitidae common across Eurasia and parts of North Africa. A mid-sized group like the suckers, they are rather similar to catfish in appearance and behaviour, feeding off the substrate and equipped with barbels to help them locate food at night or in murky conditions.
Fishes in the families Cobitidae, Balitoridae and Gyrinocheilidae are called loaches, although it seems that the last do not belong to the lineage of "true" loaches but are related to the suckers. These included all the forms now placed in the superorder Ostariophysi except the catfish, which were placed in the order Siluriformes. By this definition, the Cypriniformes were paraphyletic, so the orders Gonorhynchiformes and Gymnotiformes have been separated out to form their own monophyletic orders; the families of Cypriniformes are traditionally divided into two superfamilies. Superfamily Cyprinioidea contains the carps and minnows and the mountain carps as the family Psilorhynchidae. In 2012 Maurice Kottelat reviewed the superfamily Cobitoidei and under his revision it now consists of the following families: hillstream loaches, Botiidae, true loaches, Gastromyzontidae, sucking loaches, stone loaches, Serpenticobitidae and long-finned loaches. Catostomoidea is treated as a junior synonym of Cobitoidei.
But it seems that it could be split off the Catostomidae and Gyrinocheilidae in a distinct superfamily. While the Cyprinioidea seem more "primitive" than the loach-like forms, they were successful enough never to shift from the original ecological niche of the basal Ostariophysi. Yet, from the ecomorphologically conservative main lineage at least two major radiations branched off; these diversified from the lowlands into torrential river habitats, acquiring similar habitus and adaptations in the process. The mountain carps are apomorphic Cyprinidae close to true carps, or maybe to the danionins. While some details about the phylogenetic structures of this massively diverse family are known – e.g. that Cultrinae and Leuciscinae are rather close relatives and stand apart from Cyprininae – there is no good consensus yet on how the main lineages are interrelated. A systematic list, from the most ancient to the most modern lineages, can thus be given as: Superfamily Cyprinoidei Family Cyprinidae Bonaparte, 1840 and minnows incl.
Psilorhynchidae) Superfamily Cobitoidei Superfamily Catostomoidea Family Catostomidae Agassiz 1850 Superfamily Gyrinocheiloidea Family Gyrinocheilidae Gill 1905 Superfamily Cobitoidea Family Barbuccidae Kottelat 2012 Family Serpenticobitidae Kottelat 2012 Family Botiidae Berg 1940 Family Vaillantellidae Nalbant & Bănărescu 1977 Family Cobitidae Swainson 1838 Family Balitoridae Swainson 1839 Family Gastromyzontidae Fowler 1905 Family Ellopostomatidae Bohlen & Šlechtová 2009 Family Nemacheilidae Regan 1911 Phylogeny based on the work of the following works Cypriniformes include the most primitive of the Ostariophysi in the narrow sense. This is evidenced n
Sven O. Kullander
Sven Oscar Kullander is a Swedish biologist specialised in ichthyology. He researches cichlids – notably the genus Apistogramma and the Cichlasoma-complex – and other tropical fresh water fishes, he has been working with endangered fish species in Sweden. He studied at the universities of Umeå and Stockholm, took his Ph. D. in Stockholm in 1984. He is senior curator at the Swedish Museum of Natural History in Stockholm, with the responsibility for the ichthyologic and herpetologic collections. Kullander coordinates the museum's contributions to FishBase. Kullander has produced more than 100 scientific and popular publications on fishes, described many groups and new species of cichlids; the Swedish aquarists' magazine Tidskriften Akvariet gave him "Akvariets Oscar" in 1996 for his great contribution to the aquarium hobby. His wife Fang Fang Kullander was an ichthyologist at the Swedish Museum. Kullander, S. O. 1983. A revision of the South American Cichlid genus Cichlasoma. Swedish Museum of Natural History, Stockholm.
ISBN 91-86510-01-0 Kullander, S. O. 1986. Cichlid fishes of the Amazon River drainage of Peru. Department of Vertebrate Zoology, Swedish Museum of Natural History, Stockholm. ISBN 91-86510-04-5 Kullander, S. O. & H. Nijssen. 1989. The cichlids of Surinam: Teleostei, Labroidei. E. J. Brill, Leiden. ISBN 90-04-09077-0 Kullander, S. O. T. Stach, H. G. Hansson, B. Delling & H. Blom. 2011. Nationalnyckeln till Sveriges flora och fauna. Ryggsträngsdjur: lansettfiskar – broskfiskar, Chordata: Branchiostomatidae – Chondrichthyes. ArtDatabanken, Uppsala Sven O. Kullander Swedish Museum of Natural History Kullander's publications
Wikispecies is a wiki-based online project supported by the Wikimedia Foundation. Its aim is to create a comprehensive free content catalogue of all species. Jimmy Wales stated that editors are not required to fax in their degrees, but that submissions will have to pass muster with a technical audience. Wikispecies is available under the GNU Free Documentation License and CC BY-SA 3.0. Started in September 2004, with biologists across the world invited to contribute, the project had grown a framework encompassing the Linnaean taxonomy with links to Wikipedia articles on individual species by April 2005. Benedikt Mandl co-ordinated the efforts of several people who are interested in getting involved with the project and contacted potential supporters in early summer 2004. Databases were evaluated and the administrators contacted, some of them have agreed on providing their data for Wikispecies. Mandl defined two major tasks: Figure out how the contents of the data base would need to be presented—by asking experts, potential non-professional users and comparing that with existing databases Figure out how to do the software, which hardware is required and how to cover the costs—by asking experts, looking for fellow volunteers and potential sponsorsAdvantages and disadvantages were discussed by the wikimedia-I mailing list.
The board of directors of the Wikimedia Foundation voted by 4 to 0 in favor of the establishment of a Wikispecies. The project is hosted at species.wikimedia.org. It was merged to a sister project of Wikimedia Foundation on September 14, 2004. On October 10, 2006, the project exceeded 75,000 articles. On May 20, 2007, the project exceeded 100,000 articles with a total of 5,495 registered users. On September 8, 2008, the project exceeded 150,000 articles with a total of 9,224 registered users. On October 23, 2011, the project reached 300,000 articles. On June 16, 2014, the project reached 400,000 articles. On January 7, 2017, the project reached 500,000 articles. On October 30, 2018, the project reached 600,000 articles, a total of 1.12 million pages. Wikispecies comprises taxon pages, additionally pages about synonyms, taxon authorities, taxonomical publications, institutions or repositories holding type specimen. Wikispecies asks users to use images from Wikimedia Commons. Wikispecies does not allow the use of content.
All Species Foundation Catalogue of Life Encyclopedia of Life Tree of Life Web Project List of online encyclopedias The Plant List Wikispecies, The free species directory that anyone can edit Species Community Portal The Wikispecies Charter, written by Wales
Animals are multicellular eukaryotic organisms that form the biological kingdom Animalia. With few exceptions, animals consume organic material, breathe oxygen, are able to move, can reproduce sexually, grow from a hollow sphere of cells, the blastula, during embryonic development. Over 1.5 million living animal species have been described—of which around 1 million are insects—but it has been estimated there are over 7 million animal species in total. Animals range in length from 8.5 millionths of a metre to 33.6 metres and have complex interactions with each other and their environments, forming intricate food webs. The category includes humans, but in colloquial use the term animal refers only to non-human animals; the study of non-human animals is known as zoology. Most living animal species are in the Bilateria, a clade whose members have a bilaterally symmetric body plan; the Bilateria include the protostomes—in which many groups of invertebrates are found, such as nematodes and molluscs—and the deuterostomes, containing the echinoderms and chordates.
Life forms interpreted. Many modern animal phyla became established in the fossil record as marine species during the Cambrian explosion which began around 542 million years ago. 6,331 groups of genes common to all living animals have been identified. Aristotle divided animals into those with those without. Carl Linnaeus created the first hierarchical biological classification for animals in 1758 with his Systema Naturae, which Jean-Baptiste Lamarck expanded into 14 phyla by 1809. In 1874, Ernst Haeckel divided the animal kingdom into the multicellular Metazoa and the Protozoa, single-celled organisms no longer considered animals. In modern times, the biological classification of animals relies on advanced techniques, such as molecular phylogenetics, which are effective at demonstrating the evolutionary relationships between animal taxa. Humans make use of many other animal species for food, including meat and eggs. Dogs have been used in hunting, while many aquatic animals are hunted for sport.
Non-human animals have appeared in art from the earliest times and are featured in mythology and religion. The word "animal" comes from the Latin animalis, having soul or living being; the biological definition includes all members of the kingdom Animalia. In colloquial usage, as a consequence of anthropocentrism, the term animal is sometimes used nonscientifically to refer only to non-human animals. Animals have several characteristics. Animals are eukaryotic and multicellular, unlike bacteria, which are prokaryotic, unlike protists, which are eukaryotic but unicellular. Unlike plants and algae, which produce their own nutrients animals are heterotrophic, feeding on organic material and digesting it internally. With few exceptions, animals breathe oxygen and respire aerobically. All animals are motile during at least part of their life cycle, but some animals, such as sponges, corals and barnacles become sessile; the blastula is a stage in embryonic development, unique to most animals, allowing cells to be differentiated into specialised tissues and organs.
All animals are composed of cells, surrounded by a characteristic extracellular matrix composed of collagen and elastic glycoproteins. During development, the animal extracellular matrix forms a flexible framework upon which cells can move about and be reorganised, making the formation of complex structures possible; this may be calcified, forming structures such as shells and spicules. In contrast, the cells of other multicellular organisms are held in place by cell walls, so develop by progressive growth. Animal cells uniquely possess the cell junctions called tight junctions, gap junctions, desmosomes. With few exceptions—in particular, the sponges and placozoans—animal bodies are differentiated into tissues; these include muscles, which enable locomotion, nerve tissues, which transmit signals and coordinate the body. There is an internal digestive chamber with either one opening or two openings. Nearly all animals make use of some form of sexual reproduction, they produce haploid gametes by meiosis.
These fuse to form zygotes, which develop via mitosis into a hollow sphere, called a blastula. In sponges, blastula larvae swim to a new location, attach to the seabed, develop into a new sponge. In most other groups, the blastula undergoes more complicated rearrangement, it first invaginates to form a gastrula with a digestive chamber and two separate germ layers, an external ectoderm and an internal endoderm. In most cases, a third germ layer, the mesoderm develops between them; these germ layers differentiate to form tissues and organs. Repeated instances of mating with a close relative during sexual reproduction leads to inbreeding depression within a population due to the increased prevalence of harmful recessive traits. Animals have evolved numerous mechanisms for avoiding close inbreeding. In some species, such as the splendid fairywren, females benefit by mating with multiple males, thus producing more offspring of higher genetic quality; some animals are capable of asexual reproduction, which results
The Cyprinidae are the family of freshwater fishes, collectively called cyprinids, that includes the carps, the true minnows, their relatives. Called the "carp family", or "minnow family", Cyprinidae is the largest and most diverse fish family and the largest vertebrate animal family in general, with about 3,000 species of which only 1,270 remain extant, divided into about 370 genera.. They range from about 12 mm to the 3-meter Catlocarpio siamensis; this family of fish is one of the few. The family belongs to the ostariophysian order Cypriniformes, of whose genera and species the cyprinids make more than two-thirds; the family name is derived from the Ancient Greek kyprînos. Cyprinids are stomachless fish with toothless jaws. So, food can be chewed by the gill rakers of the specialized last gill bow; these pharyngeal teeth allow the fish to make chewing motions against a chewing plate formed by a bony process of the skull. The pharyngeal teeth are used by scientists to identify species. Strong pharyngeal teeth allow fish such as the common carp and ide to eat hard baits such as snails and bivalves.
Hearing is a well-developed sense in the cyprinids since they have the Weberian organ, three specialized vertebral processes that transfer motion of the gas bladder to the inner ear. The vertebral processes of the Weberian organ permit a cyprinid to detect changes in motion of the gas bladder due to atmospheric conditions or depth changes; the cyprinids are considered physostomes because the pneumatic duct is retained in adult stages and the fish are able to gulp air to fill the gas bladder, or they can dispose excess gas to the gut. Cyprinids are native to North America and Eurasia; the largest known cyprinid is the giant barb, which may grow up to 3 m in length and 300 kg in weight. Other large species that can surpass 2 m are the golden mahseer and mangar; the largest North American species is the Colorado pikeminnow, which can reach up to 1.8 m in length. Conversely, many species are smaller than 5 cm; the smallest known fish is Paedocypris progenetica, reaching 10.3 mm at the longest. All fish in this family most do not guard their eggs.
The bitterlings of subfamily Acheilognathinae are notable for depositing their eggs in bivalve molluscs, where the young develop until able to fend for themselves. Most cyprinids feed on invertebrates and vegetation due to the lack of teeth and stomach. Many species, such as the ide and the common rudd, prey on small fish when individuals become large enough. Small species, such as the moderlieschen, are opportunistic predators that will eat larvae of the common frog in artificial circumstances; some cyprinids, such as the grass carp, are specialized herbivores. For this reason, cyprinids are introduced as a management tool to control various factors in the aquatic environment, such as aquatic vegetation and diseases transmitted by snails. Unlike most fish species, cyprinids increase in abundance in eutrophic lakes. Here, they contribute towards positive feedback as they are efficient at eating the zooplankton that would otherwise graze on the algae, reducing its abundance. Cyprinids are important food fish.
In land-locked countries in particular, cyprinids are the major species of fish eaten because they make the largest part of biomass in most water types except for fast-flowing rivers. In Eastern Europe, they are prepared with traditional methods such as drying and salting; the prevalence of inexpensive frozen fish products made this less important now than it was in earlier times. Nonetheless, in certain places, they remain popular for food, as well as recreational fishing, have been deliberately stocked in ponds and lakes for centuries for this reason. Cyprinids are popular for angling for match fishing and fishing for common carp because of its size and strength. Several cyprinids have been introduced to waters outside their natural ranges to provide food, sport, or biological control for some pest species; the common carp and the grass carp are the most important for example in Florida. In some cases, such as the Asian carp in the Mississippi Basin, they have become invasive species that compete with native fishes or disrupt the environment.
Carp in particular can stir up sediment, reducing the clarity of the water and making it difficult for plants to grow. Numerous cyprinids have become important in the aquarium and fishpond hobbies, most famously the goldfish, bred in China from the Prussian carp. First imported into Europe around 1728, it was much fancied by Chinese nobility as early as 1150 AD and after it arrived there in 1502 in Japan. In the latter country, from the 18th century onwards, the common carp was bred into the ornamental variety known as koi – or more nishikigoi, as koi means "common carp" in Japanese. Other popular aquarium cyprinids include danionins and true barbs. Larger species are bred by the thousands in outdoor ponds in Southeast Asia, trade in these aquarium fishes
A chordate is an animal constituting the phylum Chordata. During some period of their life cycle, chordates possess a notochord, a dorsal nerve cord, pharyngeal slits, an endostyle, a post-anal tail: these five anatomical features define this phylum. Chordates are bilaterally symmetric; the Chordata and Ambulacraria together form the superphylum Deuterostomia. Chordates are divided into three subphyla: Vertebrata. There are extinct taxa such as the Vetulicolia. Hemichordata has been presented as a fourth chordate subphylum, but now is treated as a separate phylum: hemichordates and Echinodermata form the Ambulacraria, the sister phylum of the Chordates. Of the more than 65,000 living species of chordates, about half are bony fish that are members of the superclass Osteichthyes. Chordate fossils have been found from as early as the Cambrian explosion, 541 million years ago. Cladistically, vertebrates - chordates with the notochord replaced by a vertebral column during development - are considered to be a subgroup of the clade Craniata, which consists of chordates with a skull.
The Craniata and Tunicata compose the clade Olfactores. Chordates form a phylum of animals that are defined by having at some stage in their lives all of the following anatomical features: A notochord, a stiff rod of cartilage that extends along the inside of the body. Among the vertebrate sub-group of chordates the notochord develops into the spine, in wholly aquatic species this helps the animal to swim by flexing its tail. A dorsal neural tube. In fish and other vertebrates, this develops into the spinal cord, the main communications trunk of the nervous system. Pharyngeal slits; the pharynx is the part of the throat behind the mouth. In fish, the slits are modified to form gills, but in some other chordates they are part of a filter-feeding system that extracts particles of food from the water in which the animals live. Post-anal tail. A muscular tail that extends backwards behind the anus. An endostyle; this is a groove in the ventral wall of the pharynx. In filter-feeding species it produces mucus to gather food particles, which helps in transporting food to the esophagus.
It stores iodine, may be a precursor of the vertebrate thyroid gland. There are soft constraints that separate chordates from certain other biological lineages, but are not part of the formal definition: All chordates are deuterostomes; this means. All chordates are based on a bilateral body plan. All chordates are coelomates, have a fluid filled body cavity called a coelom with a complete lining called peritoneum derived from mesoderm; the following schema is from the third edition of Vertebrate Palaeontology. The invertebrate chordate classes are from Fishes of the World. While it is structured so as to reflect evolutionary relationships, it retains the traditional ranks used in Linnaean taxonomy. Phylum Chordata †Vetulicolia? Subphylum Cephalochordata – Class Leptocardii Clade Olfactores Subphylum Tunicata – Class Ascidiacea Class Thaliacea Class Appendicularia Class Sorberacea Subphylum Vertebrata Infraphylum incertae sedis Cyclostomata Superclass'Agnatha' paraphyletic Class Myxini Class Petromyzontida or Hyperoartia Class †Conodonta Class †Myllokunmingiida Class †Pteraspidomorphi Class †Thelodonti Class †Anaspida Class †Cephalaspidomorphi Infraphylum Gnathostomata Class †Placodermi Class Chondrichthyes Class †Acanthodii Superclass Osteichthyes Class Actinopterygii Class Sarcopterygii Superclass Tetrapoda Class Amphibia Class Sauropsida Class Synapsida Craniates, one of the three subdivisions of chordates, all have distinct skulls.
They include the hagfish. Michael J. Benton commented that "craniates are characterized by their heads, just as chordates, or all deuterostomes, are by their tails". Most craniates are vertebrates; these consist of a series of bony or cartilaginous cylindrical vertebrae with neural arches that protect the spinal cord, with projections that link the vertebrae. However hagfish have incomplete braincases and no vertebrae, are therefore not regarded as vertebrates, but as members of the craniates, the group from which vertebrates are thought to have evolved; however the cladistic exclusion of hagfish from the vertebrates is controversial, as they ma