A primate is a eutherian mammal constituting the taxonomic order Primates. Primates arose 85–55 million years ago from small terrestrial mammals, which adapted to living in the trees of tropical forests: many primate characteristics represent adaptations to life in this challenging environment, including large brains, visual acuity, color vision, altered shoulder girdle, dexterous hands. Primates range in size from Madame Berthe's mouse lemur, which weighs 30 g, to the eastern gorilla, weighing over 200 kg. There are 190 -- 448 species of living primates, depending on. New primate species continue to be discovered: over 25 species were described in the first decade of the 2000s, eleven since 2010. Primates are divided into two distinct suborders; the first is the strepsirrhines - lemurs and lorisids. The second is haplorhines - the "dry-nosed" primates - tarsier and ape clades, the last of these including humans. Simians are monkeys and apes, cladistically including: the catarrhines consisting of the Old World monkeys and apes.
Forty million years ago, simians from Africa migrated to South America by drifting on debris, gave rise to the New World monkeys. Twenty five million years ago the remaining Old World simians split into Old World monkeys. Common names for the simians are the baboons, macaques and great apes. Primates have large brains compared to other mammals, as well as an increased reliance on visual acuity at the expense of the sense of smell, the dominant sensory system in most mammals; these features are more developed in monkeys and apes, noticeably less so in lorises and lemurs. Some primates are trichromats, with three independent channels for conveying color information. Except for apes, primates have tails. Most primates have opposable thumbs. Many species are sexually dimorphic. Primates have slower rates of development than other sized mammals, reach maturity and have longer lifespans. Depending on the species, adults may live in solitude, in mated pairs, or in groups of up to hundreds of members; some primates, including gorillas and baboons, are terrestrial rather than arboreal, but all species have adaptations for climbing trees.
Arboreal locomotion techniques used include leaping from tree to tree and swinging between branches of trees. Primates are among the most social of animals, forming pairs or family groups, uni-male harems, multi-male/multi-female groups. Non-human primates have at four types of social systems, many defined by the amount of movement by adolescent females between groups. Most primate species remain at least arboreal: the exceptions are some great apes and humans, who left the trees for the ground and now inhabit every continent. Close interactions between humans and non-human primates can create opportunities for the transmission of zoonotic diseases virus diseases, including herpes, ebola and hepatitis. Thousands of non-human primates are used in research around the world because of their psychological and physiological similarity to humans. About 60% of primate species are threatened with extinction. Common threats include deforestation, forest fragmentation, monkey drives, primate hunting for use in medicines, as pets, for food.
Large-scale tropical forest clearing for agriculture most threatens primates. The English name "primates" is derived from Old French or French primat, from a noun use of Latin primat-, from primus; the name was given by Carl Linnaeus. The relationships among the different groups of primates were not understood until recently, so the used terms are somewhat confused. For example, "ape" has been used either as an alternative for "monkey" or for any tailless human-like primate. Sir Wilfrid Le Gros Clark was one of the primatologists who developed the idea of trends in primate evolution and the methodology of arranging the living members of an order into an "ascending series" leading to humans. Used names for groups of primates such as "prosimians", "monkeys", "lesser apes", "great apes" reflect this methodology. According to our current understanding of the evolutionary history of the primates, several of these groups are paraphyletic: a paraphyletic group is one which does not include all the descendants of the group's common ancestor.
In contrast with Clark's methodology, modern classifications identify only those groupings that are monophyletic. The cladogram below shows one possible classification sequence of the living primates: groups that use common names are shown on the right. All groups with scientific names are monophyletic, the sequence of scientific classification reflects the evolution
The public domain consists of all the creative works to which no exclusive intellectual property rights apply. Those rights may have been forfeited, expressly waived, or may be inapplicable; the works of William Shakespeare and Beethoven, most early silent films, are in the public domain either by virtue of their having been created before copyright existed, or by their copyright term having expired. Some works are not covered by copyright, are therefore in the public domain—among them the formulae of Newtonian physics, cooking recipes, all computer software created prior to 1974. Other works are dedicated by their authors to the public domain; the term public domain is not applied to situations where the creator of a work retains residual rights, in which case use of the work is referred to as "under license" or "with permission". As rights vary by country and jurisdiction, a work may be subject to rights in one country and be in the public domain in another; some rights depend on registrations on a country-by-country basis, the absence of registration in a particular country, if required, gives rise to public-domain status for a work in that country.
The term public domain may be interchangeably used with other imprecise or undefined terms such as the "public sphere" or "commons", including concepts such as the "commons of the mind", the "intellectual commons", the "information commons". Although the term "domain" did not come into use until the mid-18th century, the concept "can be traced back to the ancient Roman Law, as a preset system included in the property right system." The Romans had a large proprietary rights system where they defined "many things that cannot be owned" as res nullius, res communes, res publicae and res universitatis. The term res nullius was defined as things not yet appropriated; the term res communes was defined as "things that could be enjoyed by mankind, such as air and ocean." The term res publicae referred to things that were shared by all citizens, the term res universitatis meant things that were owned by the municipalities of Rome. When looking at it from a historical perspective, one could say the construction of the idea of "public domain" sprouted from the concepts of res communes, res publicae, res universitatis in early Roman law.
When the first early copyright law was first established in Britain with the Statute of Anne in 1710, public domain did not appear. However, similar concepts were developed by French jurists in the 18th century. Instead of "public domain", they used terms such as publici juris or propriété publique to describe works that were not covered by copyright law; the phrase "fall in the public domain" can be traced to mid-19th century France to describe the end of copyright term. The French poet Alfred de Vigny equated the expiration of copyright with a work falling "into the sink hole of public domain" and if the public domain receives any attention from intellectual property lawyers it is still treated as little more than that, left when intellectual property rights, such as copyright and trademarks, expire or are abandoned. In this historical context Paul Torremans describes copyright as a, "little coral reef of private right jutting up from the ocean of the public domain." Copyright law differs by country, the American legal scholar Pamela Samuelson has described the public domain as being "different sizes at different times in different countries".
Definitions of the boundaries of the public domain in relation to copyright, or intellectual property more regard the public domain as a negative space. According to James Boyle this definition underlines common usage of the term public domain and equates the public domain to public property and works in copyright to private property. However, the usage of the term public domain can be more granular, including for example uses of works in copyright permitted by copyright exceptions; such a definition regards work in copyright as private property subject to fair-use rights and limitation on ownership. A conceptual definition comes from Lange, who focused on what the public domain should be: "it should be a place of sanctuary for individual creative expression, a sanctuary conferring affirmative protection against the forces of private appropriation that threatened such expression". Patterson and Lindberg described the public domain not as a "territory", but rather as a concept: "here are certain materials – the air we breathe, rain, life, thoughts, ideas, numbers – not subject to private ownership.
The materials that compose our cultural heritage must be free for all living to use no less than matter necessary for biological survival." The term public domain may be interchangeably used with other imprecise or undefined terms such as the "public sphere" or "commons", including concepts such as the "commons of the mind", the "intellectual commons", the "information commons". A public-domain book is a book with no copyright, a book, created without a license, or a book where its copyrights expired or have been forfeited. In most countries the term of protection of copyright lasts until January first, 70 years after the death of the latest living author; the longest copyright term is in Mexico, which has life plus 100 years for all deaths since July 1928. A notable exception is the United States, where every book and tale published prior to 1924 is in the public domain.
The Hominidae, whose members are known as great apes or hominids, are a taxonomic family of primates that includes eight extant species in four genera: Pongo, the Bornean and Tapanuli orangutan. Several revisions in classifying the great apes have caused the use of the term "hominid" to vary over time, its original meaning referred only to their closest extinct relatives. That restrictive meaning has now been assumed by the term "hominin", which comprises all members of the human clade after the split from the chimpanzees; the current, 21st-century meaning of "hominid" includes all the great apes including humans. Usage still varies and some scientists and laypersons still use "hominid" in the original restrictive sense. Within the taxon Hominidae, a number of extant and known extinct, that is, genera are grouped with the humans and gorillas in the subfamily Homininae; the most recent common ancestor of all Hominidae lived 14 million years ago, when the ancestors of the orangutans speciated from the ancestral line of the other three genera.
Those ancestors of the family Hominidae had speciated from the family Hylobatidae 15 million to 20 million years ago. In the early Miocene, about 22 million years ago, there were many species of arboreally adapted primitive catarrhines from East Africa. Fossils at 20 million years ago include fragments attributed to Victoriapithecus, the earliest Old World monkey. Among the genera thought to be in the ape lineage leading up to 13 million years ago are Proconsul, Dendropithecus, Nacholapithecus, Nyanzapithecus, Afropithecus and Kenyapithecus, all from East Africa. At sites far distant from East Africa, the presence of other generalized non-cercopithecids, that is, non-monkey primates, of middle Miocene age—Otavipithecus from cave deposits in Namibia, Pierolapithecus and Dryopithecus from France and Austria—is further evidence of a wide diversity of ancestral ape forms across Africa and the Mediterranean basin during the warm and equable climatic regimes of the early and middle Miocene; the most recent of these far-flung Miocene apes is Oreopithecus, from the fossil-rich coal beds in northern Italy and dated to 9 million years ago.
Molecular evidence indicates that the lineage of gibbons, the lesser apes, diverged from that of the great apes some 18–12 million years ago, that of orangutans diverged from the other great apes at about 12 million years. There are no fossils that document the ancestry of gibbons, which may have originated in a still-unknown South East Asian hominoid population. Species close to the last common ancestor of gorillas and humans may be represented by Nakalipithecus fossils found in Kenya and Ouranopithecus found in Greece. Molecular evidence suggests that between 8 and 4 million years ago, first the gorillas, the chimpanzees split off from the line leading to the humans. Human DNA is 98.4% identical to that of chimpanzees when comparing single nucleotide polymorphisms. The fossil record, however, of gorillas and chimpanzees is limited. Other hominins adapted to the drier environments outside the African equatorial belt; the wet equatorial belt contracted after about 8 million years ago, there is little fossil evidence for the divergence of the hominin lineage from that of gorillas and chimpanzees—which split was thought to have occurred around that time.
The earliest fossils argued by some to belong to the human lineage are Sahelanthropus tchadensis and Orrorin tugenensis, followed by Ardipithecus, with species Ar. kadabba and Ar. ramidus. The classification of the great apes has been revised several times in the last few decades; the original meaning of the term referred to only humans and their closest relatives—what is now the modern meaning of the term "hominin". The meaning of the taxon Hominidae changed leading to a different usage of "hominid" that today includes all the great apes including humans; the term hominid is confused with a number of similar words: A hominoid called an ape, is a member of the superfamily Hominoidea: extant members are the gibbons and the hominids. A hominid is a member of the family Hominidae, the great apes: orangutans, gorillas and humans. A hominine is a member of the subfamily Homininae: gorillas and humans. A hominin is a member of the tribe Hominini: humans. A homininan, following a suggestion by Wood and Richmond, would be a member of the subtribe Homin
Neanderthals are an extinct species or subspecies of archaic humans in the genus Homo, who lived within Eurasia from circa 400,000 until 40,000 years ago. The earliest fossils of Neanderthals in Europe are dated between 450,000 and 430,000 years ago, thereafter Neanderthals expanded into Southwest and Central Asia, they are known from numerous fossils, as well as stone tool assemblages. All assemblages younger than 160,000 years are of the so-called Mousterian techno-complex, characterised by tools made out of stone flakes; the type specimen is Neanderthal 1, found in Neander Valley in the German Rhineland, in 1856. Compared to modern humans, Neanderthals were stockier, with bigger bodies. In conformance with Bergmann's rule, as well as Allen's rule, this was was an adaptation to preserve heat in cold climates. Male and female Neanderthals had cranial capacities averaging 1,600 cm3 and 1,300 cm3 within the range of the values for anatomically modern humans. Average males stood around females 152 to 156 cm tall.
There has been growing evidence for admixture between Neanderthals and anatomically modern humans, reflected in the genomes of all modern non-African populations but not in the genomes of most sub-Saharan Africans. This suggests that interbreeding between Neanderthals and anatomically modern humans took place after the recent "out of Africa" migration, around 70,000 years ago. Recent admixture analyses have added to the complexity, finding that Eastern Neanderthals derived up to 2% of their ancestry from anatomically modern humans who left Africa some 100,000 years ago. Neanderthals are named after one of the first sites where their fossils were discovered in the mid-19th century in the Neander Valley, just east of Düsseldorf, at the time in the Rhine Province of the Kingdom of Prussia; the valley itself was named for Joachim Neander, Neander being the graecicized form of the surname Neumann. The German spelling of Thal "Valley" was current in the 19th century. Neanderthal 1 was known as the "Neanderthal cranium" or "Neanderthal skull" in anthropological literature, the individual reconstructed on the basis of the skull was called "the Neanderthal man".
The binomial name Homo neanderthalensis—extending the name "Neanderthal man" from the individual type specimen to the entire group—was first proposed by the Anglo-Irish geologist William King in a paper read to the British Association in 1863, although in the following year he stated that the specimen was not human and rejected the name. King's name had priority over the proposal put forward in 1866 by Homo stupidus. Popular English usage of "Neanderthal" as shorthand for "Neanderthal man", as in "the Neanderthals" or "a Neanderthal", emerged in the popular literature of the 1920s. Since the historical spelling -th- in German represents the phoneme /t/ or /tʰ/, not the fricative /θ/, standard British pronunciation of "Neanderthal" is with /t/; because of the usual sound represented by digraph ⟨th⟩ in English, "Neanderthal" is pronounced with the voiceless fricative /θ/, at least in "layman's American English". The spelling Neandertal is seen in English in scientific publications. Since "Neanderthal", or "Neandertal", is a common name, there is no authoritative prescription on its spelling, unlike the spelling of the binominal name H. neanderthalensis, predicated by King 1864.
The common name in German is always invariably Neandertaler, not Neandertal, but the spelling of the name of the Neander Valley itself has been affected by the species name, the names of the Neanderthal Museum and of Neanderthal station persisting with pre-1900 orthography. Since the discovery of the Neanderthal fossils, expert opinion has been divided as to whether Neanderthals should be considered a separate species or a subspecies relative to modern humans. Pääbo described such "taxonomic wars" as unresolveable in principle, "since there is no definition of species describing the case." The question depends on the definition of Homo sapiens as a chronospecies, in flux throughout the 20th century. Authorities preferring classification of Neanderthals as subspecies have introduced the subspecies name Homo sapiens sapiens for the anatomically modern Cro-Magnon population which lived in Europe at the same time as Neanderthals, while authorities preferring classification as separate species use Homo sapiens as equivalent to "anatomically modern humans".
During the early 20th century, a prevailing view of Neanderthals as "simian", influenced by Arthur Keith and Marcellin Boule, tended to exaggerate the anatomical differences between Neanderthals and Cro Magnon. Beginning in the 1930s, revised reconstructions of Neanderthals emphasized the similarity rather than differences from modern humans. From the 1940s throughout the 1970s, it was common to use the subspecies classification of Homo sapiens neanderthalensis vs. Homo sapiens sapiens; the hypothesis of "multiregional origin" of modern man was formulated in the 1980s on such grounds, arguing for the presence of an unbroken succession of fossil sites in both Europe and Asia. Hybridization between Neanderthals and Cro Magnon had been suggested on skeletal and craniological grounds since the early 20th century, found increasing support in the 20th century, until Neanderthal admixture was found to be present in modern populations genet
A number of varieties of Homo are grouped into the broad category of archaic humans in the period contemporary to and predating the emergence of the earliest anatomically modern humans over 315 ka. The term includes Homo neanderthalensis, Homo rhodesiensis, Homo heidelbergensis, Homo antecessor. There is no universal consensus on this terminology, varieties of "archaic humans" are included under the binomial name of either Homo sapiens or Homo erectus by some authors. Archaic humans had a brain size averaging 1,200 to 1,400 cubic centimeters, which overlaps with the range of modern humans. Archaics are distinguished from anatomically modern humans by having a thick skull, prominent supraorbital ridges and the lack of a prominent chin. Anatomically modern humans appear from over 160,000 years ago in Ethiopia and after 70,000 years ago supplanting the "archaic" human varieties. Non-modern varieties of Homo are certain to have survived until after 30,000 years ago, until as as 12,000 years ago.
Which of these, if any, are included under the term "archaic human" is a matter of definition and varies among authors. Nonetheless, according to recent genetic studies, modern humans may have bred with "at least two groups" of ancient humans: Neanderthals and Denisovans. Other studies have cast doubt on admixture being the source of the shared genetic markers between archaic and modern humans, pointing to an ancestral origin of the traits which originated 500,000–800,000 years ago. Another group may have been extant as as 11,500 years ago, the Red Deer Cave people of China. Chris Stringer of the Natural History Museum in London has suggested that these people could be a result of mating between Denisovans and modern humans. Other scientists remain skeptical, suggesting that the unique features are within the variations expected for modern human populations; the category archaic human lacks a single, agreed upon definition. According to one definition, Homo sapiens is a single species comprising several subspecies that include the archaics and modern humans.
Under this definition, modern humans are referred to as Homo sapiens sapiens and archaics are designated with the prefix "Homo sapiens". For example, the Neanderthals are Homo sapiens neanderthalensis, Homo heidelbergensis is Homo sapiens heidelbergensis. Other taxonomists prefer not to consider archaics and modern humans as a single species but as several different species. In this case the standard taxonomy is used, Homo neanderthalensis; the evolutionary dividing lines that separate modern humans from archaic humans and archaic humans from Homo erectus are unclear. The earliest known fossils of anatomically modern humans such as the Omo remains from 195,000 years ago, Homo sapiens idaltu from 160,000 years ago, Qafzeh remains from 90,000 years ago are recognizably modern humans. However, these early modern humans do possess a number of archaic traits, such as moderate, but not prominent, brow ridges; the emergence of archaic humans is sometimes used as an example of punctuated equilibrium.
This occurs when a species undergoes significant biological evolution within a short period. Subsequently, the species undergoes little change for long periods until the next punctuation; the brain size of archaic humans expanded from 900 cm3 in erectus to 1,300 cm3. Since the peak of human brain size during the archaics, it has begun to decline. Robin Dunbar has argued. Based on his analysis of the relationship between brain size and hominin group size, he concluded that because archaic humans had large brains, they must have lived in groups of over 120 individuals. Dunbar argues that it was not possible for hominins to live in such large groups without using language, otherwise there could be no group cohesion and the group would disintegrate. By comparison, chimpanzees live in smaller groups of up to 50 individuals. Atapuerca Mountains, Sima de los Huesos Saldanha Man Altamura Man Kabwe Skull Steinheim Skull Ndutu cranium Early and Late "Archaic" Homo Sapiens and "Anatomically Modern" Homo Sapiens Origins of Modern Humans: Multiregional or Out of Africa?
Homo sapiens, Museum of Natural History Human Timeline – Smithsonian, National Museum of Natural History
Jebel Irhoud is an archaeological site located just north of the locality known as Tlet Ighoud, about 50 km south-east of the city of Safi in Morocco. It is noted for the hominin fossils that have been found there since the site's discovery in 1960. Thought to be Neanderthals, the specimens have since been assigned to Homo sapiens and have been dated at 315,000 years old; the site is the remnants of a solutional cave filled with 8 meters of deposits from the Pleistocene era, located on the eastern side of a karstic outcrop of limestone at an elevation of 562 meters. It was discovered in 1960. A miner discovered a skull in the wall of the cave, extracted it and gave it to an engineer, who kept it as a souvenir for a time, it was handed over to the University of Rabat, who organized a joint French-Moroccan expedition to the site in 1961, headed by the French researcher Émile Ennouchi. Ennouchi's team identified the remains of around 30 species of mammals, some of which are associated with the Middle Pleistocene, but the stratigraphic provenance is unknown.
Another excavation was carried out by Jacques Tixier and Roger de Bayle des Hermens in 1967 and 1969 in which 22 layers were identified in the cave. The lower 13 layers were found to contain signs of human habitation including an industry classified as Levallois Mousterian; the site is noted for the hominin fossils found there. Ennouchi discovered a skull which he termed Irhoud 1 and is now on display in the Rabat Archaeological Museum, he discovered part of another skull, designated Irhoud 2, the following year and subsequently uncovered the lower mandible of a child, designated Irhoud 3. Tixier's excavation found 1,267 recorded objects including skulls, a humerus designated Irhoud 4 and a hip bone recorded as Irhoud 5. Further excavations were carried out by American researchers in the 1990s and by a team led by Jean-Jacques Hublin from 2004. Animal remains, it was quite different to the present and represented a dry and steppe-like environment roamed by equids, gazelles and various predators.
The finds were interpreted as Neanderthal, as the stone tools found with them were believed to be associated with Neanderthals. They had archaic features believed to be representative of the Neanderthals, rather than Homo sapiens, they were thought to be around 40,000 years old, but this was thrown into doubt by faunal evidence suggesting a Middle Pleistocene date, around 160,000 years ago. The fossils were reappraised as representing an archaic form of Homo sapiens or a population of Homo sapiens that had interbred with Neanderthals; this was consistent with the idea that the oldest known remains of a Homo sapiens idaltu, dated to around 195,000 years ago and found in Omo Kibish, indicated an eastern African origin for humans at around 200,000 years ago. However, dating carried out by the Max Planck Institute for Evolutionary Anthropology in Leipzig revealed that the Jebel Irhoud site was far older than first thought. Fresh excavations carried out in 2004 by the Hublin team revealed more than 20 new bones from the remains of at least five people, a number of stone tools.
The finds included part of a skull, a jawbone and limb bones which had come from three adults, a juvenile, a child aged about seven and a half years old. The bones looked similar facially to those of humans today but had much larger lower jaws and elongated braincases, they have similar features to the Florisbad Skull dating to 260,000 years ago found at the other end of the continent, in Florisbad, South Africa, attributed to Homo sapiens on the basis of the Jebel Irhoud finds. The tools were found alongside gazelle bones and lumps of charcoal, indicating the presence of fire and of cooking in the cave; the gazelle bones showed characteristic signs of butchery and cooking, such as cut marks, notches consistent with marrow extraction, charring. Some of the tools had been burned due to fires being lit on top of them after they had been discarded; this enabled the researchers to use thermoluminescence dating to ascertain when the burning had happened, by proxy the age of the fossil bones, which were found in the same deposit layer.
The burnt tools were dated to around 315,000 years ago, indicating that the fossils are of about the same age. This conclusion was confirmed by recalculating the age of the Irhoud 3 mandible, which produced an age range compatible with that of the tools at 280,000 to 350,000 years old. If they hold up, these dates would make the remains by far the earliest known examples of Homo sapiens; this suggests that, rather than arising in East Africa around 200,000 years ago, modern humans may have been present across the length of Africa 100,000 years earlier. According to study author Jean-Jacques Hublin, "The idea is that early Homo sapiens dispersed around the continent and elements of human modernity appeared in different places, so different parts of Africa contributed to the emergence of what we call modern humans today." Early humans may have comprised a large, interbreeding population dispersed across Africa whose spread was facilitated by a wetter climate that created a "green Sahara", around 330,000 to 300,000 years ago.
The rise of modern humans may thus have taken place on a continental scale rather than being confined to a particular corner of Africa. Hublin and his team attempted to obtain a DNA sample from these fossils, but these attempts were unsuccessful. Genomic analysis would have provided necess