The eudicots, Eudicotidae or eudicotyledons are a clade of flowering plants, called tricolpates or non-magnoliid dicots by previous authors. The botanical terms were introduced in 1991 by evolutionary botanist James A. Doyle and paleobotanist Carol L. Hotton to emphasize the evolutionary divergence of tricolpate dicots from earlier, less specialized, dicots; the close relationships among flowering plants with tricolpate pollen grains was seen in morphological studies of shared derived characters. These plants have a distinct trait in their pollen grains of exhibiting three colpi or grooves paralleling the polar axis. Molecular evidence confirmed the genetic basis for the evolutionary relationships among flowering plants with tricolpate pollen grains and dicotyledonous traits; the term means "true dicotyledons", as it contains the majority of plants that have been considered dicots and have characteristics of the dicots. The term "eudicots" has subsequently been adopted in botany to refer to one of the two largest clades of angiosperms, monocots being the other.
The remaining angiosperms include magnoliids and what are sometimes referred to as basal angiosperms or paleodicots, but these terms have not been or adopted, as they do not refer to a monophyletic group. The other name for the eudicots is tricolpates, a name which refers to the grooved structure of the pollen. Members of the group have tricolpate pollen; these pollens have three or more pores set in furrows called colpi. In contrast, most of the other seed plants produce monosulcate pollen, with a single pore set in a differently oriented groove called the sulcus; the name "tricolpates" is preferred by some botanists to avoid confusion with the dicots, a nonmonophyletic group. Numerous familiar plants are eudicots, including many common food plants and ornamentals; some common and familiar eudicots include members of the sunflower family such as the common dandelion, the forget-me-not and other members of its family, buttercup and macadamia. Most leafy trees of midlatitudes belong to eudicots, with notable exceptions being magnolias and tulip trees which belong to magnoliids, Ginkgo biloba, not an angiosperm.
The name "eudicots" is used in the APG system, of 1998, APG II system, of 2003, for classification of angiosperms. It is applied to a monophyletic group, which includes most of the dicots. "Tricolpate" is a synonym for the "Eudicot" monophyletic group, the "true dicotyledons". The number of pollen grain furrows or pores helps classify the flowering plants, with eudicots having three colpi, other groups having one sulcus. Pollen apertures are any modification of the wall of the pollen grain; these modifications include thinning and pores, they serve as an exit for the pollen contents and allow shrinking and swelling of the grain caused by changes in moisture content. The elongated apertures/ furrows in the pollen grain are called colpi, along with pores, are a chief criterion for identifying the pollen classes; the eudicots can be divided into two groups: the basal eudicots and the core eudicots. Basal eudicot is an informal name for a paraphyletic group; the core eudicots are a monophyletic group.
A 2010 study suggested the core eudicots can be divided into two clades, Gunnerales and a clade called "Pentapetalae", comprising all the remaining core eudicots. The Pentapetalae can be divided into three clades: Dilleniales superrosids consisting of Saxifragales and rosids superasterids consisting of Santalales, Berberidopsidales and asteridsThis division of the eudicots is shown in the following cladogram: The following is a more detailed breakdown according to APG IV, showing within each clade and orders: clade Eudicots order Ranunculales order Proteales order Trochodendrales order Buxales clade Core eudicots order Gunnerales order Dilleniales clade Superrosids order Saxifragales clade Rosids order Vitales clade Fabids order Fabales order Rosales order Fagales order Cucurbitales order Oxalidales order Malpighiales order Celastrales order Zygophyllales clade Malvids order Geraniales order Myrtales order Crossosomatales order Picramniales order Malvales order Brassicales order Huerteales order Sapindales clade Superasterids order Berberidopsidales order Santalales order Caryophyllales clade Asterids order Cornales order Ericales clade Campanulids order Aquifoliales order Asterales order Escalloniales order Bruniales order Apiales order Dipsacales order Paracryphiales clade Lamiids order Solanales order Lamiales order Vahliales order Gentianales order Boraginales order Garryales order Metteniusales order Icacinales Eudicots at the Encyclopedia of Life Eudicots, Tree of Life Web Project Dicots Plant Life Forms
In the APG IV system for the classification of flowering plants, the name asterids denotes a clade. Common examples include the forget-me-nots, the common sunflower, morning glory and sweet potato, lavender, olive, honeysuckle, ash tree, snapdragon, psyllium, garden sage, table herbs such as mint and rosemary, rainforest trees such as Brazil nut. Most of the taxa belonging to this clade had been referred to the Asteridae in the Cronquist system and to the Sympetalae in earlier systems; the name asterids resembles the earlier botanical name but is intended to be the name of a clade rather than a formal ranked name, in the sense of the ICBN. The phylogenetic tree presented hereafter has been proposed by the APG IV project. Genetic analysis carried out after APG II maintains that the sister to all other asterids are the Cornales. A second order that split from the base of the asterids are the Ericales; the remaining orders cluster into two clades, the lamiids and the campanulids. The structure of both of these clades has changed in APG III.
In APG III system, the following clades were renamed: euasterids I → lamiids euasterids II → campanulids Asterids in Stevens, P. F.. Angiosperm Phylogeny Website. Version 7, May 2006
The flowering plants known as angiosperms, Angiospermae or Magnoliophyta, are the most diverse group of land plants, with 64 orders, 416 families 13,164 known genera and c. 369,000 known species. Like gymnosperms, angiosperms are seed-producing plants. However, they are distinguished from gymnosperms by characteristics including flowers, endosperm within the seeds, the production of fruits that contain the seeds. Etymologically, angiosperm means a plant; the term comes from the Greek words sperma. The ancestors of flowering plants diverged from gymnosperms in the Triassic Period, 245 to 202 million years ago, the first flowering plants are known from 160 mya, they diversified extensively during the Early Cretaceous, became widespread by 120 mya, replaced conifers as the dominant trees from 100 to 60 mya. Angiosperms differ from other seed plants in several ways, described in the table below; these distinguishing characteristics taken together have made the angiosperms the most diverse and numerous land plants and the most commercially important group to humans.
Angiosperm stems are made up of seven layers. The amount and complexity of tissue-formation in flowering plants exceeds that of gymnosperms; the vascular bundles of the stem are arranged such that the phloem form concentric rings. In the dicotyledons, the bundles in the young stem are arranged in an open ring, separating a central pith from an outer cortex. In each bundle, separating the xylem and phloem, is a layer of meristem or active formative tissue known as cambium. By the formation of a layer of cambium between the bundles, a complete ring is formed, a regular periodical increase in thickness results from the development of xylem on the inside and phloem on the outside; the soft phloem becomes crushed, but the hard wood persists and forms the bulk of the stem and branches of the woody perennial. Owing to differences in the character of the elements produced at the beginning and end of the season, the wood is marked out in transverse section into concentric rings, one for each season of growth, called annual rings.
Among the monocotyledons, the bundles are more numerous in the young stem and are scattered through the ground tissue. They once formed the stem increases in diameter only in exceptional cases; the characteristic feature of angiosperms is the flower. Flowers show remarkable variation in form and elaboration, provide the most trustworthy external characteristics for establishing relationships among angiosperm species; the function of the flower is to ensure fertilization of the ovule and development of fruit containing seeds. The floral apparatus may arise terminally from the axil of a leaf; as in violets, a flower arises singly in the axil of an ordinary foliage-leaf. More the flower-bearing portion of the plant is distinguished from the foliage-bearing or vegetative portion, forms a more or less elaborate branch-system called an inflorescence. There are two kinds of reproductive cells produced by flowers. Microspores, which will divide to become pollen grains, are the "male" cells and are borne in the stamens.
The "female" cells called megaspores, which will divide to become the egg cell, are contained in the ovule and enclosed in the carpel. The flower may consist only of these parts, as in willow, where each flower comprises only a few stamens or two carpels. Other structures are present and serve to protect the sporophylls and to form an envelope attractive to pollinators; the individual members of these surrounding structures are known as petals. The outer series is green and leaf-like, functions to protect the rest of the flower the bud; the inner series is, in general, white or brightly colored, is more delicate in structure. It functions to attract bird pollinators. Attraction is effected by color and nectar, which may be secreted in some part of the flower; the characteristics that attract pollinators account for the popularity of flowers and flowering plants among humans. While the majority of flowers are perfect or hermaphrodite, flowering plants have developed numerous morphological and physiological mechanisms to reduce or prevent self-fertilization.
Heteromorphic flowers have short carpels and long stamens, or vice versa, so animal pollinators cannot transfer pollen to the pistil. Homomorphic flowers may employ a biochemical mechanism called self-incompatibility to discriminate between self and non-self pollen grains. In other species, the male and female parts are morphologically separated, developing on different flowers; the botanical term "Angiosperm", from the Ancient Greek αγγείον, angeíon and σπέρμα, was coined in the form Angiospermae by Paul Hermann in 1690, as the name of one of his primary divisions of the plant kingdom. This included flowering plants possessing seeds enclosed in capsules, distinguished from his Gymnospermae, or flowering plants with achenial or schizo-carpic fruits, the whole fruit or each of its pieces being here regarded as a seed and naked; the term and its antonym were maintained by Carl Linnaeus with the same sense, but with restricted application, in the names of the orders of his class Didynamia. Its use with any
Calyceraceae is a plant family in the order Asterales. The natural distribution of the about sixty species belonging to this family is restricted to the southern half of South-America; the species of the family resemble both the Dipsacaceae. Calyceraceae are annual herbs. There may be a many branched stems that may be without hair or with soft silky hairs; the leaves may set alternately along the stems. Stipules are lacking; the leafblade may be lobed to pinnatisect. The margin of the leaves may be toothed; the inflorescences are flowerheads comparable to those in the sunflower family. They are at the top of the stems or opposite leaves, may have a flowerstem or be seated, while each flowerhead may be on its own or in a cyme; each individual flowerhead is surrounded by an involucre, consisting of one or two rows of bracts that are leaf-like and not merged. The base of the flowerhead may be conical, convex or sometimes spheroidal. On the base of the flowerhead, at the base of each individual flower, are linear to narrowly lanceolate, chaffy scales that become woody when seeds are ripening.
Each flowerhead may contain a few or up to over one hundred hermaphrodite or unisexual, star-symmetric or mirror-symmetric flowers. The petals are fused to form a funnel-shaped or sometimes cylinder-shaped corolla, split into four to six lobes at the top; the corolla's remains stay on the top of the one-seeded dry fruit at maturity. Four or five stamens alternate with the corolla lobes; the lower third of these filaments are fused with the corolla tube, while sometimes filaments may be attached to their neighbors. Filaments carry nectaries; the anthers stand upright, with pollen freed from a slit at the top. The style is thread-like without hairs, sticking out above the corolla tube, while the stigma at its tip is club-shaped or split in two; the ovary consists of two carpels with only one ovule, pendulous and anatropous. The fruit is an achene, with a persistent calyx which may consists of spines, contains one seed, only enclosed by a thin pericarp and has fleshy endosperm; the sepals may be free or fused calyx lobes, woody on the outside.
Fruits may be dispersed separately when ripe or can remain on the floral base that breaks free of the plant. Both Calyceraceae and Asteraceae have their flowers set in heads with a common floral base; the bracts surrounding the flowerhead in the Calyceraceae are leaf-like while the involucral bracts in the Asteraceae differ from the leaves. The anthers are form a tube in the Asteraceae; the filaments in the Calyceraceae are, at least in their lower third, fused to the corolla, whereas in the Asteraceae the filaments are free or connected. Both Calyceraceae and Dipsacaceae have persistent calyces, but these become lignified or spiny in Calyceraceae, but are cup-shaped or consist of a circle of hairs in the Dipsacaceae. Stamens are alternating with the corolla lobes and anthers open at their top in the Calyceraceae whereas stamens are centered on the petals and anthers open toward the middle of the flower in the Dipsacaceae. Six genera are assigned to this family: Acicarpha, Calycera, Gamocarpha and Nastanthus.
The majority of species in this family occur in Argentina, seven of which are endemics, with the highest species density south of the tropics. Calycera and Acicarpha both are widespread along the Andes from northern Argentina through to the Altiplano of Peru. Most of the thirteen species of Boopsis occur in the south of Argentina and Chile but some species are found in the tropics. Moschopsis grows in the Salta Province in Argentina. Acicarpha tribuloides occurs as an introduced weed along roads in Florida
Natural Resources Conservation Service
Natural Resources Conservation Service known as the Soil Conservation Service, is an agency of the United States Department of Agriculture that provides technical assistance to farmers and other private landowners and managers. Its name was changed in 1994 during the presidency of Bill Clinton to reflect its broader mission, it is a small agency comprising about 12,000 employees. Its mission is to improve and conserve natural resources on private lands through a cooperative partnership with state and local agencies. While its primary focus has been agricultural lands, it has made many technical contributions to soil surveying and water quality improvement. One example is the Conservation Effects Assessment Project, set up to quantify the benefits of agricultural conservation efforts promoted and supported by programs in the Farm Security and Rural Investment Act of 2002. NRCS is the leading agency in this project; the agency was founded through the efforts of Hugh Hammond Bennett, a soil conservation pioneer who worked for the Department of Agriculture from 1903 to 1952.
Bennett's motivation was based on his knowledge of the detrimental effects of soil erosion and the impacts on U. S lands. On September 13, 1933, the Soil Erosion Service was formed in the Department of the Interior, with Bennett as chief; the service was transferred to the Department of Agriculture on March 23, 1935, was shortly thereafter combined with other USDA units to form the Soil Conservation Service by the Soil Conservation and Domestic Allotment Act of 1935. The Soil Conservation Service was in charge of 500 Civilian Conservation Corps camps between 1933 and 1942; the primary purpose of these camps was erosion control. Hugh Bennett continued as chief, a position he held until his retirement in 1952. On October 20, 1994, the agency was renamed to the Natural Resources Conservation Service as part of the Federal Crop Insurance Reform and Department of Agriculture Reorganization Act of 1994. NRCS offers financial assistance to farmers and ranchers; the financial assistance is authorized by the Farm Bill, a law, renewed every five years.
The 2014 Farm Bill consolidated 23 programs into 15. NRCS offers these services to private land owners, conservation districts and other types of organizations. NRCS collects and shares information on the nation's soil, water and plants; the Conservation Title of the Farm Bill provides the funding to agricultural producers, a conservation plan must be included. All of these programs are voluntary; the main programs include: The purpose of EQIP is to provide assistance to landowners to help them improve their soil and related natural resources, including grazing lands and wildlife habitat. Conservation Stewardship Program CSP is targeted to a producers who maintain a higher level of environmental stewardship. Regional Conservation Partnership Program RCPP consolidated four programs from the prior 2008 Farm Bill, it aims at more watershed scale projects, rather than individual farms and ranches. Agricultural Conservation Easement Program ACEP was another consolidation effort of the 2014 Farm Bill, which includes the former Grasslands Reserve Program and Ranch Lands Protection Program, Wetlands Reserve Program.
ACEP includes technical and financial help to maintain or improve land for agriculture or environmental benefits. Landowners volunteer to protect forests in 30 or 10 year contracts; this program hands assisting funds to participants. The objectives of HFRP are to: Promote the recovery of endangered and threatened species under the Endangered Species Act Improve plant and animal biodiversity Enhance carbon sequestration Serves 10 states in the Midwest United States in helping to reduce Nitrate levels in soil due to runoff from fertilized farmland; the project began in 2010 and focused on the Mississippi Basin area. The main goal of the project is to implement better methods of managing water drainage from agricultural uses, in place of letting the water drain as it had done in the past. In October 2011, The National "Managing Water, Harvesting Results" Summit was held to promote the drainage techniques used in hopes of people adopting them nationwide. Includes water supply forecasts and the Surface Water Supply Index for Alaska and other Western states.
NRCS agents collect data from snowpack and mountain sites to predict spring runoff and summer streamflow amounts. These predictions are used in decision making for agriculture, wildlife management and development, several other areas; these predictions are available within the first 5 days of each month from January to June. Is a blanket program which involves conservation efforts on soil and water conservation, as well as management of agricultural wastes and general longterm sustainability. NRCS and related agencies work with landowners, communities, or developers to protect the environment. Serve to guide people to comply with acts such as the Highly Erodible Land and Conservation Compliance Provisions acts; the CTA can cover projects by state and federal governments. Is a program to assist gulf bordering states improve water quality and use sustainable methods of farming and other industry; the program will deliver up to 50 million dollars over 2011-2013 to apply these sustainable methods, as well as wildlife habitat management systems that do not hinder agricultural productivity, prevent future over use of water resources to protect native endangered spe
Antonio José Cavanilles
Antonio José Cavanilles was a leading Spanish taxonomic botanist of the 18th century. He named many plants from Oceania, he named at least 100 genera, about 54 of which were still used in 2004, including Dahlia, Cobaea and Oleandra. Cavanilles was born in Valencia, he lived in Paris from 1777 to 1781, where he followed careers as a clergyman and a botanist, thanks to André Thouin and Antoine Laurent de Jussieu. He was one of the first Spanish scientists to use the classification method invented by Carl Linnaeus. From Paris he moved to Madrid, where he was director of the Royal Botanical Garden and Professor of botany from 1801 to 1804, he died in Madrid in 1804. Icones et descriptiones plantarum, quae aut sponte in Hispania crescunt, aut in hortis hospitantur... Madrid, 1791-1801 List of plants of Caatinga vegetation of Brazil List of plants of Cerrado vegetation of Brazil List of Roman Catholic scientist-clerics "Cavanilles, Antonio José (1745-1804". JSTOR Global Plants. ITHAKA. 2013. Biography by the Australian National Botanic Gardens Malpighiaceae/Cavanilles Monadelphiæ classis dissertationes decem on the Internet Archive Chronophobia Scans of 160 plates from Monadelphiæ classis dissertationes decem Antonio José Cavanilles.
Polymath Virtual Library, Fundación Ignacio Larramendi
Plants are multicellular, predominantly photosynthetic eukaryotes of the kingdom Plantae. Plants were treated as one of two kingdoms including all living things that were not animals, all algae and fungi were treated as plants. However, all current definitions of Plantae exclude the fungi and some algae, as well as the prokaryotes. By one definition, plants form the clade Viridiplantae, a group that includes the flowering plants and other gymnosperms and their allies, liverworts and the green algae, but excludes the red and brown algae. Green plants obtain most of their energy from sunlight via photosynthesis by primary chloroplasts that are derived from endosymbiosis with cyanobacteria, their chloroplasts contain b, which gives them their green color. Some plants are parasitic or mycotrophic and have lost the ability to produce normal amounts of chlorophyll or to photosynthesize. Plants are characterized by sexual reproduction and alternation of generations, although asexual reproduction is common.
There are about 320 thousand species of plants, of which the great majority, some 260–290 thousand, are seed plants. Green plants provide a substantial proportion of the world's molecular oxygen and are the basis of most of Earth's ecosystems on land. Plants that produce grain and vegetables form humankind's basic foods, have been domesticated for millennia. Plants have many cultural and other uses, as ornaments, building materials, writing material and, in great variety, they have been the source of medicines and psychoactive drugs; the scientific study of plants is known as a branch of biology. All living things were traditionally placed into one of two groups and animals; this classification may date from Aristotle, who made the distincton between plants, which do not move, animals, which are mobile to catch their food. Much when Linnaeus created the basis of the modern system of scientific classification, these two groups became the kingdoms Vegetabilia and Animalia. Since it has become clear that the plant kingdom as defined included several unrelated groups, the fungi and several groups of algae were removed to new kingdoms.
However, these organisms are still considered plants in popular contexts. The term "plant" implies the possession of the following traits multicellularity, possession of cell walls containing cellulose and the ability to carry out photosynthesis with primary chloroplasts; when the name Plantae or plant is applied to a specific group of organisms or taxon, it refers to one of four concepts. From least to most inclusive, these four groupings are: Another way of looking at the relationships between the different groups that have been called "plants" is through a cladogram, which shows their evolutionary relationships; these are not yet settled, but one accepted relationship between the three groups described above is shown below. Those which have been called "plants" are in bold; the way in which the groups of green algae are combined and named varies between authors. Algae comprise several different groups of organisms which produce food by photosynthesis and thus have traditionally been included in the plant kingdom.
The seaweeds range from large multicellular algae to single-celled organisms and are classified into three groups, the green algae, red algae and brown algae. There is good evidence that the brown algae evolved independently from the others, from non-photosynthetic ancestors that formed endosymbiotic relationships with red algae rather than from cyanobacteria, they are no longer classified as plants as defined here; the Viridiplantae, the green plants – green algae and land plants – form a clade, a group consisting of all the descendants of a common ancestor. With a few exceptions, the green plants have the following features in common, they undergo closed mitosis without centrioles, have mitochondria with flat cristae. The chloroplasts of green plants are surrounded by two membranes, suggesting they originated directly from endosymbiotic cyanobacteria. Two additional groups, the Rhodophyta and Glaucophyta have primary chloroplasts that appear to be derived directly from endosymbiotic cyanobacteria, although they differ from Viridiplantae in the pigments which are used in photosynthesis and so are different in colour.
These groups differ from green plants in that the storage polysaccharide is floridean starch and is stored in the cytoplasm rather than in the plastids. They appear to have had a common origin with Viridiplantae and the three groups form the clade Archaeplastida, whose name implies that their chloroplasts were derived from a single ancient endosymbiotic event; this is the broadest modern definition of the term'plant'. In contrast, most other algae not only have different pigments but have chloroplasts with three or four surrounding membranes, they are not close relatives of the Archaeplastida having acquired chloroplasts separately from ingested or symbiotic green and red algae. They are thus not included in the broadest modern definition of the plant kingdom, although they were in the past; the green plants or Viridiplantae were traditionally divided into the green algae (including