A lichen is a composite organism that arises from algae or cyanobacteria living among filaments of multiple fungi species in a mutualistic relationship. The combined lichen has properties different from those of its component organisms. Lichens come in many colors and forms; the properties are sometimes plant-like, but lichens are not plants. Lichens may have tiny, leafless branches, flat leaf-like structures, flakes that lie on the surface like peeling paint, a powder-like appearance, or other growth forms. A macrolichen is a lichen, either bush-like or leafy. Here, "macro" and "micro" do not refer to size, but to the growth form. Common names for lichens may contain the word moss, lichens may superficially look like and grow with mosses, but lichens are not related to mosses or any plant. Lichens do not have roots that absorb water and nutrients as plants do, but like plants, they produce their own nutrition by photosynthesis; when they grow on plants, they do not live as parasites, but instead use the plants as a substrate.
Lichens occur from sea level to high alpine elevations, in many environmental conditions, can grow on any surface. Lichens are abundant growing on bark, mosses, on other lichens, hanging from branches "living on thin air" in rain forests and in temperate woodland, they grow on rock, gravestones, exposed soil surfaces, in the soil as part of a biological soil crust. Different kinds of lichens have adapted to survive in some of the most extreme environments on Earth: arctic tundra, hot dry deserts, rocky coasts, toxic slag heaps, they can live inside solid rock, growing between the grains. It is estimated that 6% of Earth's land surface is covered by lichens. There are about 20,000 known species of lichens; some lichens have lost the ability to reproduce sexually, yet continue to speciate. Lichens can be seen as being self-contained miniature ecosystems, where the fungi, algae, or cyanobacteria have the potential to engage with other microorganisms in a functioning system that may evolve as an more complex composite organism.
Lichens may be long-lived, with some considered to be among the oldest living things. They are among the first living things to grow on fresh rock exposed after an event such as a landslide; the long life-span and slow and regular growth rate of some lichens can be used to date events. In American English, "lichen" is pronounced the same as the verb "liken". In British English, both this pronunciation and one rhyming with "kitchen" ) are used. English lichen derives from Greek λειχήν leichēn via Latin lichen; the Greek noun, which means "licker", derives from the verb λείχειν leichein, "to lick". Lichens grow in a wide range of shapes and forms; the shape of a lichen is determined by the organization of the fungal filaments. The nonreproductive tissues, or vegetative body parts, are called the thallus. Lichens are grouped by thallus type, since the thallus is the most visually prominent part of the lichen. Thallus growth forms correspond to a few basic internal structure types. Common names for lichens come from a growth form or color, typical of a lichen genus.
Common groupings of lichen thallus growth forms are: fruticose – growing like a tuft or multiple-branched leafless mini-shrub, upright or hanging down, 3-dimensional branches with nearly round cross section or flattened foliose – growing in 2-dimensional, leaf-like lobes crustose – crust-like, adhering to a surface like a thick coat of paint squamulose – formed of small leaf-like scales crustose below but free at the tips leprose – powdery gelatinous – jelly like filamentous – stringy or like matted hair byssoid – wispy, like teased wool structurelessThere are variations in growth types in a single lichen species, grey areas between the growth type descriptions, overlapping between growth types, so some authors might describe lichens using different growth type descriptions. When a crustose lichen gets old, the center may start to crack up like old-dried paint, old-broken asphalt paving, or like the polygonal "islands" of cracked-up mud in a dried lakebed; this is called being rimose or areolate, the "island" pieces separated by the cracks are called areolas.
The areolas appear separated, but are connected by an underlying "prothallus" or "hypothallus". When a crustose lichen grows from a center and appears to radiate out, it is called crustose placodioid; when the edges of the areolas lift up from the substrate, it is called squamulose. These growth form groups are not defined. Foliose lichens may sometimes branch and appear to be fruticose. Fruticose lichens may have flattened branching parts and appear leafy. Squamulose lichens may appear where the edges lift up. Gelatinous lichens may appear leafy when dry. Means of telling them apart in these cases are in the sections below. Structures involved in reproduction appear as discs, bumps, or squiggly lines on the surface of the thallus; the thallus is not always the part of the lichen, most visually noticeable. Some lichens can grow inside solid rock between the grains, with only the sexual fruiting part visible growing outside the rock; these may be dramatic in color or appearance. Forms of these sexual parts are not in the above growth form categories.
The most visually noticeable reproductive parts are circular, plate-like or disc-like outgrowths, with crinkly edges, are described in sections below. Lichens come in many colors. Coloratio
The Appalachian Mountains called the Appalachians, are a system of mountains in eastern North America. The Appalachians first formed 480 million years ago during the Ordovician Period, they once reached elevations similar to those of the Alps and the Rocky Mountains before experiencing natural erosion. The Appalachian chain is a barrier to east–west travel, as it forms a series of alternating ridgelines and valleys oriented in opposition to most highways and railroads running east–west. Definitions vary on the precise boundaries of the Appalachians; the United States Geological Survey defines the Appalachian Highlands physiographic division as consisting of thirteen provinces: the Atlantic Coast Uplands, Eastern Newfoundland Atlantic, Maritime Acadian Highlands, Maritime Plain, Notre Dame and Mégantic Mountains, Western Newfoundland Mountains, Blue Ridge and Ridge, Saint Lawrence Valley, Appalachian Plateaus, New England province, the Adirondack areas. A common variant definition does not include the Adirondack Mountains, which geologically belong to the Grenville Orogeny and have a different geological history from the rest of the Appalachians.
The mountain range is in the United States but it extends into southeastern Canada, forming a zone from 100 to 300 mi wide, running from the island of Newfoundland 1,500 mi southwestward to Central Alabama in the United States. The range covers parts of the islands of Saint Pierre and Miquelon, which comprise an overseas territory of France; the system is divided into a series of ranges, with the individual mountains averaging around 3,000 ft. The highest of the group is Mount Mitchell in North Carolina at 6,684 feet, the highest point in the United States east of the Mississippi River; the term Appalachian refers to several different regions associated with the mountain range. Most broadly, it refers to the entire mountain range with its surrounding hills and the dissected plateau region; the term is used more restrictively to refer to regions in the central and southern Appalachian Mountains including areas in the states of Kentucky, Virginia, West Virginia, North Carolina, as well as sometimes extending as far south as northern Alabama and western South Carolina, as far north as Pennsylvania, southern Ohio, parts of southern upstate New York.
The Ouachita Mountains in Arkansas and Oklahoma were part of the Appalachians as well but became disconnected through geologic history. While exploring inland along the northern coast of Florida in 1528, the members of the Narváez expedition, including Álvar Núñez Cabeza de Vaca, found a Native American village near present-day Tallahassee, Florida whose name they transcribed as Apalchen or Apalachen; the name was soon altered by the Spanish to Apalachee and used as a name for the tribe and region spreading well inland to the north. Pánfilo de Narváez's expedition first entered Apalachee territory on June 15, 1528, applied the name. Now spelled "Appalachian," it is the fourth-oldest surviving European place-name in the US. After the de Soto expedition in 1540, Spanish cartographers began to apply the name of the tribe to the mountains themselves; the first cartographic appearance of Apalchen is on Diego Gutierrez's map of 1562. The name was not used for the whole mountain range until the late 19th century.
A competing and more popular name was the "Allegheny Mountains", "Alleghenies", "Alleghania". In the early 19th century, Washington Irving proposed renaming the United States either Appalachia or Alleghania. In U. S. dialects in the southern regions of the Appalachians, the word is pronounced, with the third syllable sounding like "latch". In northern parts of the mountain range, it is pronounced or. There is great debate between the residents of the regions as to which pronunciation is the more correct one. Elsewhere, a accepted pronunciation for the adjective Appalachian is, with the last two syllables "-ian" pronounced as in the word "Romanian"; the whole system may be divided into three great sections: Northern: The northern section runs from the Canadian province of Newfoundland and Labrador to the Hudson River. It includes the Long Range Mountains and Annieopsquotch Mountains on the island of Newfoundland, Chic-Choc Mountains and Notre Dame Range in Quebec and New Brunswick, scattered elevations and small ranges elsewhere in Nova Scotia and New Brunswick, the Longfellow Mountains in Maine, the White Mountains in New Hampshire, the Green Mountains in Vermont, The Berkshires in Massachusetts and Connecticut.
The Metacomet Ridge Mountains in Connecticut and south-central Massachusetts, although contained within the Appalachian province, is a younger system and not geologically associated with the Appalachians. The Monteregian Hills, which cross the Green Mountains in Quebec, are unassociated with the Appalachians. Central: The central section goes from the Hudson Valley to the New River running through Virginia and West Virginia, it comprises the Valley Ridges between the Allegheny Front of the Allegheny Plateau and the Great Appalachian Valley, the New York–New Jersey Highlands, the Taconic Mountains in New York, a large portion of the Blue Ridge. Southern: The southern section runs from the New River onwards, it consists of the prolongation of the Blue Ridge, divided into the Western Blue Ridge Front and the Eastern Blue Ridge Front, the Ridge-and-Valley Appalachians, the Cumberland Plateau. The Adirondack Mountains in New Y
Tennessee is a state located in the southeastern region of the United States. Tennessee is the 16th most populous of the 50 United States. Tennessee is bordered by Kentucky to the north, Virginia to the northeast, North Carolina to the east, Georgia and Mississippi to the south, Arkansas to the west, Missouri to the northwest; the Appalachian Mountains dominate the eastern part of the state, the Mississippi River forms the state's western border. Nashville is the state's capital and largest city, with a 2017 population of 667,560. Tennessee's second largest city is Memphis, which had a population of 652,236 in 2017; the state of Tennessee is rooted in the Watauga Association, a 1772 frontier pact regarded as the first constitutional government west of the Appalachians. What is now Tennessee was part of North Carolina, part of the Southwest Territory. Tennessee was admitted to the Union as the 16th state on June 1, 1796. Tennessee was the last state to leave the Union and join the Confederacy at the outbreak of the American Civil War in 1861.
Occupied by Union forces from 1862, it was the first state to be readmitted to the Union at the end of the war. Tennessee furnished more soldiers for the Confederate Army than any other state besides Virginia, more soldiers for the Union Army than the rest of the Confederacy combined. Beginning during Reconstruction, it had competitive party politics, but a Democratic takeover in the late 1880s resulted in passage of disenfranchisement laws that excluded most blacks and many poor whites from voting; this reduced competition in politics in the state until after passage of civil rights legislation in the mid-20th century. In the 20th century, Tennessee transitioned from an agrarian economy to a more diversified economy, aided by massive federal investment in the Tennessee Valley Authority and, in the early 1940s, the city of Oak Ridge; this city was established to house the Manhattan Project's uranium enrichment facilities, helping to build the world's first atomic bombs, two of which were dropped on Imperial Japan near the end of World War II.
Tennessee's major industries include agriculture and tourism. Poultry and cattle are the state's primary agricultural products, major manufacturing exports include chemicals, transportation equipment, electrical equipment; the Great Smoky Mountains National Park, the nation's most visited national park, is headquartered in the eastern part of the state, a section of the Appalachian Trail follows the Tennessee-North Carolina border. Other major tourist attractions include the Tennessee Aquarium in Chattanooga; the earliest variant of the name that became Tennessee was recorded by Captain Juan Pardo, the Spanish explorer, when he and his men passed through an American Indian village named "Tanasqui" in 1567 while traveling inland from South Carolina. In the early 18th century, British traders encountered a Cherokee town named Tanasi in present-day Monroe County, Tennessee; the town was located on a river of the same name, appears on maps as early as 1725. It is not known whether this was the same town as the one encountered by Juan Pardo, although recent research suggests that Pardo's "Tanasqui" was located at the confluence of the Pigeon River and the French Broad River, near modern Newport.
The meaning and origin of the word are uncertain. Some accounts suggest, it has been said to mean "meeting place", "winding river", or "river of the great bend". According to ethnographer James Mooney, the name "can not be analyzed" and its meaning is lost; the modern spelling, Tennessee, is attributed to James Glen, the governor of South Carolina, who used this spelling in his official correspondence during the 1750s. The spelling was popularized by the publication of Henry Timberlake's "Draught of the Cherokee Country" in 1765. In 1788, North Carolina created "Tennessee County", the third county to be established in what is now Middle Tennessee; when a constitutional convention met in 1796 to organize a new state out of the Southwest Territory, it adopted "Tennessee" as the name of the state. Tennessee is known as The Volunteer State, a nickname some claimed was earned during the War of 1812 because of the prominent role played by volunteer soldiers from Tennessee during the Battle of New Orleans.
Other sources differ on the origin of the state nickname. This explanation is more because President Polk's call for 2,600 nationwide volunteers at the beginning of the Mexican–American War resulted in 30,000 volunteers from Tennessee alone in response to the death of Davy Crockett and appeals by former Tennessee Governor and Texas politician, Sam Houston. Tennessee borders eight other states: Virginia to the north. Tennessee is tied with Missouri as the state bordering the most other states; the state is trisected by the Tennessee River. The highest point in the state is Clingmans Dome at 6,643 feet (
A fungus is any member of the group of eukaryotic organisms that includes microorganisms such as yeasts and molds, as well as the more familiar mushrooms. These organisms are classified as a kingdom, separate from the other eukaryotic life kingdoms of plants and animals. A characteristic that places fungi in a different kingdom from plants and some protists is chitin in their cell walls. Similar to animals, fungi are heterotrophs. Fungi do not photosynthesize. Growth is their means of mobility, except for spores, which may travel through the water. Fungi are the principal decomposers in ecological systems; these and other differences place fungi in a single group of related organisms, named the Eumycota, which share a common ancestor, an interpretation, strongly supported by molecular phylogenetics. This fungal group oomycetes; the discipline of biology devoted to the study of fungi is known as mycology. In the past, mycology was regarded as a branch of botany, although it is now known fungi are genetically more related to animals than to plants.
Abundant worldwide, most fungi are inconspicuous because of the small size of their structures, their cryptic lifestyles in soil or on dead matter. Fungi include symbionts of plants, animals, or other fungi and parasites, they may become noticeable when fruiting, either as molds. Fungi perform an essential role in the decomposition of organic matter and have fundamental roles in nutrient cycling and exchange in the environment, they have long been used in the form of mushrooms and truffles. Since the 1940s, fungi have been used for the production of antibiotics, more various enzymes produced by fungi are used industrially and in detergents. Fungi are used as biological pesticides to control weeds, plant diseases and insect pests. Many species produce bioactive compounds called mycotoxins, such as alkaloids and polyketides, that are toxic to animals including humans; the fruiting structures of a few species contain psychotropic compounds and are consumed recreationally or in traditional spiritual ceremonies.
Fungi can break down manufactured materials and buildings, become significant pathogens of humans and other animals. Losses of crops due to fungal diseases or food spoilage can have a large impact on human food supplies and local economies; the fungus kingdom encompasses an enormous diversity of taxa with varied ecologies, life cycle strategies, morphologies ranging from unicellular aquatic chytrids to large mushrooms. However, little is known of the true biodiversity of Kingdom Fungi, estimated at 2.2 million to 3.8 million species. Of these, only about 120,000 have been described, with over 8,000 species known to be detrimental to plants and at least 300 that can be pathogenic to humans. Since the pioneering 18th and 19th century taxonomical works of Carl Linnaeus, Christian Hendrik Persoon, Elias Magnus Fries, fungi have been classified according to their morphology or physiology. Advances in molecular genetics have opened the way for DNA analysis to be incorporated into taxonomy, which has sometimes challenged the historical groupings based on morphology and other traits.
Phylogenetic studies published in the last decade have helped reshape the classification within Kingdom Fungi, divided into one subkingdom, seven phyla, ten subphyla. The English word fungus is directly adopted from the Latin fungus, used in the writings of Horace and Pliny; this in turn is derived from the Greek word sphongos, which refers to the macroscopic structures and morphology of mushrooms and molds. The word mycology is derived from the Greek logos, it denotes the scientific study of fungi. The Latin adjectival form of "mycology" appeared as early as 1796 in a book on the subject by Christiaan Hendrik Persoon; the word appeared in English as early as 1824 in a book by Robert Kaye Greville. In 1836 the English naturalist Miles Joseph Berkeley's publication The English Flora of Sir James Edward Smith, Vol. 5. Refers to mycology as the study of fungi. A group of all the fungi present in a particular area or geographic region is known as mycobiota, e.g. "the mycobiota of Ireland". Before the introduction of molecular methods for phylogenetic analysis, taxonomists considered fungi to be members of the plant kingdom because of similarities in lifestyle: both fungi and plants are immobile, have similarities in general morphology and growth habitat.
Like plants, fungi grow in soil and, in the case of mushrooms, form conspicuous fruit bodies, which sometimes resemble plants such as mosses. The fungi are now considered a separate kingdom, distinct from both plants and animals, from which they appear to have diverged around one billion years ago; some morphological and genetic features are shared with other organisms, while others are unique to the fungi separating them from the other kingdoms: Shared features: With other euka
Wikispecies is a wiki-based online project supported by the Wikimedia Foundation. Its aim is to create a comprehensive free content catalogue of all species. Jimmy Wales stated that editors are not required to fax in their degrees, but that submissions will have to pass muster with a technical audience. Wikispecies is available under the GNU Free Documentation License and CC BY-SA 3.0. Started in September 2004, with biologists across the world invited to contribute, the project had grown a framework encompassing the Linnaean taxonomy with links to Wikipedia articles on individual species by April 2005. Benedikt Mandl co-ordinated the efforts of several people who are interested in getting involved with the project and contacted potential supporters in early summer 2004. Databases were evaluated and the administrators contacted, some of them have agreed on providing their data for Wikispecies. Mandl defined two major tasks: Figure out how the contents of the data base would need to be presented—by asking experts, potential non-professional users and comparing that with existing databases Figure out how to do the software, which hardware is required and how to cover the costs—by asking experts, looking for fellow volunteers and potential sponsorsAdvantages and disadvantages were discussed by the wikimedia-I mailing list.
The board of directors of the Wikimedia Foundation voted by 4 to 0 in favor of the establishment of a Wikispecies. The project is hosted at species.wikimedia.org. It was merged to a sister project of Wikimedia Foundation on September 14, 2004. On October 10, 2006, the project exceeded 75,000 articles. On May 20, 2007, the project exceeded 100,000 articles with a total of 5,495 registered users. On September 8, 2008, the project exceeded 150,000 articles with a total of 9,224 registered users. On October 23, 2011, the project reached 300,000 articles. On June 16, 2014, the project reached 400,000 articles. On January 7, 2017, the project reached 500,000 articles. On October 30, 2018, the project reached 600,000 articles, a total of 1.12 million pages. Wikispecies comprises taxon pages, additionally pages about synonyms, taxon authorities, taxonomical publications, institutions or repositories holding type specimen. Wikispecies asks users to use images from Wikimedia Commons. Wikispecies does not allow the use of content.
All Species Foundation Catalogue of Life Encyclopedia of Life Tree of Life Web Project List of online encyclopedias The Plant List Wikispecies, The free species directory that anyone can edit Species Community Portal The Wikispecies Charter, written by Wales
Binomial nomenclature called binominal nomenclature or binary nomenclature, is a formal system of naming species of living things by giving each a name composed of two parts, both of which use Latin grammatical forms, although they can be based on words from other languages. Such a name is called a binomen, binominal name or a scientific name; the first part of the name – the generic name – identifies the genus to which the species belongs, while the second part – the specific name or specific epithet – identifies the species within the genus. For example, humans belong within this genus to the species Homo sapiens. Tyrannosaurus rex is the most known binomial; the formal introduction of this system of naming species is credited to Carl Linnaeus beginning with his work Species Plantarum in 1753. But Gaspard Bauhin, in as early as 1623, had introduced in his book Pinax theatri botanici many names of genera that were adopted by Linnaeus; the application of binomial nomenclature is now governed by various internationally agreed codes of rules, of which the two most important are the International Code of Zoological Nomenclature for animals and the International Code of Nomenclature for algae and plants.
Although the general principles underlying binomial nomenclature are common to these two codes, there are some differences, both in the terminology they use and in their precise rules. In modern usage, the first letter of the first part of the name, the genus, is always capitalized in writing, while that of the second part is not when derived from a proper noun such as the name of a person or place. Both parts are italicized when a binomial name occurs in normal text, thus the binomial name of the annual phlox is now written as Phlox drummondii. In scientific works, the authority for a binomial name is given, at least when it is first mentioned, the date of publication may be specified. In zoology "Patella vulgata Linnaeus, 1758"; the name "Linnaeus" tells the reader who it was that first published a description and name for this species of limpet. "Passer domesticus". The original name given by Linnaeus was Fringilla domestica; the ICZN does not require that the name of the person who changed the genus be given, nor the date on which the change was made, although nomenclatorial catalogs include such information.
In botany "Amaranthus retroflexus L." – "L." is the standard abbreviation used in botany for "Linnaeus". "Hyacinthoides italica Rothm. – Linnaeus first named this bluebell species Scilla italica. The name is composed of two word-forming elements: "bi", a Latin prefix for two, "-nomial", relating to a term or terms; the word "binomium" was used in Medieval Latin to mean a two-term expression in mathematics. Prior to the adoption of the modern binomial system of naming species, a scientific name consisted of a generic name combined with a specific name, from one to several words long. Together they formed a system of polynomial nomenclature; these names had two separate functions. First, to designate or label the species, second, to be a diagnosis or description. In a simple genus, containing only two species, it was easy to tell them apart with a one-word genus and a one-word specific name; such "polynomial names" may sometimes look like binomials, but are different. For example, Gerard's herbal describes various kinds of spiderwort: "The first is called Phalangium ramosum, Branched Spiderwort.
The other... is aptly termed Phalangium Ephemerum Virginianum, Soon-Fading Spiderwort of Virginia". The Latin phrases are short descriptions, rather than identifying labels; the Bauhins, in particular Caspar Bauhin, took some important steps towards the binomial system, by pruning the Latin descriptions, in many cases to two words. The adoption by biologists of a system of binomial nomenclature is due to Swedish botanist and physician Carl von Linné, more known by his Latinized name Carl Linnaeus, it was in his 1753 Species Plantarum that he first began using a one-word "trivial name" together with a generic name in a system of binomial nomenclature. This trivial name is what is now known as specific name; the Bauhins' genus names were retained in many of these, but the descriptive part was reduced to a single word. Linnaeus's trivial names introduced an important new idea, namely that the function of a name could be to give a species a unique label; this meant. Thus Gerard's Phalangium ephemerum virginianum became Tradescantia virgi
In biology, a monotypic taxon is a taxonomic group that contains only one subordinate taxon. A monotypic species is one. In the case of genera, the term "unispecific" or "monospecific" is sometimes preferred. In botanical nomenclature, a monotypic genus is a genus in the special case where a genus and a single species are described. In contrast an oligotypic taxon contains more than one but only a few subordinate taxa. Just as the term monotypic is used to describe a taxon including only one subdivision, one can refer to the contained taxon as monotypic within the higher-level taxon, e.g. a genus monotypic within a family. Some examples of monotypic groups are: In the order Amborellales, there is only one family and there is only one genus, in this genus there is only one species, namely Amborella trichopoda The family Cephalotaceae includes only one genus and only one species, Cephalotus follicularis – the Albany pitcher plant. Anodorhynchus hyacinthinus, the hyacinth macaw, is a monotypic species.
Panurus biarmicus, the bearded reedling, has a few subspecies across its range, but belongs to the genus Panurus, which current knowledge considers monotypic within the family Panuridae. Ozichthys albimaculosus, the cream-spotted cardinalfish, found in tropical Australia and southern New Guinea, is the type species of the monotypic genus Ozichthys; the flowering plant Breonadia salicina is the only species in the monotypic genus Breonadia. The madrone butterfly is the only species in the monotypic genus Eucheira. However, there are two sub-species of this butterfly, E. socialis socialis and E. socialis westwoodi. Glossary of scientific naming Monophyly Race