1.
Life
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Various forms of life exist, such as plants, animals, fungi, protists, archaea, and bacteria. The criteria can at times be ambiguous and may or may not define viruses, viroids, biology is the primary science concerned with the study of life, although many other sciences are involved. The definition of life is controversial, the current definition is that organisms maintain homeostasis, are composed of cells, undergo metabolism, can grow, adapt to their environment, respond to stimuli, and reproduce. However, many other definitions have been proposed, and there are some borderline cases. Modern definitions are more complex, with input from a diversity of scientific disciplines, biophysicists have proposed many definitions based on chemical systems, there are also some living systems theories, such as the Gaia hypothesis, the idea that the Earth itself is alive. Another theory is that life is the property of systems, and yet another is elaborated in complex systems biology. Abiogenesis describes the process of life arising from non-living matter. Properties common to all organisms include the need for certain chemical elements to sustain biochemical functions. Life on Earth first appeared as early as 4.28 billion years ago, soon after ocean formation 4.41 billion years ago, Earths current life may have descended from an RNA world, although RNA-based life may not have been the first. The mechanism by which began on Earth is unknown, though many hypotheses have been formulated and are often based on the Miller–Urey experiment. The earliest known forms are microfossils of bacteria. In July 2016, scientists reported identifying a set of 355 genes believed to be present in the last universal ancestor of all living organisms. Since its primordial beginnings, life on Earth has changed its environment on a time scale. To survive in most ecosystems, life must often adapt to a range of conditions. Some microorganisms, called extremophiles, thrive in physically or geochemically extreme environments that are detrimental to most other life on Earth, Aristotle was the first person to classify organisms. Later, Carl Linnaeus introduced his system of nomenclature for the classification of species. Eventually new groups and categories of life were discovered, such as cells and microorganisms, cells are sometimes considered the smallest units and building blocks of life. There are two kinds of cells, prokaryotic and eukaryotic, both of which consist of cytoplasm enclosed within a membrane and contain many such as proteins
2.
Domain (biology)
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In biological taxonomy, a domain is the highest taxonomic rank of organisms in the three-domain system of taxonomy designed by Carl Woese, an American microbiologist and biophysicist. According to the Woese system, introduced in 1990, the tree of life consists of three domains, Archaea, Bacteria, and Eukarya, the first two are all prokaryotic microorganisms, or single-celled organisms whose cells have no nucleus. All life that has a nucleus and membrane-bound organelles, and multicellular organisms, is included in the Eukarya, stefan Luketa in 2012 proposed the five-domain system of life with both cellular and non-cellular organisms. The term domain is proposed by Woese et al. in his three-domain system and this term represents a synonym for the category of dominion, introduced by Moore in 1974. However, only S. Luketa uses the term dominion, each of these three domains of life recognized by biologists today contain unique rRNA. This fact in itself forms the basis of the three-domain system, Archaea are prokaryotic cells, typically characterized by membrane lipids that are branched hydrocarbon chains attached to glycerol by ether linkages. The presence of ether linkages in Archaea adds to their ability to withstand extreme temperatures and highly acidic conditions. Halophiles, organisms that thrive in salty environments, and hyperthermophiles. Archaea evolved many cell sizes, but all are relatively small and their size ranges from 0.1 to 15 μ diameter and up to 200 μ long. They are about the size of bacteria or similar to the size of a mitochondrion in a eukaryotic cell, members of the genus Thermoplasma are the smallest of the archaea. Even though bacteria are prokaryotic cells just like Archaea, their membranes are made of unbranched fatty acid chains attached to glycerol by ester linkages, cyanobacteria and mycoplasmas are two examples of bacteria. They characteristically do not have ether linkages like Archaea, and they are grouped into a different category—and hence a different domain. There is a deal of diversity in this domain. Members of the domain Eukarya have membrane-bound organelles and are represented by four kingdoms, Plantae, Protista, Animalia, none of the three systems currently include non-cellular life. As of 2011 there is talk about Nucleocytoplasmic large DNA viruses possibly being a branch domain of life. Stefan Luketa in 2012 proposed a system, adding Prionobiota and Virusobiota to the traditional three domains. Alternative classifications of life so far proposed include, The two-empire system or superdomain system, the eocyte hypothesis, first proposed by James A. Lake et al. in 1984, which posits two domains
3.
Kingdom (biology)
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In biology, kingdom is the second highest taxonomic rank below domain. Kingdoms are divided into groups called phyla. Traditionally, textbooks from the United States used a system of six kingdoms while textbooks in Great Britain, India, Australia, Latin America and other countries used five kingdoms. Later two further main ranks were introduced, making the kingdom, phylum or division, class, order, family, genus. In the 1960s a rank was introduced above kingdom, namely domain, prefixes can be added so subkingdom and infrakingdom are the two ranks immediately below kingdom. Superkingdom may be considered as an equivalent of domain or empire or as an independent rank between kingdom and domain or subdomain, in some classification systems the additional rank branch can be inserted between subkingdom and infrakingdom. Taxonomic ranks, including kingdoms, were to be groups of organisms with a common ancestor, based on such RNA studies, Carl Woese thought life could be divided into three large divisions and referred to them as the three primary kingdom model or urkingdom model. In 1990, the domain was proposed for the highest rank. This term represents a synonym for the category of dominion, introduced by Moore in 1974, unlike Moore, Whoese et al. did not suggest a Latin term for this category, which represents a further argument supporting the accurately introduced term dominion. It was also found that the eukaryotes are more related to the Archaea than they are to the Eubacteria. Although the primacy of the Eubacteria-Archaea divide has been questioned, it has been upheld by subsequent research, there is no consensus on how many kingdoms exist in the classification scheme proposed by Woese. In 2004, a article by Simpson and Roger noted that the Protista were a grab-bag for all eukaryotes that are not animals. On this basis, the diagram opposite showed the real kingdoms of the eukaryotes, a classification which followed this approach was produced in 2005 for the International Society of Protistologists, by a committee which worked in collaboration with specialists from many societies. It divided the eukaryotes into the same six supergroups, the published classification deliberately did not use formal taxonomic ranks, including that of kingdom. In this system the multicellular animals are descended from the same ancestor as both the unicellular choanoflagellates and the fungi which form the Opisthokonta, Plants are thought to be more distantly related to animals and fungi. Beyond this, there does not appear to be a consensus, rogozin et al. in 2009 noted that The deep phylogeny of eukaryotes is an extremely difficult and controversial problem. The classification of living things into animals and plants is an ancient one, aristotle classified animal species in his History of Animals, while his pupil Theophrastus wrote a parallel work, the Historia Plantarum, on plants. Carl Linnaeus laid the foundations for biological nomenclature, now regulated by the Nomenclature Codes
4.
Phylum
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In biology, a phylum is a taxonomic rank below kingdom and above class. Traditionally, in botany the term division was used instead of phylum, although from 1993 the International Code of Nomenclature for algae, fungi, depending on definitions, the kingdom Animalia contains approximately 35 phyla, Plantae contains about 12, and Fungi contains around 7. Current research in phylogenetics is uncovering the relationships between phyla, which are contained in larger clades, like Ecdysozoa and Embryophyta, the definitions of zoological phyla have changed from their origins in the six Linnaean classes and the four embranchements of Georges Cuvier. Haeckel introduced the term phylum, based on the Greek word phylon, in plant taxonomy, Eichler classified plants into five groups, named divisions. Informally, phyla can be thought of as grouping organisms based on general specialization of body plan, the most important objective measure in the above definitions is the certain degree—how unrelated do organisms need to be to be members of different phyla. The minimal requirement is that all organisms in a phylum should be more closely related to one another than to any other group. So phyla can be merged or split if it becomes apparent that they are related to one another or not, a definition of a phylum based on body plan has been proposed by paleontologists Graham Budd and Sören Jensen. By Budd and Jensens definition, a phylum is defined by a set of shared by all its living representatives. This approach brings some small problems—for instance, ancestral characters common to most members of a phylum may have been lost by some members, also, this definition is based on an arbitrary point of time, the present. However, as it is based, it is easy to apply to the fossil record. A greater problem is that it relies on a decision about which groups of organisms should be considered as phyla. However, proving that a fossil belongs to the group of a phylum is difficult. Furthermore, organisms in the group of a phylum can possess the body plan of the phylum without all the characteristics necessary to fall within it. This weakens the idea that each of the phyla represents a body plan. A classification using this definition may be affected by the chance survival of rare groups. Representatives of many modern phyla did not appear until long after the Cambrian, the kingdom Plantae is defined in various ways by different biologists. All definitions include the living embryophytes, to which may be added the two green algae divisions, Chlorophyta and Charophyta, to form the clade Viridiplantae. The table below follows the influential Cavalier-Smith system in equating Plantae with Archaeplastida, a group containing Viridiplantae, the definition and classification of plants at the division level also varies from source to source, and has changed progressively in recent years
5.
Class (biology)
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In biological classification, class is, a taxonomic rank. Other well-known ranks in descending order of size are life, domain, kingdom, phylum, order, family, genus, as for the other well-known ranks, there is the option of an immediately lower rank, indicated by the prefix sub-, subclass. A taxonomic unit, a taxon, in that rank, in that case the plural is classes The composition of each class is determined by a taxonomist. Often there is no agreement, with different taxonomists taking different positions. There are no rules that a taxonomist needs to follow in describing a class. For example, dogs are assigned to the phylum Chordata, in the class Mammalia. In botany, classes are now rarely discussed, for some clades, a number of alternative classifications are used. In the first edition of his Systema Naturae, carl Linnaeus divided all three of his kingdoms of Nature into classes. The class was considered the highest level of the hierarchy until George Cuviers embranchements. Cladistics List of animal classes Phylogenetics Systematics Taxonomy
6.
Order (biology)
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In biological classification, the order is a taxonomic rank used in the classification of organisms and recognized by the nomenclature codes. Other well-known ranks are life, domain, kingdom, phylum, class, family, genus, an immediately higher rank, superorder, may be added directly above order, while suborder would be a lower rank. A taxonomic unit, a taxon, in that rank, in that case the plural is orders. Example, The Juglans and Hickory trees belong to the family Juglandaceae, what does and does not belong to each order is determined by a taxonomist, as is whether a particular order should be recognized at all. Often there is no agreement, with different taxonomists each taking a different position. There are no rules that a taxonomist needs to follow in describing or recognizing an order. Some taxa are accepted almost universally, while others are recognised only rarely, for some groups of organisms, consistent suffixes are used to denote that the rank is an order. The Latin suffix -formes meaning having the form of is used for the name of orders of birds and fishes. The suffix -ales is for the name of orders of plants, fungi, for some clades covered by the International Code of Zoological Nomenclature, a number of additional classifications are sometimes used, although not all of these are officially recognised. In their 1997 classification of mammals, McKenna and Bell used two levels between superorder and order, grandorder and mirorder. Michael Novacek inserted them at the same position, michael Benton inserted them between superorder and magnorder instead. This position was adopted by Systema Naturae 2000 and others, in botany, the ranks of subclass and suborder are secondary ranks pre-defined as respectively above and below the rank of order. Any number of further ranks can be used as long as they are clearly defined, the superorder rank is commonly used, with the ending -anae that was initiated by Armen Takhtajans publications from 1966 onwards. Carl Linnaeus was the first to apply it consistently to the division of all three kingdoms of nature in his Systema Naturae, for plants, Linnaeus orders in the Systema Naturae and the Species Plantarum were strictly artificial, introduced to subdivide the artificial classes into more comprehensible smaller groups. In French botanical publications, from Michel Adansons Familles naturelles des plantes and until the end of the 19th century, some of the plant families still retain the names of Linnaean natural orders or even the names of pre-Linnaean natural groups recognised by Linnaeus as orders in his natural classification. Such names are known as family names. In zoology, the Linnaean orders were used more consistently and that is, the orders in the zoology part of the Systema Naturae refer to natural groups. Some of his names are still in use
7.
Family (biology)
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In biological classification, family is one of the eight major taxonomic ranks, it is classified between order and genus. A family may be divided into subfamilies, which are intermediate ranks above the rank of genus. In vernacular usage, a family may be named one of its common members, for example, walnuts and hickory trees belong to the family Juglandaceae. What does or does not belong to a family—or whether a family should be recognized at all—are proposed and determined by practicing taxonomists. There are no rules for describing or recognizing a family. Taxonomists often take different positions about descriptions of taxa, and there may be no broad consensus across the community for some time. Some described taxa are accepted broadly and quickly, but others only rarely, if at all, the naming of families is codified by various international codes. In zoological nomenclature, the names of animals end with the suffix -idae. The concept of rank at time was not yet settled, and in the preface to the Prodromus Magnol spoke of uniting his families into larger genera. Carolus Linnaeus used the word familia in his Philosophia botanica to denote groups of plants, trees, herbs, ferns, palms. He used this term only in the section of the book. In zoology, the family as an intermediate between order and genus was introduced by Pierre André Latreille in his Précis des caractères génériques des insectes. He used families in some but not in all his orders of insects, families can be used for evolutionary, palaeontological and generic studies because they are more stable than lower taxonomic levels such as genera and species
8.
Genus
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A genus is a taxonomic rank used in the biological classification of living and fossil organisms in biology. In the hierarchy of classification, genus comes above species. In binomial nomenclature, the name forms the first part of the binomial species name for each species within the genus. Felis catus and Felis silvestris are two species within the genus Felis, Felis is a genus within the family Felidae. The composition of a genus is determined by a taxonomist, the standards for genus classification are not strictly codified, so different authorities often produce different classifications for genera. Moreover, genera should be composed of units of the same kind as other genera. The term comes from the Latin genus, a noun form cognate with gignere, linnaeus popularized its use in his 1753 Species Plantarum, but the French botanist Joseph Pitton de Tournefort is considered the founder of the modern concept of genera. The scientific name of a genus may be called the name or generic epithet. It plays a role in binomial nomenclature, the system of naming organisms. The rules for the names of organisms are laid down in the Nomenclature Codes. The standard way of scientifically describing species and other lower-ranked taxa is by binomial nomenclature, the generic name forms its first half. For example, the gray wolfs binomial name is Canis lupus, with Canis being the name shared by the wolfs close relatives. The specific name is written in lower-case and may be followed by names in zoology or a variety of infraspecific names in botany. Especially with these names, when the generic name is known from context. Because animals are typically only grouped within subspecies, it is written as a trinomen with a third name. Dog breeds, meanwhile, are not scientifically distinguished, there are several divisions of plant species and therefore their infraspecific names generally include contractions explaining the relation. For example, the genus Hibiscus includes hundreds of other species apart from the Rose of Sharon or common garden hibiscus, Rose of Sharon doesnt have subspecies but has cultivars that carry desired traits, such as the bright white H. syriaca Diana. Hawaiian hibiscus, meanwhile, includes several separate species, since not all botanists agree on the divisions or names between species, it is common to specify the source of the name using author abbreviations
9.
Species
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In biology, a species is the basic unit of biological classification and a taxonomic rank. A species is defined as the largest group of organisms in which two individuals can produce fertile offspring, typically by sexual reproduction. While this definition is often adequate, looked at more closely it is problematic, for example, with hybridisation, in a species complex of hundreds of similar microspecies, or in a ring species, the boundaries between closely related species become unclear. Other ways of defining species include similarity of DNA, morphology, all species are given a two-part name, a binomial. The first part of a binomial is the genus to which the species belongs, the second part is called the specific name or the specific epithet. For example, Boa constrictor is one of four species of the Boa genus, Species were seen from the time of Aristotle until the 18th century as fixed kinds that could be arranged in a hierarchy, the great chain of being. In the 19th century, biologists grasped that species could evolve given sufficient time, Charles Darwins 1859 book The Origin of Species explained how species could arise by natural selection. Genes can sometimes be exchanged between species by horizontal transfer, and species may become extinct for a variety of reasons. In his biology, Aristotle used the term γένος to mean a kind, such as a bird or fish, a kind was distinguished by its attributes, for instance, a bird has feathers, a beak, wings, a hard-shelled egg, and warm blood. A form was distinguished by being shared by all its members, Aristotle believed all kinds and forms to be distinct and unchanging. His approach remained influential until the Renaissance, when observers in the Early Modern period began to develop systems of organization for living things, they placed each kind of animal or plant into a context. Many of these early delineation schemes would now be considered whimsical, animals likewise that differ specifically preserve their distinct species permanently, one species never springs from the seed of another nor vice versa. In the 18th century, the Swedish scientist Carl Linnaeus classified organisms according to shared physical characteristics and he established the idea of a taxonomic hierarchy of classification based upon observable characteristics and intended to reflect natural relationships. At the time, however, it was widely believed that there was no organic connection between species, no matter how similar they appeared. However, whether or not it was supposed to be fixed, by the 19th century, naturalists understood that species could change form over time, and that the history of the planet provided enough time for major changes. Jean-Baptiste Lamarck, in his 1809 Zoological Philosophy, described the transmutation of species, proposing that a species could change over time, in 1859, Charles Darwin and Alfred Russel Wallace provided a compelling account of evolution and the formation of new species. Darwin argued that it was populations that evolved, not individuals and this required a new definition of species. Darwin concluded that species are what appear to be, ideas
10.
Taxonomic rank
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In biological classification, rank is the relative level of a group of organisms in a taxonomic hierarchy. Examples of taxonomic ranks are species, genus, family, order, class, phylum, kingdom, domain, a given rank subsumes under it less general categories, that is, more specific descriptions of life forms. Above it, each rank is classified within more general categories of organisms, the rank of any species and the description of its genus is basic, which means that to identify a particular organism, it is usually not necessary to specify ranks other than these first two. Consider a particular species, the red fox Vulpes vulpes, its next rank, as one group of the class Mammalia, all of the above are classified among those with backbones in the Chordata phylum rank, and with them among all the animals in the Animalia kingdom rank. Finally, all of the above will find their earliest relatives somewhere in their domain rank Eukarya, today, nomenclature is regulated by the nomenclature codes. There are seven main ranks, kingdom, phylum or division, class, order, family, genus. In addition, the domain is now used as one of the fundamental ranks. Also, this represents a synonym for the category of dominion. Unlike Moore, Whoese et al. did not suggest a Latin term for this category, a taxon is usually assigned a rank when it is given its formal name. The basic ranks are species and genus, when an organism is given a species name it is assigned to a genus, and the genus name is part of the species name. The species name is called a binomial, that is. For example, the name for the human species is Homo sapiens. This is usually italicized in print and underlined when italics are not available, in this case, Homo is the generic name and it is capitalized, sapiens indicates the species and it is not capitalized. There are definitions of the taxonomic ranks in the International Code of Zoological Nomenclature, superfamily, family, subfamily, tribe, subtribe, genus, subgenus. The International Code of Zoological Nomenclature divides names into family-group names, genus-group names, in the genus group and species group no further ranks are allowed. At higher ranks a lower level may be denoted by adding the prefix infra, meaning lower, a taxon above the rank of species has a scientific name in one part. A species has a composed of two parts, generic name + specific name, for example Canis lupus. A subspecies has a composed of three parts, generic name + specific name + subspecific name, for example Canis lupus familiaris
11.
Taxonomy (biology)
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Taxonomy is the science of defining groups of biological organisms on the basis of shared characteristics and giving names to those groups. The exact definition of taxonomy varies from source to source, but the core of the remains, the conception, naming. There is some disagreement as to whether biological nomenclature is considered a part of taxonomy, the broadest meaning of taxonomy is used here. The word taxonomy was introduced in 1813 by Candolle, in his Théorie élémentaire de la botanique, the term alpha taxonomy is primarily used today to refer to the discipline of finding, describing, and naming taxa, particularly species. In earlier literature, the term had a different meaning, referring to morphological taxonomy, ideals can, it may be said, never be completely realized. They have, however, a value of acting as permanent stimulants. Some of us please ourselves by thinking we are now groping in a beta taxonomy, turrill thus explicitly excludes from alpha taxonomy various areas of study that he includes within taxonomy as a whole, such as ecology, physiology, genetics, and cytology. He further excludes phylogenetic reconstruction from alpha taxonomy, thus, Ernst Mayr in 1968 defined beta taxonomy as the classification of ranks higher than species. This activity is what the term denotes, it is also referred to as beta taxonomy. How species should be defined in a group of organisms gives rise to practical and theoretical problems that are referred to as the species problem. The scientific work of deciding how to define species has been called microtaxonomy, by extension, macrotaxonomy is the study of groups at higher taxonomic ranks, from subgenus and above only, than species. While some descriptions of taxonomic history attempt to date taxonomy to ancient civilizations, earlier works were primarily descriptive, and focused on plants that were useful in agriculture or medicine. There are a number of stages in scientific thinking. Early taxonomy was based on criteria, the so-called artificial systems. Later came systems based on a complete consideration of the characteristics of taxa, referred to as natural systems, such as those of de Jussieu, de Candolle and Bentham. The publication of Charles Darwins Origin of Species led to new ways of thinking about classification based on evolutionary relationships and this was the concept of phyletic systems, from 1883 onwards. This approach was typified by those of Eichler and Engler, the advent of molecular genetics and statistical methodology allowed the creation of the modern era of phylogenetic systems based on cladistics, rather than morphology alone. Taxonomy has been called the worlds oldest profession, and naming and classifying our surroundings has likely been taking place as long as mankind has been able to communicate
12.
Taxon
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In biology, a taxon is a group of one or more populations of an organism or organisms seen by taxonomists to form a unit. Although neither is required, a taxon is known by a particular name and given a particular ranking, especially if. It is not uncommon, however, for taxonomists to remain at odds over what belongs to a taxon, if a taxon is given a formal scientific name, its use is then governed by one of the nomenclature codes specifying which scientific name is correct for a particular grouping. Many modern systematists, such as advocates of phylogenetic nomenclature, use cladistic methods that require taxa to be monophyletic and their basic unit, therefore, is the clade rather than the taxon. Similarly, among those contemporary taxonomists working with the traditional Linnean nomenclature, an example of a well-established taxon that is not also a clade is the class Reptilia, the reptiles, birds are descendants of reptiles but are not included in the Reptilia. The term taxon was first used in 1926 by Adolf Meyer-Abich for animal groups, for plants, it was proposed by Herman Johannes Lam in 1948, and it was adopted at the VII International Botanical Congress, held in 1950. The Glossary of the International Code of Zoological Nomenclature defines a taxon, a taxon encompasses all included taxa of lower rank and individual organisms. A taxon can be assigned a rank, usually when it is given a formal name. Phylum applies formally to any domain, but traditionally it was always used for animals, whereas Division was traditionally often used for plants, fungi. A prefix is used to indicate a ranking of lesser importance, the prefix super- indicates a rank above, the prefix sub- indicates a rank below. In zoology the prefix infra- indicates a rank below sub-, for instance, among the additional ranks of class are superclass, subclass and infraclass. Rank is relative, and restricted to a particular systematic schema, for example, liverworts have been grouped, in various systems of classification, as a family, order, class, or division. Many cladists do not see any need to depart from traditional nomenclature as governed by the ICZN, ICN, etc
13.
Dog
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The domestic dog is a member of genus Canis that forms part of the wolf-like canids, and is the most widely abundant carnivore. The dog and the extant gray wolf are sister taxa, with modern wolves not closely related to the wolves that were first domesticated, the dog was the first domesticated species and has been selectively bred over millennia for various behaviors, sensory capabilities, and physical attributes. Their long association with humans has led dogs to be attuned to human behavior. Dogs vary widely in shape, size and colours, dogs perform many roles for people, such as hunting, herding, pulling loads, protection, assisting police and military, companionship and, more recently, aiding handicapped individuals. This influence on society has given them the sobriquet mans best friend. The term domestic dog is used for both domesticated and feral varieties. The English word dog comes from Middle English dogge, from Old English docga, the term may possibly derive from Proto-Germanic *dukkōn, represented in Old English finger-docce. The word also shows the familiar petname diminutive -ga also seen in frogga frog, picga pig, stagga stag, wicga beetle, worm, the term dog may ultimately derive from the earliest layer of Proto-Indo-European vocabulary. In 14th-century England, hound was the word for all domestic canines, and dog referred to a subtype of hound. It is believed this dog type was so common, it became the prototype of the category hound. By the 16th century, dog had become the general word, the word hound is ultimately derived from the Proto-Indo-European word *kwon-, dog. This semantic shift may be compared with in German, where the corresponding words Dogge, a male canine is referred to as a dog, while a female is called a bitch. The father of a litter is called the sire, and the mother is called the dam, the process of birth is whelping, from the Old English word hwelp, the modern English word whelp is an alternate term for puppy. A litter refers to the offspring at one birth which are called puppies or pups from the French poupée, doll. The term dog typically is applied both to the species as a whole, and any male member of the same. An adult female is a bitch, in some countries, especially in North America, dog is used instead due to the vulgar connotation of bitch. An adult male capable of reproduction is a stud, an adult female capable of reproduction is a brood bitch, or brood mother. Immature males or females are pups or puppies, a group of pups from the same gestation period is a litter
14.
Mammal
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Mammals are any vertebrates within the class Mammalia, a clade of endothermic amniotes distinguished from reptiles by the possession of a neocortex, hair, three middle ear bones and mammary glands. All female mammals nurse their young with milk, secreted from the mammary glands, Mammals include the largest animals on the planet, the great whales. The basic body type is a quadruped, but some mammals are adapted for life at sea, in the air, in trees. The largest group of mammals, the placentals, have a placenta, Mammals range in size from the 30–40 mm bumblebee bat to the 30-meter blue whale. With the exception of the five species of monotreme, all modern mammals give birth to live young, most mammals, including the six most species-rich orders, belong to the placental group. The largest orders are the rodents, bats and Soricomorpha, the next three biggest orders, depending on the biological classification scheme used, are the Primates, the Cetartiodactyla, and the Carnivora. Living mammals are divided into the Yinotheria and Theriiformes There are around 5450 species of mammal, in some classifications, extant mammals are divided into two subclasses, the Prototheria, that is, the order Monotremata, and the Theria, or the infraclasses Metatheria and Eutheria. The marsupials constitute the group of the Metatheria, and include all living metatherians as well as many extinct ones. Much of the changes reflect the advances of cladistic analysis and molecular genetics, findings from molecular genetics, for example, have prompted adopting new groups, such as the Afrotheria, and abandoning traditional groups, such as the Insectivora. The mammals represent the only living Synapsida, which together with the Sauropsida form the Amniota clade, the early synapsid mammalian ancestors were sphenacodont pelycosaurs, a group that produced the non-mammalian Dimetrodon. At the end of the Carboniferous period, this group diverged from the line that led to todays reptiles. Some mammals are intelligent, with some possessing large brains, self-awareness, Mammals can communicate and vocalize in several different ways, including the production of ultrasound, scent-marking, alarm signals, singing, and echolocation. Mammals can organize themselves into fission-fusion societies, harems, and hierarchies, most mammals are polygynous, but some can be monogamous or polyandrous. They provided, and continue to provide, power for transport and agriculture, as well as commodities such as meat, dairy products, wool. Mammals are hunted or raced for sport, and are used as model organisms in science, Mammals have been depicted in art since Palaeolithic times, and appear in literature, film, mythology, and religion. Defaunation of mammals is primarily driven by anthropogenic factors, such as poaching and habitat destruction, Mammal classification has been through several iterations since Carl Linnaeus initially defined the class. No classification system is accepted, McKenna & Bell and Wilson & Reader provide useful recent compendiums. Though field work gradually made Simpsons classification outdated, it remains the closest thing to a classification of mammals
15.
Flowering plant
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The flowering plants, also known as Angiospermae or Magnoliophyta, are the most diverse group of land plants, with 416 families, approx. 13,164 known genera and a total of c.295,383 known species, etymologically, angiosperm means a plant that produces seeds within an enclosure, in other words, a fruiting plant. The term angiosperm comes from the Greek composite word meaning enclosed seeds, the ancestors of flowering plants diverged from gymnosperms in the Triassic Period, during the range 245 to 202 million years ago, and the first flowering plants are known from 160 mya. They diversified extensively during the Lower Cretaceous, became widespread by 120 mya, angiosperms differ from other seed plants in several ways, described in the table. These distinguishing characteristics taken together have made the angiosperms the most diverse and numerous land plants, the amount and complexity of tissue-formation in flowering plants exceeds that of gymnosperms. The vascular bundles of the stem are arranged such that the xylem and phloem form concentric rings, in the dicotyledons, the bundles in the very young stem are arranged in an open ring, separating a central pith from an outer cortex. In each bundle, separating the xylem and phloem, is a layer of meristem or active formative tissue known as cambium, the soft phloem becomes crushed, but the hard wood persists and forms the bulk of the stem and branches of the woody perennial. Among the monocotyledons, the bundles are more numerous in the stem and are scattered through the ground tissue. They contain no cambium and once formed the stem increases in diameter only in exceptional cases, the characteristic feature of angiosperms is the flower. Flowers show remarkable variation in form and elaboration, and provide the most trustworthy external characteristics for establishing relationships among angiosperm species, the function of the flower is to ensure fertilization of the ovule and development of fruit containing seeds. The floral apparatus may arise terminally on a shoot or from the axil of a leaf, occasionally, as in violets, a flower arises singly in the axil of an ordinary foliage-leaf. There are two kinds of cells produced by flowers. Microspores, which divide to become pollen grains, are the male cells and are borne in the stamens. The female cells called megaspores, which divide to become the egg cell, are contained in the ovule. The flower may consist only of parts, as in willow. Usually, other structures are present and serve to protect the sporophylls, the individual members of these surrounding structures are known as sepals and petals. The outer series is usually green and leaf-like, and functions to protect the rest of the flower, the inner series is, in general, white or brightly colored, and is more delicate in structure. It functions to attract insect or bird pollinators, attraction is effected by color, scent, and nectar, which may be secreted in some part of the flower
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Clade
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A clade is a group of organisms that consists of a common ancestor and all its lineal descendants, and represents a single branch on the tree of life. The common ancestor may be an individual, a population, a species, clades are nested, one in another, as each branch in turn splits into smaller branches. These splits reflect evolutionary history as populations diverged and evolved independently, over the last few decades, the cladistic approach has revolutionized biological classification and revealed surprising evolutionary relationships among organisms. Increasingly, taxonomists try to avoid naming taxa that are not clades, the term clade was coined in 1957 by the biologist Julian Huxley to refer to the result of cladogenesis, a concept Huxley borrowed from Bernhard Rensch. Many commonly named groups are clades, for example, rodents or insects, because in each case, their name refers to a common ancestor with all its descendant branches. Rodents, for example, are a branch of mammals that split off after the end of the period when the clade Dinosauria stopped being the dominant terrestrial vertebrates 66 million years ago, the original population and all its descendants are a clade. The rodent clade corresponds to the order Rodentia, and insects to the class Insecta and these clades include smaller clades, such as chipmunk or ant, each of which consists of even smaller clades. The clade rodent is in included in the mammal, vertebrate. The phenomenon of convergent evolution is, however, responsible for cases where there are misleading similarities in the morphology of groups that evolved from different lineages. The publication of Darwins theory of evolution in 1859 gave this view increasing weight, thomas Henry Huxley, an early advocate of evolutionary theory, proposed a revised taxonomy based on clades. For example, he grouped birds with reptiles, based on fossil evidence, german biologist Emil Hans Willi Hennig is considered to be the founder of cladistics. Taxonomists have increasingly worked to make the taxonomic system reflect evolution, when it comes to naming, however, this principle is not always compatible with the traditional rank-based nomenclature. For these and other reasons, phylogenetic nomenclature has been developed, a clade is by definition monophyletic, meaning that it contains one ancestor and all its descendants. The ancestor can be known or unknown, any and all members of a clade can be extant or extinct, the science that tries to reconstruct phylogenetic trees and thus discover clades is called phylogenetics or cladistics, the latter term coined by Ernst Mayr, derived from clade. The results of analyses are tree-shaped diagrams called cladograms, they. Three methods of defining clades are featured in phylogenetic nomenclature, node-, stem-, the relationship between clades can be described in several ways, A clade located within a clade is said to be nested within that clade. In the diagram, the clade, i. e. the apes. Two clades are sisters if they have a common ancestor
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Tetrapod
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The superclass Tetrapoda contains the four-limbed vertebrates known as tetrapods, it includes living and extinct amphibians, reptiles and mammals, as well as earlier extinct groups. The first tetrapods appeared by the late Devonian,367, the first tetrapods were primarily aquatic. Their descendants drove most amphibians to extinction as they began to dominate, the amniotes include the tetrapods that further evolved for flight—such as birds from among the dinosaurs, and bats from among the mammals. The change from a plan for breathing and navigating in water to a body plan enabling the animal to move on land is one of the most profound evolutionary changes known. It is also becoming increasingly well-understood as a result of more transitional fossil finds, many tetrapods have returned to partially aquatic or fully aquatic lives throughout the history of the group. Tetrapods can be defined in cladistics as the nearest common ancestor of all living amphibians and all living amniotes, the group so defined is the crown group, or crown tetrapods. The term tetrapodomorph is used for the definition, any animal that is more closely related to living amphibians, reptiles, birds. The group so defined is known as the total group. Stegocephalia is a larger group equivalent to some uses of the word tetrapod. Batrachomorphs are all sharing a more recent common ancestry with living amphibians than with living amniotes. Reptiliomorphs are all sharing a more recent common ancestry with living amniotes than with living amphibians. Tetrapoda includes four classes, amphibians, reptiles, mammals, however, that diversification process was interrupted at least a few times by major biological crises, such as the Permian–Triassic extinction event, which at least affected amniotes. As biodiversity has grown, so has the number of niches that tetrapods have occupied, the first tetrapods were aquatic and fed primarily on fish. Today, the Earth supports a diversity of tetrapods that live in many habitats. Tetrapods evolved from early bony fishes, specifically from the branch of lobe-finned fishes. The first tetrapods probably evolved in the Emsian stage of the Early Devonian from Tetrapodomorph fish living in water environments. The very earliest tetrapods would have been similar to Acanthostega, with legs and lungs as well as gills. The earliest tetrapods inhabited saltwater, brackish-water, and freshwater environments and these traits were shared with many early lobed-finned fishes
18.
Maxillopoda
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Maxillopoda is a diverse class of crustaceans including barnacles, copepods and a number of related animals. It does not appear to be a group, and no single character unites all the members. With the exception of some barnacles, maxillopodans are mostly small, including the smallest known arthropod and they often have short bodies, with the abdomen reduced in size, and generally lacking any appendages. This may have arisen through paedomorphosis, apart from barnacles, which use their legs for filter feeding, most maxillopodans feed with their maxillae. They have a bauplan comprising 5 cephalic segments,6 thoracic segments and 4 abdominal segments, the fossil record of the group extends back into the Cambrian, with fossils of both barnacles and tongue worms known from that period. Six subclasses are generally recognised, although many works have included the ostracods among the Maxillopoda. Skaracarida Media related to Maxillopoda at Wikimedia Commons
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Sauropsida
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Sauropsida is a group of amniotes that includes all existing birds and reptiles as well as their fossil ancestors and other extinct relatives. Large land animals are either in this group or in its group, Synapsida. This clade includes Parareptilia and other extinct clades, all living sauropsids are members of the sub-group Diapsida, the Parareptilia clade having died out 200 million years ago. The term originated in 1864 with Thomas Henry Huxley, who grouped birds with reptiles based on fossil evidence, the term Sauropsida has a long history, and hails back to Thomas Henry Huxley, and his opinion that birds had risen from the dinosaurs. He based this chiefly on the fossils of Hesperornis and Archaeopteryx, early in the following year he proposed the names Sauropsida and Ichthyopsida for the two latter. Huxley did however include groups on the line like Dicynodon among the sauropsids. Thus, under the definition, Sauropsida contained not only the groups usually associated with it today. By the early 20th century, the fossils of Permian synapsids from South Africa had become well known, the term Sauropsida was taken up by E. S. Goodrich in 1916 much like Huxleys, to include lizards, birds and he distinguished them from mammals and their extinct relatives, which he included in the sister group Theropsida. Goodrichs classification thus differs somewhat from Huxleys, in which the non-mammalian synapsids fell under the sauropsids, Goodrich supported this division by the nature of the hearts and blood vessels in each group, and other features such as the structure of the forebrain. According to Goodrich, both lineages evolved from a stem group, the Protosauria, which included some Paleozoic amphibians as well as early reptiles predating the sauropsid/synapsid split. Watson observed that sauropsids and synapsids diverged very early in the evolutionary history. He also reinterpreted the Sauropsida and Theropsida to exclude birds and mammals respectively, making them paraphyletic, thus his Sauropsida included Procolophonia, Eosuchia, Millerosauria, Chelonia, Squamata, Rhynchocephalia, Crocodilia, thecodonts, non-avian dinosaurs, pterosaurs, ichthyosaurs, and sauropyterygians. The class Reptilia has been known to be a grade rather than a clade for as long as evolution has been recognised. Reclassifying reptiles has been among the key aims of phylogenetic nomenclature, the term Sauropsida had from the mid 20th century been used to denote all species not on the synapsid side after the synapsid/sauropsid split, a branch-based clade. Some later cladistic work has used Sauropsida more restrictively, to signify the group, i. e. all descendants of the last common ancestor of extant reptiles. Sauropsids evolved from basal amniotes stock approximately 320 million years ago in the Paleozoic Era, in the Mesozoic Era, sauropsids were the largest animals on land, in the water, and in the air. The Mesozoic is sometimes called the Age of Reptiles, sixty-six million years ago, the large-bodied sauropsids died out in the global extinction event at the end of the Mesozoic era
20.
Theria
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Theria is a subclass of mammals amongst the Theriiformes. Theria includes the eutherians and the metatherians, therian mammals give birth to live young without using a shelled egg. It is possible thanks to key proteins called syncytins, which allow exchanges between the mother and its offspring through a placenta even rudimental such as the marsupial ones, genetic studies have enlighted the viral origin of syncytins through endogenization process. Therian mammals no longer have the coracoid bone, contrary to their cousins monotremes, pinnae are also a distinctive trait that is a therian exclusivity. The earliest known therian mammal fossil is Juramaia, from the Middle Jurassic of China, however, molecular data suggests that therians may have originated even earlier, during the Early Jurassic. The rank of Theria may vary depending on the system used. The textbook classification system by Vaughan et al. gives the following, marsupials Monotremes Placental mammals Theria — supercohort — Tree of Life
21.
Thecostraca
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Thecostraca is a subclass of marine invertebrates containing about 1,320 described species. Many species have larvae which become sessile or parasitic as adults. The most important subgroup are the barnacles, constituting about 1,220 known species, the subgroup Facetotecta contains a single genus, Hansenocaris, known only from the tiny planktonic nauplii called y-larvae. These larvae have no known form, though it is suspected that they are parasites. The group Ascothoracida contains about 100 species, all parasites of coelenterates and echinoderms
22.
Avialae
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Avialae is a clade of flying dinosaurs containing their only living representatives, the birds. It is usually defined as all dinosaurs more closely related to modern birds than to deinonychosaurs. Archaeopteryx lithographica, from the late Jurassic Period Solnhofen Formation of Germany, is the earliest known avialan which may have had the capability of powered flight, however, several older avialans are known from the late Jurassic Tiaojishan Formation of China, dated to about 160 million years ago. Most researchers define Avialae as branch-based clade, though definitions vary, many authors have used a definition similar to all theropods closer to birds than to Deinonychus. A nearly identical definition, the group that includes all taxa closer to Passer than to Dromaeosaurus, was used by Agnolín. Additionally, beginning in the late 2000s and early 2010s, several groups of researchers began adding the genus Troodon as a specifier in the definition of Avialae. Avialae is also defined as an apomorphy-based clade. Jacques Gauthier, who named Avialae in 1986, re-defined it in 2001 as all dinosaurs that possessed feathered wings used in flapping flight, and the birds that descended from them. Gauthier identified four conflicting ways of defining the term Aves, which is a problem because the biological name is being used four different ways. Gauthier proposed a solution, number 4 below, which is to reserve the term Aves only for the crown group and he assigned other names to the other groups. Under the fourth definition Archaeopteryx is an avialan, and not a member of Aves, gauthiers proposals have been adopted by many researchers in the field of paleontology and bird evolution, though the exact definitions applied have been inconsistent. Avialae, initially proposed to replace the traditional content of Aves, is sometimes used synonymously with the vernacular term bird by these researchers. The earliest known fossils come from the Tiaojishan Formation of China. The avialan species from time period include Anchiornis huxleyi, Xiaotingia zhengi. The well-known early avialan, Archaeopteryx, dates from slightly later Jurassic rocks from Germany, many of these early avialans shared unusual anatomical features that may be ancestral to modern birds, but were later lost during bird evolution. Avialans diversified into a variety of forms during the Cretaceous Period. Many groups retained primitive characteristics, such as clawed wings and teeth, though the latter were lost independently in a number of avialan groups, in the late Cretaceous, around 95 million years ago, the ancestor of all modern birds also evolved a better sense of smell
23.
Barnacle
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A barnacle is a type of arthropod constituting the infraclass Cirripedia in the subphylum Crustacea, and is hence related to crabs and lobsters. Barnacles are exclusively marine, and tend to live in shallow and tidal waters and they are sessile suspension feeders, and have two nektonic larval stages. Around 1,220 barnacle species are currently known, the name Cirripedia is Latin, meaning curl-footed. Barnacles are encrusters, attaching themselves permanently to a hard substrate, the most common, acorn barnacles, are sessile, growing their shells directly onto the substrate. The order Pedunculata attach themselves by means of a stalk, free-living barnacles are attached to the substratum by cement glands that form the base of the first pair of antennae, in effect, the animal is fixed upside down by means of its forehead. In some barnacles, the cement glands are fixed to a long, muscular stalk, a ring of plates surrounds the body, homologous with the carapace of other crustaceans. These consist of the rostrum, two plates, two carinolaterals, and a carina. In sessile barnacles, the apex of the ring of plates is covered by an operculum, the plates are held together by various means, depending on species, in some cases being solidly fused. Inside the carapace, the lies on its back, with its limbs projecting upwards. Segmentation is usually indistinct, and the body is more or less divided between the head and thorax, with little, if any, abdomen. Adult barnacles have few appendages on their heads, with only a single, vestigial pair of antennae, attached to the cement gland. The six pairs of limbs are referred to as cirri. Barnacles have no heart, although a sinus close to the esophagus performs similar function. The blood vascular system is minimal, similarly, they have no gills, absorbing oxygen from the water through their limbs and the inner membrane of their carapaces. The excretory organs of barnacles are maxillary glands, the main sense of barnacles appears to be touch, with the hairs on the limbs being especially sensitive. The adult also has an eye, although this is probably only capable of sensing the difference between light and dark. This eye is derived from the naupliar eye. Barnacles have two larval stages, the nauplius and the cyprid, before developing into a mature adult
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Bird
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Birds, a subgroup of Reptiles, are the last living examples of Dinosaurs. Birds live worldwide and range in size from the 5 cm bee hummingbird to the 2.75 m ostrich. They rank as the class of tetrapods with the most living species, at ten thousand. Birds are the closest living relatives of crocodilians, the fossil record indicates that birds evolved from feathered ancestors within the theropod group of saurischian dinosaurs. True birds first appeared during the Cretaceous period, around 100 million years ago, Birds, especially those in the southern continents, survived this event and then migrated to other parts of the world while diversifying during periods of global cooling. Primitive bird-like dinosaurs that lie outside class Aves proper, in the broader group Avialae, have been found dating back to the mid-Jurassic period, around 170 million years ago. Birds have wings which are more or less developed depending on the species, the digestive and respiratory systems of birds are also uniquely adapted for flight. Some bird species of aquatic environments, particularly seabirds and some waterbirds, have evolved for swimming. Many species annually migrate great distances, Birds are social, communicating with visual signals, calls, and bird songs, and participating in such social behaviours as cooperative breeding and hunting, flocking, and mobbing of predators. The vast majority of species are socially monogamous, usually for one breeding season at a time, sometimes for years. Other species have breeding systems that are polygynous or, rarely, Birds produce offspring by laying eggs which are fertilised through sexual reproduction. They are usually laid in a nest and incubated by the parents, most birds have an extended period of parental care after hatching. Some birds, such as hens, lay eggs even when not fertilised, songbirds, parrots, and other species are popular as pets. Guano is harvested for use as a fertiliser, Birds prominently figure throughout human culture. About 120–130 species have become extinct due to human activity since the 17th century, human activity threatens about 1,200 bird species with extinction, though efforts are underway to protect them. Recreational birdwatching is an important part of the ecotourism industry, the first classification of birds was developed by Francis Willughby and John Ray in their 1676 volume Ornithologiae. Carl Linnaeus modified that work in 1758 to devise the taxonomic classification system currently in use, Birds are categorised as the biological class Aves in Linnaean taxonomy. Phylogenetic taxonomy places Aves in the dinosaur clade Theropoda, Aves and a sister group, the clade Crocodilia, contain the only living representatives of the reptile clade Archosauria
25.
France
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France, officially the French Republic, is a country with territory in western Europe and several overseas regions and territories. The European, or metropolitan, area of France extends from the Mediterranean Sea to the English Channel and the North Sea, Overseas France include French Guiana on the South American continent and several island territories in the Atlantic, Pacific and Indian oceans. France spans 643,801 square kilometres and had a population of almost 67 million people as of January 2017. It is a unitary republic with the capital in Paris. Other major urban centres include Marseille, Lyon, Lille, Nice, Toulouse, during the Iron Age, what is now metropolitan France was inhabited by the Gauls, a Celtic people. The area was annexed in 51 BC by Rome, which held Gaul until 486, France emerged as a major European power in the Late Middle Ages, with its victory in the Hundred Years War strengthening state-building and political centralisation. During the Renaissance, French culture flourished and a colonial empire was established. The 16th century was dominated by civil wars between Catholics and Protestants. France became Europes dominant cultural, political, and military power under Louis XIV, in the 19th century Napoleon took power and established the First French Empire, whose subsequent Napoleonic Wars shaped the course of continental Europe. Following the collapse of the Empire, France endured a succession of governments culminating with the establishment of the French Third Republic in 1870. Following liberation in 1944, a Fourth Republic was established and later dissolved in the course of the Algerian War, the Fifth Republic, led by Charles de Gaulle, was formed in 1958 and remains to this day. Algeria and nearly all the colonies became independent in the 1960s with minimal controversy and typically retained close economic. France has long been a centre of art, science. It hosts Europes fourth-largest number of cultural UNESCO World Heritage Sites and receives around 83 million foreign tourists annually, France is a developed country with the worlds sixth-largest economy by nominal GDP and ninth-largest by purchasing power parity. In terms of household wealth, it ranks fourth in the world. France performs well in international rankings of education, health care, life expectancy, France remains a great power in the world, being one of the five permanent members of the United Nations Security Council with the power to veto and an official nuclear-weapon state. It is a member state of the European Union and the Eurozone. It is also a member of the Group of 7, North Atlantic Treaty Organization, Organisation for Economic Co-operation and Development, the World Trade Organization, originally applied to the whole Frankish Empire, the name France comes from the Latin Francia, or country of the Franks
26.
Botany
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Botany, also called plant science, plant biology or phytology, is the science of plant life and a branch of biology. A botanist or plant scientist is a scientist who specialises in this field, the term botany comes from the Ancient Greek word βοτάνη meaning pasture, grass, or fodder, βοτάνη is in turn derived from βόσκειν, to feed or to graze. Nowadays, botanists study approximately 410,000 species of plants of which some 391,000 species are vascular plants. Medieval physic gardens, often attached to monasteries, contained plants of medical importance and they were forerunners of the first botanical gardens attached to universities, founded from the 1540s onwards. One of the earliest was the Padua botanical garden and these gardens facilitated the academic study of plants. Efforts to catalogue and describe their collections were the beginnings of plant taxonomy, in the last two decades of the 20th century, botanists exploited the techniques of molecular genetic analysis, including genomics and proteomics and DNA sequences to classify plants more accurately. Modern botany is a broad, multidisciplinary subject with inputs from most other areas of science, dominant themes in 21st century plant science are molecular genetics and epigenetics, which are the mechanisms and control of gene expression during differentiation of plant cells and tissues. Botany originated as herbalism, the study and use of plants for their medicinal properties, many records of the Holocene period date early botanical knowledge as far back as 10,000 years ago. This early unrecorded knowledge of plants was discovered in ancient sites of human occupation within Tennessee, the early recorded history of botany includes many ancient writings and plant classifications. Examples of early works have been found in ancient texts from India dating back to before 1100 BC, in archaic Avestan writings. His major works, Enquiry into Plants and On the Causes of Plants, constitute the most important contributions to science until the Middle Ages. De Materia Medica was widely read for more than 1,500 years, important contributions from the medieval Muslim world include Ibn Wahshiyyas Nabatean Agriculture, Abū Ḥanīfa Dīnawarīs the Book of Plants, and Ibn Bassals The Classification of Soils. In the early 13th century, Abu al-Abbas al-Nabati, and Ibn al-Baitar wrote on botany in a systematic and scientific manner and these gardens continued the practical value of earlier physic gardens, often associated with monasteries, in which plants were cultivated for medical use. They supported the growth of botany as an academic subject, lectures were given about the plants grown in the gardens and their medical uses demonstrated. Botanical gardens came much later to northern Europe, the first in England was the University of Oxford Botanic Garden in 1621, throughout this period, botany remained firmly subordinate to medicine. German physician Leonhart Fuchs was one of the three German fathers of botany, along with theologian Otto Brunfels and physician Hieronymus Bock, Fuchs and Brunfels broke away from the tradition of copying earlier works to make original observations of their own. Bock created his own system of plant classification, physician Valerius Cordus authored a botanically and pharmacologically important herbal Historia Plantarum in 1544 and a pharmacopoeia of lasting importance, the Dispensatorium in 1546. Naturalist Conrad von Gesner and herbalist John Gerard published herbals covering the medicinal uses of plants, naturalist Ulisse Aldrovandi was considered the father of natural history, which included the study of plants
27.
Joseph Pitton de Tournefort
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Joseph Pitton de Tournefort was a French botanist, notable as the first to make a clear definition of the concept of genus for plants. The botanist Charles Plumier had been his pupil and accompanied him on his voyages, Tournefort was born in Aix-en-Provence and studied at the Jesuit convent there. It was intended that he enter the Church, but the death of his father allowed him to follow his interest in botany, after two years collecting, he studied medicine at Montpellier, but was appointed professor of botany at the Jardin des Plantes in Paris in 1683. During this time he travelled through Western Europe, particularly the Pyrenees and he was accompanied by the German botanist Andreas Gundelsheimer and the artist Claude Aubriet. His description of this journey was published posthumously, he himself having been killed by a carriage in Paris and his principal work was the 1694 Eléments de botanique, ou Méthode pour reconnaître les Plantes. The principal artist was Claude Aubriet who later became the principal artist at the Jardin des Plantes, Tournefort is often credited with being the first to make a clear distinction between genus and species. Though he did indeed cluster the 7,000 plant species that he described into around 700 genera, concepts of genus and species had been framed as early as the 16th century, and Kaspar Bauhin in particular consistently distinguished genera and species. Augustus Quirinus Rivinus had even advocated the use of binary nomenclature shortly before Tourneforts work was published, the word herbarium also seems to have been an invention of Tournefort, previously herbaria had been called by a variety of names, such as Hortus siccus. His herbarium collection of 6,963 specimens was housed in Paris, now part of the Muséum national dhistoire naturelle. Media related to Joseph Pitton de Tournefort at Wikimedia Commons KUL Digital version of Elemens de botanique ou methode pour connoitre les plantes 1694–1695
28.
Systema Naturae
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Systema Naturae is one of the major works of the Swedish botanist, zoologist and physician Carl Linnaeus and introduced the Linnaean taxonomy. The first edition was published in 1735, the tenth edition of this book is considered the starting point of zoological nomenclature. In 1766–1768 Linnaeus published the much enhanced 12th edition, the last under his authorship, another again enhanced work in the same style and entitled Systema Naturae was published by Johann Friedrich Gmelin between 1788 and 1793. Since at least the early 1900s zoologists commonly recognized this as the last edition belonging to this series and it was also officially regarded by the International Commission on Zoological Nomenclature in Opinion 296 as the 13th edition of Systema Naturae. Linnaeus published the first edition of Systema Naturae in the year 1735, as was customary for the scientific literature of its day, the book was published in Latin. In it, he outlined his ideas for the classification of the natural world, dividing it into the animal kingdom, the plant kingdom. Linnaeuss Systema Naturae lists only about 10,000 species of organisms, Linnaeus developed his classification of the plant kingdom in an attempt to describe and understand the natural world as a reflection of the logic of Gods creation. His sexual system, where species with the number of stamens were treated in the same group, was convenient. Linnaeus believed in Gods creation, and that there were no deeper relationships to be expressed and he is frequently quoted to have said. The classification of animals was more natural, for instance, humans were for the first time placed together with other primates, as Anthropomorpha. As a result of the popularity of the work, and the number of new specimens sent to him from around the world, Linnaeus kept publishing new and it grew from eleven very large pages in the first edition to 2,400 pages in the 12th edition. In this same edition, he introduced two-part names for animal species, the system eventually developed into modern Linnaean taxonomy, a hierarchically organized biological classification. After Linnaeus health declined in the early 1770s, publication of editions of Systema Naturae went in two directions, another Swedish scientist, Johan Andreas Murray issued the Regnum Vegetabile section separately in 1774 as the Systema Vegetabilium, rather confusingly labelled the 13th edition. Meanwhile, a 13th edition of the entire Systema appeared in parts between 1788-1793 and it was as the Systema Vegetabilium that Linnaeus work became widely known in England following translation from the Latin by the Lichfield Botanical Society, as A System of Vegetables. In his Imperium Naturæ, Linnaeus established three kingdoms, namely Regnum Animale, Regnum Vegetabile and Regnum Lapideum. This approach, the Animal, Vegetable and Mineral Kingdoms, survives today in the popular mind, notably in the form of parlour games, Is it animal. The classification was based on five levels, kingdom, class, order, genus, while species and genus was seen as God-given, the three higher levels were seen by Linnaeus as constructs. The concept behind the set ranks being applied to all groups was to make a system that was easy to remember and navigate, Linnaeuss work had a huge impact on science, it was indispensable as a foundation for biological nomenclature, now regulated by the Nomenclature Codes
29.
Carl Linnaeus
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Carl Linnaeus, also known after his ennoblement as Carl von Linné, was a Swedish botanist, physician, and zoologist, who formalised the modern system of naming organisms called binomial nomenclature. He is known by the father of modern taxonomy. Many of his writings were in Latin, and his name is rendered in Latin as Carolus Linnæus, Linnaeus was born in the countryside of Småland, in southern Sweden. He received most of his education at Uppsala University. He lived abroad between 1735 and 1738, where he studied and also published a first edition of his Systema Naturae in the Netherlands and he then returned to Sweden, where he became professor of medicine and botany at Uppsala. In the 1740s, he was sent on journeys through Sweden to find and classify plants. In the 1750s and 1760s, he continued to collect and classify animals, plants, and minerals, at the time of his death, he was one of the most acclaimed scientists in Europe. The philosopher Jean-Jacques Rousseau sent him the message, Tell him I know no man on earth. The German writer Johann Wolfgang von Goethe wrote, With the exception of Shakespeare and Spinoza, Swedish author August Strindberg wrote, Linnaeus was in reality a poet who happened to become a naturalist. Among other compliments, Linnaeus has been called Princeps botanicorum, The Pliny of the North and he is also considered as one of the founders of modern ecology. In botany, the abbreviation used to indicate Linnaeus as the authority for species names is L. In older publications, sometimes the abbreviation Linn. is found, Linnæus was born in the village of Råshult in Småland, Sweden, on 23 May 1707. He was the first child of Nicolaus Ingemarsson and Christina Brodersonia and his siblings were Anna Maria Linnæa, Sofia Juliana Linnæa, Samuel Linnæus, and Emerentia Linnæa. One of a line of peasants and priests, Nils was an amateur botanist, a Lutheran minister. Christina was the daughter of the rector of Stenbrohult, Samuel Brodersonius, a year after Linnæus birth, his grandfather Samuel Brodersonius died, and his father Nils became the rector of Stenbrohult. The family moved into the rectory from the curates house, even in his early years, Linnæus seemed to have a liking for plants, flowers in particular. Whenever he was upset, he was given a flower, which calmed him. Nils spent much time in his garden and often showed flowers to Linnaeus, soon Linnæus was given his own patch of earth where he could grow plants
30.
Mineral
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A mineral is a naturally occurring chemical compound, usually of crystalline form and abiogenic in origin. A mineral has one specific chemical composition, whereas a rock can be an aggregate of different minerals or mineraloids, the study of minerals is called mineralogy. There are over 5,300 known mineral species, over 5,070 of these have been approved by the International Mineralogical Association, the silicate minerals compose over 90% of the Earths crust. The diversity and abundance of species is controlled by the Earths chemistry. Silicon and oxygen constitute approximately 75% of the Earths crust, which translates directly into the predominance of silicate minerals, minerals are distinguished by various chemical and physical properties. Differences in chemical composition and crystal structure distinguish the various species, changes in the temperature, pressure, or bulk composition of a rock mass cause changes in its minerals. Minerals can be described by their various properties, which are related to their chemical structure. Common distinguishing characteristics include crystal structure and habit, hardness, lustre, diaphaneity, colour, streak, tenacity, cleavage, fracture, parting, more specific tests for describing minerals include magnetism, taste or smell, radioactivity and reaction to acid. Minerals are classified by key chemical constituents, the two dominant systems are the Dana classification and the Strunz classification, the silicate class of minerals is subdivided into six subclasses by the degree of polymerization in the chemical structure. All silicate minerals have a unit of a 4− silica tetrahedron—that is, a silicon cation coordinated by four oxygen anions. These tetrahedra can be polymerized to give the subclasses, orthosilicates, disilicates, cyclosilicates, inosilicates, phyllosilicates, other important mineral groups include the native elements, sulfides, oxides, halides, carbonates, sulfates, and phosphates. The first criterion means that a mineral has to form by a natural process, stability at room temperature, in the simplest sense, is synonymous to the mineral being solid. More specifically, a compound has to be stable or metastable at 25 °C, modern advances have included extensive study of liquid crystals, which also extensively involve mineralogy. Minerals are chemical compounds, and as such they can be described by fixed or a variable formula, many mineral groups and species are composed of a solid solution, pure substances are not usually found because of contamination or chemical substitution. Finally, the requirement of an ordered atomic arrangement is usually synonymous with crystallinity, however, crystals are also periodic, an ordered atomic arrangement gives rise to a variety of macroscopic physical properties, such as crystal form, hardness, and cleavage. There have been recent proposals to amend the definition to consider biogenic or amorphous substances as minerals. The formal definition of an approved by the IMA in 1995, A mineral is an element or chemical compound that is normally crystalline. However, if geological processes were involved in the genesis of the compound, Mineral classification schemes and their definitions are evolving to match recent advances in mineral science
31.
Plant
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Plants are mainly multicellular, predominantly photosynthetic eukaryotes of the kingdom Plantae. The term is generally limited to the green plants, which form an unranked clade Viridiplantae. This includes the plants, conifers and other gymnosperms, ferns, clubmosses, hornworts, liverworts, mosses and the green algae. Green plants have cell walls containing cellulose and obtain most of their energy from sunlight via photosynthesis by primary chloroplasts and their chloroplasts contain chlorophylls a and b, which gives them their green color. Some plants are parasitic and have lost the ability to produce amounts of chlorophyll or to photosynthesize. Plants are characterized by sexual reproduction and alternation of generations, although reproduction is also common. There are about 300–315 thousand species of plants, of which the great majority, green plants provide most of the worlds molecular oxygen and are the basis of most of Earths ecologies, especially on land. Plants that produce grains, fruits and vegetables form humankinds basic foodstuffs, Plants play many roles in culture. They are used as ornaments and, until recently and in variety, they have served as the source of most medicines. The scientific study of plants is known as botany, a branch of biology, Plants are one of the two groups into which all living things were traditionally divided, the other is animals. The division goes back at least as far as Aristotle, who distinguished between plants, which generally do not move, and animals, which often are mobile to catch their food. Much later, when Linnaeus created the basis of the system of scientific classification. Since then, it has become clear that the plant kingdom as originally defined included several unrelated groups, however, these organisms are still often considered plants, particularly in popular contexts. When the name Plantae or plant is applied to a group of organisms or taxon. The evolutionary history of plants is not yet settled. Those which have been called plants are in bold, the way in which the groups of green algae are combined and named varies considerably between authors. Algae comprise several different groups of organisms which produce energy through photosynthesis, most conspicuous among the algae are the seaweeds, multicellular algae that may roughly resemble land plants, but are classified among the brown, red and green algae. Each of these groups also includes various microscopic and single-celled organisms
32.
Animal
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Animals are multicellular, eukaryotic organisms of the kingdom Animalia. The animal kingdom emerged as a clade within Apoikozoa as the group to the choanoflagellates. Animals are motile, meaning they can move spontaneously and independently at some point in their lives and their body plan eventually becomes fixed as they develop, although some undergo a process of metamorphosis later in their lives. All animals are heterotrophs, they must ingest other organisms or their products for sustenance, most known animal phyla appeared in the fossil record as marine species during the Cambrian explosion, about 542 million years ago. Animals can be divided broadly into vertebrates and invertebrates, vertebrates have a backbone or spine, and amount to less than five percent of all described animal species. They include fish, amphibians, reptiles, birds and mammals, the remaining animals are the invertebrates, which lack a backbone. These include molluscs, arthropods, annelids, nematodes, flatworms, cnidarians, ctenophores, the study of animals is called zoology. The word animal comes from the Latin animalis, meaning having breath, the biological definition of the word refers to all members of the kingdom Animalia, encompassing creatures as diverse as sponges, jellyfish, insects, and humans. Aristotle divided the world between animals and plants, and this was followed by Carl Linnaeus, in the first hierarchical classification. In Linnaeuss original scheme, the animals were one of three kingdoms, divided into the classes of Vermes, Insecta, Pisces, Amphibia, Aves, and Mammalia. Since then the last four have all been subsumed into a single phylum, in 1874, Ernst Haeckel divided the animal kingdom into two subkingdoms, Metazoa and Protozoa. The protozoa were later moved to the kingdom Protista, leaving only the metazoa, thus Metazoa is now considered a synonym of Animalia. Animals have several characteristics that set apart from other living things. Animals are eukaryotic and multicellular, which separates them from bacteria and they are heterotrophic, generally digesting food in an internal chamber, which separates them from plants and algae. They are also distinguished from plants, algae, and fungi by lacking cell walls. All animals are motile, if only at life stages. In most animals, embryos pass through a stage, which is a characteristic exclusive to animals. With a few exceptions, most notably the sponges and Placozoa and these include muscles, which are able to contract and control locomotion, and nerve tissues, which send and process signals
33.
Linnaean taxonomy
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Linnaean taxonomy can mean either of two related concepts, the particular form of biological classification set up by Carl Linnaeus, as set forth in his Systema Naturae and subsequent works. In the taxonomy of Linnaeus there are three kingdoms, divided into classes, and they, in turn, into orders, families, genera, a term for rank-based classification of organisms, in general. That is, taxonomy in the sense of the word. This term is used as opposed to cladistic systematics, which groups organisms into clades. It is attributed to Linnaeus, although he neither invented the concept of ranked classification nor gave it its present form, in his Imperium Naturae, Linnaeus established three kingdoms, namely Regnum Animale, Regnum Vegetabile and Regnum Lapideum. This approach, the Animal, Vegetable and Mineral Kingdoms, survives today in the mind, notably in the form of the parlour game question, Is it animal. The work of Linnaeus had a impact on science, it was indispensable as a foundation for biological nomenclature. However, the impact he had on science was not because of the value of his taxonomy and his classes and orders of plants, according to his Systema Sexuale, were never intended to represent natural groups but only for use in identification. They were used for that purpose well into the nineteenth century, within each class were several orders. The Linnaean classes for plants, in the Sexual System, were, monandria, flowers with 1 stamen Classis 2. Diandria, flowers with 2 stamens Classis 3, triandria, flowers with 3 stamens Classis 4. Tetrandria, flowers with 4 stamens Classis 5, pentandria, flowers with 5 stamens Classis 6. Hexandria, flowers with 6 stamens Hexandria monogynia pp. 285–352 Hexandria polygynia pp. 342–343 Classis 7, heptandria, flowers with 7 stamens Classis 8. Octandria, flowers with 8 stamens Classis 9, enneandria, flowers with 9 stamens Classis 10. Decandria, flowers with 10 stamens Classis 11, dodecandria, flowers with 12 stamens Classis 12. Icosandria, flowers with 20 stamens, perigynous Classis 13, polyandria, flowers with many stamens, inserted on the receptacle Classis 14. Didynamia, flowers with 4 stamens,2 long and 2 short Classis 15, tetradynamia, flowers with 6 stamens,4 long and 2 short Classis 16. Monadelphia, flowers with the separate, but the filaments united
34.
Georges Cuvier
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Jean Léopold Nicolas Frédéric Cuvier, known as Georges Cuvier, was a French naturalist and zoologist, sometimes referred to as the father of paleontology. Cuvier is also known for establishing extinction as a fact—at the time, in his Essay on the Theory of the Earth Cuvier was interpreted to have proposed that new species were created after periodic catastrophic floods. In this way, Cuvier became the most influential proponent of catastrophism in geology in the early 19th century and his study of the strata of the Paris basin with Alexandre Brongniart established the basic principles of biostratigraphy. Cuvier is also remembered for strongly opposing theories of evolution, which at the time were mainly proposed by Jean-Baptiste de Lamarck, Cuvier believed there was no evidence for evolution, but rather evidence for cyclical creations and destructions of life forms by global extinction events such as deluges. Cuvier supported function and rejected Lamarcks thinking and his most famous work is Le Règne Animal. In 1819, he was created a peer for life in honor of his scientific contributions, thereafter, he was known as Baron Cuvier. He died in Paris during an epidemic of cholera, some of Cuviers most influential followers were Louis Agassiz on the continent and in the United States, and Richard Owen in Britain. His name is one of the 72 names inscribed on the Eiffel Tower, Cuvier was born in Montbéliard, France, where his Protestant ancestors had lived since the time of the Reformation. His father, Jean George Cuvier, was a lieutenant in the Swiss Guards, at the time, the town, which was annexed to France on 10 October 1793, belonged to the Duchy of Württemberg. His mother, who was younger than his father, tutored him diligently throughout his early years. During his gymnasium years, he had trouble acquiring Latin and Greek, and was always at the head of his class in mathematics, history. At the age of 10, soon entering the gymnasium, he encountered a copy of Conrad Gessners Historiae Animalium. He then began frequent visits to the home of a relation, all of these he read and reread, retaining so much of the information, that by the age of 12, he was as familiar with quadrupeds and birds as a first-rate naturalist. He remained at the gymnasium for four years, Cuvier spent an additional four years at the Caroline Academy in Stuttgart, where he excelled in all of his coursework. Although he knew no German on his arrival, after nine months of study. Upon graduation, he had no money on which to live as he awaited appointment to an academic office, so in July 1788, he took a job at Fiquainville chateau in Normandy as tutor to the only son of the Comte dHéricy, a Protestant noble. There, during the early 1790s, he began his comparisons of fossils with extant forms, Cuvier regularly attended meetings held at the nearby town of Valmont for the discussion of agricultural topics. There, he acquainted with Henri Alexandre Tessier, who had assumed a false identity
35.
Ernst Haeckel
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The published artwork of Haeckel includes over 100 detailed, multi-colour illustrations of animals and sea creatures. As a philosopher, Ernst Haeckel wrote Die Welträtsel, the genesis for the term world riddle, Ernst Haeckel was born on 16 February 1834, in Potsdam. In 1852, Haeckel completed studies at the Domgymnasium, the high school of Merseburg. He then studied medicine in Berlin and Würzburg, particularly with Albert von Kölliker, Franz Leydig, Rudolf Virchow, together with Hermann Steudner he attended botany lectures in Würzburg. In 1857, Haeckel attained a doctorate in medicine, and afterwards he received a license to practice medicine, the occupation of physician appeared less worthwhile to Haeckel, after contact with suffering patients. Between 1859 and 1866, Haeckel worked on many phyla such radiolarians, poriferans, during a trip to the Mediterranean, Haeckel named nearly 150 new species of radiolarians. In 1867, he married Agnes Huschke and their son Walter was born in 1868, their daughters Elizabeth in 1871 and Emma in 1873. In 1869, he traveled as a researcher to Norway, in 1871 to Croatia, and in 1873 to Egypt, Turkey, Haeckel retired from teaching in 1909, and in 1910 he withdrew from the Evangelical church. Haeckels wife, Agnes, died in 1915, and Haeckel became substantially frailer, with a broken leg and he sold his Villa Medusa in Jena in 1918 to the Carl Zeiss foundation, and it presently contains a historic library. Haeckel died on 9 August 1919, Haeckels political beliefs were influenced by his affinity for the German Romantic movement coupled with his acceptance of a form of Lamarckism. Rather than being a strict Darwinian, Haeckel believed that the characteristics of an organism were acquired through interactions with the environment and he believed the social sciences to be instances of applied biology, and that phrase was picked up and used for Nazi propaganda. In 1905, Haeckel founded a group called the Monist League to promote his religious and this group lasted until 1933 and included such notable members as Wilhelm Ostwald, Georg von Arco, Helene Stöcker and Walter Arthur Berendsohn. Haeckel was the first person known to use the term First World War. Shortly after the start of the war Haeckel wrote, There is no doubt that the course, Haeckel was a zoologist, an accomplished artist and illustrator, and later a professor of comparative anatomy. For example, Haeckel described and named hypothetical ancestral microorganisms that have never been found and he was one of the first to consider psychology as a branch of physiology. He also proposed the kingdom Protista in 1866 and his chief interests lay in evolution and life development processes in general, including development of nonrandom form, which culminated in the beautifully illustrated Kunstformen der Natur. Haeckel did not support natural selection, rather believing in Lamarckism and it proposed a link between ontogeny and phylogeny, summed up by Haeckel in the phrase ontogeny recapitulates phylogeny. His concept of recapitulation has been refuted in the form he gave it, the strong recapitulation hypothesis views ontogeny as repeating forms of the ancestors, while weak recapitulation means that what is repeated is the ancestral embryonic development process
36.
Cladistics
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Therefore, members of a group are assumed to share a common history and are considered to be closely related. The techniques and nomenclature of cladistics have been applied to other disciplines, Cladistics in the original sense refers to a particular set of methods used in phylogenetic analysis, although it is now sometimes used to refer to the whole field. What is now called the cladistic method appeared as early as 1901 with a work by Peter Chalmers Mitchell for birds and subsequently by Robert John Tillyard in 1921, Hennig referred to his own approach as phylogenetic systematics. Phenetics was championed at this time by the numerical taxonomists Peter Sneath and Robert Sokal, originally conceived, if only in essence, by Willi Hennig in a book published in 1950, cladistics did not flourish until its translation into English in 1966. Today, cladistics is the most popular method for constructing phylogenies from morphological, unlike phenetics, cladistics is specifically aimed at reconstructing evolutionary histories. The way for computational phylogenetics was paved by phenetics, a set of commonly used from the 1950s to 1980s. Phenetics did not try to reconstruct phylogenetic trees, rather, it tried to build dendrograms from similarity data, the cladistic method interprets each character state transformation implied by the distribution of shared character states among taxa as a potential piece of evidence for grouping. The outcome of an analysis is a cladogram – a tree-shaped diagram that is interpreted to represent the best hypothesis of phylogenetic relationships. Cladists contend that these models are unjustified, every cladogram is based on a particular dataset analyzed with a particular method. Different datasets and different methods, not to mention violations of the mentioned assumptions, only scientific investigation can show which is more likely to be correct. Since the cladograms provide competing accounts of events, at most one of them is correct. Within the primates, all anthropoids are hypothesized to have had a common ancestor all of whose descendants were anthropoids, the prosimians, on the other hand, form a paraphyletic taxon. When two or more taxa that are not nested within each other share a plesiomorphy, it is a symplesiomorphy, symplesiomorphies do not mean that the taxa that exhibit that character state are necessarily closely related. For example, Reptilia is traditionally characterized by being cold-blooded, whereas birds are warm-blooded, an apomorphy or derived state is an innovation. It can thus be used to diagnose a clade – or even to help define a clade name in phylogenetic nomenclature, features that are derived in individual taxa are called autapomorphies. Autapomorphies express nothing about relationships among groups, clades are identified by synapomorphies, for example, the possession of digits that are homologous with those of Homo sapiens is a synapomorphy within the vertebrates. The tetrapods can be singled out as consisting of the first vertebrate with such digits homologous to those of Homo sapiens together with all descendants of this vertebrate and it is therefore inferred to have evolved by convergence or reversal. Both mammals and birds are able to maintain a constant body temperature
37.
Phylogenetics
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In biology, phylogenetics /ˌfaɪloʊdʒəˈnɛtɪks, -lə-/ is the study of the evolutionary history and relationships among individuals or groups of organisms. These relationships are discovered through phylogenetic inference methods that evaluate observed heritable traits, the result of these analyses is a phylogeny – a diagrammatic hypothesis about the history of the evolutionary relationships of a group of organisms. The tips of a tree can be living organisms or fossils. Phylogenetic analyses have become central to understanding biodiversity, evolution, ecology, taxonomy is the classification, identification and naming of organisms. It is usually richly informed by phylogenetics, but remains a methodologically and logically distinct discipline, usual methods of phylogenetic inference involve computational approaches implementing the optimality criteria and methods of parsimony, maximum likelihood, and MCMC-based Bayesian inference. All these depend upon an implicit or explicit mathematical model describing the evolution of characters observed, prior to 1990, phylogenetic inferences were generally presented as narrative scenarios. Such methods are often ambiguous and lack explicit criteria for evaluating alternative hypotheses, the term phylogeny derives from the German Phylogenie, introduced by Haeckel in 1866, and the Darwinian approach to classification became known as the phyletic approach. During the late 19th century, Ernst Haeckels recapitulation theory, or biogenetic fundamental law, was widely accepted, but this theory has long been rejected. O. See, Huerta-Cepas, Jaime, Dopazo, Joaquín, Gabaldón, ETE, A python Environment for Tree Exploration. PhylomeDB, A public database hosting thousands of gene phylogenies ranging many different species, See, Huerta-Cepas, J. Capella-Gutierrez, S. Pryszcz, L. P. Denisov, I. Kormes, D. Marcet-Houben, M. Gabaldón, T. PhylomeDB v3.0, An expanding repository of genome-wide collections of trees, alignments, lents, N. H. Cifuentes, O. E. Carpi, A. Teaching the Process of Molecular Phylogeny and Systematics, A Multi-Part Inquiry-Based Exercise
38.
Systematics
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Biological systematics is the study of the diversification of living forms, both past and present, and the relationships among living things through time. Relationships are visualized as evolutionary trees, phylogenies have two components, branching order and branch length. Phylogenetic trees of species and higher taxa are used to study the evolution of traits, Systematics, in other words, is used to understand the evolutionary history of life on Earth. John Lindley provided a definition of systematics in 1830, although he wrote of systematic botany rather than using the term systematics. This is a field with a history that in recent years has experienced a notable renaissance. Part of the material has to do with evolutionary areas. Taxonomy is that part of Systematics concerned with topics to above, however, in modern usage, they can all be considered synonyms of each other. For example, Websters 9th New Collegiate Dictionary of 1987 treats classification, taxonomy, according to this work the terms originated in 1790, c. Europeans tend to use the terms systematics and biosystematics for the field of the study of biodiversity as a whole, all of these biological disciplines can deal both with extinct and with extant organisms. Scientific classifications are aids in recording and reporting information to other scientists, the systematist, a scientist who specializes in systematics, must, therefore, be able to use existing classification systems, or at least know them well enough to skilfully justify not using them. Phenetics was an attempt to determine the relationships of organisms through a measure of overall similarity, from the late-20th century onwards, it was superseded by cladistics, which rejects plesiomorphies in attempting to resolve the phylogeny of Earths various organisms through time. Todays systematists generally make use of molecular biology and of computer programs to study organisms. Taxonomic characters are the attributes that can be used to provide the evidence from which relationships between taxa are inferred. Adaptation, or evolutionary mechanisms Scientific classification and Taxonomy - the result of research in systematics Schuh, Randall T. biological Systematics, Principles and Applications, 2nd edn. ISBN 978-0-8014-4799-0 Simpson, Michael G.2005, ISBN 978-0-12-644460-5 Society of Australian Systematic Biologists Society of Systematic Biologists Willi Hennig Society
39.
Ernst Mayr
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Not to be confused with Ernst Mayr, Ernst Mayer, Ernst Meyer, Ernest Mayer or Ernest May. Ernst Walter Mayr was one of the 20th centurys leading evolutionary biologists and he was also a renowned taxonomist, tropical explorer, ornithologist, philosopher of biology, and historian of science. Although Charles Darwin and others posited that multiple species could evolve from a common ancestor. Ernst Mayr approached the problem with a new definition for species, the most significant and rapid genetic reorganization occurs in extremely small populations that have been isolated. Mayr was the son of Helene Pusinelli and Dr. Otto Mayr. His father was a jurist but took an interest in natural history and he learnt all the local birds in Würzburg from his elder brother Otto. He also had access to a history magazine for amateurs. His father died just before he was thirteen, the family then moved to Dresden and he studied at the Staatsgymnasium in Dresden-Neustadt and completed his high school education there. In April 1922, while still in school, he joined the newly founded Saxony Ornithologists’ Association. Here he met Rudolf Zimmermann, who became his ornithological mentor, in February 1923, Mayr passed his high school examination and his mother rewarded him with a pair of binoculars. On 23 March 1923 on the lakes of Moritzburg, the Frauenteich, the species had not been seen in Saxony since 1845 and the local club argued about the identity. Raimund Schelcher of the club suggested that Mayr visit his classmate Erwin Stresemann on his way to Greifswald. After a tough interrogation, Stresemann accepted and published the sighting as authentic, Stresemann was very impressed and suggested that, between semesters, Mayr could work as a volunteer in the ornithological section of the museum. Mayr wrote about this event, It was as if someone had given me the key to heaven, Mayr was endlessly interested in ornithology and chose Greifswald at the Baltic for my studies for no other reason than that. It was situated in the ornithologically most interesting area, although he ostensibly planned to become a physician, he was first and foremost an ornithologist. During the first semester break Stresemann gave him a test to identify treecreepers, Stresemann declared that Mayr was a born systematist. Mayr completed his doctorate in ornithology at the University of Berlin under Dr. Carl Zimmer, on 1 July he accepted the position offered to him at the museum for a monthly salary of 330.54 Reichsmark. In New Guinea, Mayr collected several thousand bird skins and, during his stay in New Guinea, he was invited to accompany the Whitney South Seas Expedition to the Solomon Islands
40.
International Standard Book Number
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The International Standard Book Number is a unique numeric commercial book identifier. An ISBN is assigned to each edition and variation of a book, for example, an e-book, a paperback and a hardcover edition of the same book would each have a different ISBN. The ISBN is 13 digits long if assigned on or after 1 January 2007, the method of assigning an ISBN is nation-based and varies from country to country, often depending on how large the publishing industry is within a country. The initial ISBN configuration of recognition was generated in 1967 based upon the 9-digit Standard Book Numbering created in 1966, the 10-digit ISBN format was developed by the International Organization for Standardization and was published in 1970 as international standard ISO2108. Occasionally, a book may appear without a printed ISBN if it is printed privately or the author does not follow the usual ISBN procedure, however, this can be rectified later. Another identifier, the International Standard Serial Number, identifies periodical publications such as magazines, the ISBN configuration of recognition was generated in 1967 in the United Kingdom by David Whitaker and in 1968 in the US by Emery Koltay. The 10-digit ISBN format was developed by the International Organization for Standardization and was published in 1970 as international standard ISO2108, the United Kingdom continued to use the 9-digit SBN code until 1974. The ISO on-line facility only refers back to 1978, an SBN may be converted to an ISBN by prefixing the digit 0. For example, the edition of Mr. J. G. Reeder Returns, published by Hodder in 1965, has SBN340013818 -340 indicating the publisher,01381 their serial number. This can be converted to ISBN 0-340-01381-8, the check digit does not need to be re-calculated, since 1 January 2007, ISBNs have contained 13 digits, a format that is compatible with Bookland European Article Number EAN-13s. An ISBN is assigned to each edition and variation of a book, for example, an ebook, a paperback, and a hardcover edition of the same book would each have a different ISBN. The ISBN is 13 digits long if assigned on or after 1 January 2007, a 13-digit ISBN can be separated into its parts, and when this is done it is customary to separate the parts with hyphens or spaces. Separating the parts of a 10-digit ISBN is also done with either hyphens or spaces, figuring out how to correctly separate a given ISBN number is complicated, because most of the parts do not use a fixed number of digits. ISBN issuance is country-specific, in that ISBNs are issued by the ISBN registration agency that is responsible for country or territory regardless of the publication language. Some ISBN registration agencies are based in national libraries or within ministries of culture, in other cases, the ISBN registration service is provided by organisations such as bibliographic data providers that are not government funded. In Canada, ISBNs are issued at no cost with the purpose of encouraging Canadian culture. In the United Kingdom, United States, and some countries, where the service is provided by non-government-funded organisations. Australia, ISBNs are issued by the library services agency Thorpe-Bowker
41.
Superphylum
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In biology, a phylum is a taxonomic rank below kingdom and above class. Traditionally, in botany the term division was used instead of phylum, although from 1993 the International Code of Nomenclature for algae, fungi, depending on definitions, the kingdom Animalia contains approximately 35 phyla, Plantae contains about 12, and Fungi contains around 7. Current research in phylogenetics is uncovering the relationships between phyla, which are contained in larger clades, like Ecdysozoa and Embryophyta, the definitions of zoological phyla have changed from their origins in the six Linnaean classes and the four embranchements of Georges Cuvier. Haeckel introduced the term phylum, based on the Greek word phylon, in plant taxonomy, Eichler classified plants into five groups, named divisions. Informally, phyla can be thought of as grouping organisms based on general specialization of body plan, the most important objective measure in the above definitions is the certain degree—how unrelated do organisms need to be to be members of different phyla. The minimal requirement is that all organisms in a phylum should be more closely related to one another than to any other group. So phyla can be merged or split if it becomes apparent that they are related to one another or not, a definition of a phylum based on body plan has been proposed by paleontologists Graham Budd and Sören Jensen. By Budd and Jensens definition, a phylum is defined by a set of shared by all its living representatives. This approach brings some small problems—for instance, ancestral characters common to most members of a phylum may have been lost by some members, also, this definition is based on an arbitrary point of time, the present. However, as it is based, it is easy to apply to the fossil record. A greater problem is that it relies on a decision about which groups of organisms should be considered as phyla. However, proving that a fossil belongs to the group of a phylum is difficult. Furthermore, organisms in the group of a phylum can possess the body plan of the phylum without all the characteristics necessary to fall within it. This weakens the idea that each of the phyla represents a body plan. A classification using this definition may be affected by the chance survival of rare groups. Representatives of many modern phyla did not appear until long after the Cambrian, the kingdom Plantae is defined in various ways by different biologists. All definitions include the living embryophytes, to which may be added the two green algae divisions, Chlorophyta and Charophyta, to form the clade Viridiplantae. The table below follows the influential Cavalier-Smith system in equating Plantae with Archaeplastida, a group containing Viridiplantae, the definition and classification of plants at the division level also varies from source to source, and has changed progressively in recent years
