# Clymeniida

Clymeniida | |
---|---|

Clymenia laevigata
| |

Scientific classification | |

Kingdom: | Animalia |

Phylum: | Mollusca |

Class: | Cephalopoda |

Subclass: | †Ammonoidea |

Order: | †ClymeniidaHyatt, 1884 |

Suborders | |

**Clymeniida** is an order of ammonoid cephalopods from the Upper Devonian characterized by having an unusual dorsal siphuncle. They measured about 4 cm (1.6 in) in diameter and are restricted to Europe, North Africa, and possibly Australia.

## Morphologic characteristics[edit]

Clymeniids produced a variety of shells ranging from smooth to ribbed and spinose and from evolute with all whorls exposed to strongly involute with the last whorl covering the previous; some were even triangular as viewed from the side (along the axis of coiling). With the exception of the first few chambers, all have a siphuncle that runs along the dorsal margin, along the inside of the coils, opposite that of most ammonoids.

The siphuncle starts off ventrally, like that in other ammonoids, but after the first few septa migrates to a definite and stable dorsal position. Septal necks are retrosiphonate, characteristic of their nautilloid ancestors, and are commonly very long, forming an almost continuous siphunclular tube. Septa, characteristic of ammonoids, are convex toward the front.

## Classification[edit]

Miller, Furnish, and Schindewolf, 1957, in Part L of the Treatise on Invertebrate Paleontology ^{[1]} included three superfamilies in the Clymeniida, the Gonioclymeniaceae, Clymeniaceae, and Parawocklumeriaceae; the Gonoclymeniaceae unites five families, the Clymeniaceae three. The Parawocklumeraceae was established for a single family, the Parawocklumeriidae.

Saunders, Work, and Nikoleava, 1999,^{[2]} divided the Clymeniida into two suborders, the Gonioclymeniina and the Clymeniina; the Gonioclymeniina includes the Sellaclymeniaceae with 7 families and 20 genera, the Gonioclyemniacea, containing a single family with six genera, and the Parawocklumeraceae, also containing a single family, but with only three genera. The Clymeniina combines five families with a total of 36 genera into a single superfamily, the Clymeniaceae.

GONIAT Website ^{[3]} gives two suborders which fall into the Clymeniida, they are the Clymeniina with four superfamilies and Gonioclymeniina, established for a single superfamily. In contrast, Dieter Korn, 2006 divided the Clymeniida into the Clymeniina with superfamilies and families, and Cyrtoclymenia, named by him in 2002. with superfamilies and families.^{[4]} Shevyrev, 2006 on the other hand divided the Clymeniida into the Clymeniina and Gonioclymeniina, as with GONIAT and Saunders, Work, and Nikolaeva ^{[5]}

## Taxonomy[edit]

**Clymeniida**

- Suborder Clymeniina Hyatt, 1884
- Superfamily Clymeniaceae Edwards, 1849
- Family Clymeniidae Edwards, 1849
- Genus
*Aktuboclymenia*Bogoslovsky, 1979 - Genus
*Clymenia*Münster, 1834

- Genus
- Family Kosmoclymeniidae Korn and Price, 1987
- Subfamily Kosmoclymeniinae Korn and Price, 1987
- Genus
*Kosmoclymenia*Schindewolf, 1949 - Genus
*Linguaclymenia*Korn and Price, 1987 - Genus
*Lissoclymenia*Korn and Price, 1987 - Genus
*Muessenbiaergia*Korn and Price, 1987

- Genus
- Subfamily Rodeckiinae Korn 2002
- Genus
*Franconiclymenia*Korn and Price, 1987 - Genus
*Protoxyclymenia*Schindewolf, 1923 - Genus
*Rodeckia*Korn, 2002

- Genus

- Subfamily Kosmoclymeniinae Korn and Price, 1987

- Family Clymeniidae Edwards, 1849
- Superfamily Gonioclymeniaceae Hyatt, 1884
- Family Costaclymeniidae Ruzhencev, 1957
- Genus
*Costaclymenia*Schindewolf, 1920 - Genus
*Endosiphonites*Ansted, 1838

- Genus
- Family Gonioclymeniidae Hyatt, 1884
- Genus
*Finiclymenia*Price and Korn 1989 - Genus
*Gonioclymenia*Sepkoski, Jr., 2002 - Genus
*Kalloclymenia*Wedekind, 1914 - Genus
*Leviclymenia*Korn, 2002 - Genus
*Mesoclymenia*Bogoslovsky, 1981

- Genus
- Family Sellaclymeniidae Schindewolf, 1923
- Genus
*Sellaclymenia*Sepkoski, Jr., 2002

- Genus
- Family Sphenoclymeniidae Korn, 1992
- Genus
*Medioclymenia*Korn, 2002 - Genus
*Sphenoclymenia*Schindewolf, 1920

- Genus

- Family Costaclymeniidae Ruzhencev, 1957
- Superfamily Platyclymeniaceae Wedekind , 1914
- Family Glatziellidae Schindewolf, 1928
- Genus
*Glatziella*Renz, 1914 - Genus
*Liroclymenia*Czarnocki, 1989 - Genus
*Postglatziella*Schindewolf, 1937 - Genus
*Soliclymenia*Schindewolf, 1937

- Genus
- Family Piriclymeniidae Korn, 1992
- Genus
*Ornatoclymenia*Bogoslovsky, 1979 - Genus
*Piriclymenia*Schindewolf, 1937 - Genus
*Sulcoclymenia*Schindewolf, 1923

- Genus
- Family Platyclymeniidae Wedekind, 1914
- Subfamily Nodosoclymeniinae Korn, 2002
- Genus
*Czarnoclymenia*Korn, 1999 - Genus
*Nodosoclymenia*Czarnocki, 1989 - Genus
*Stenoclymenia*Lange, 1929

- Genus
- Subfamily Platyclymeniinae Wedekind, 1914
- Genus
*Fasciclymenia*Korn and Price, 1987 - Genus
*Platyclymenia*Sepkoski, Jr., 2002 - Genus
*Progonioclymenia*Schindewolf, 1937 - Genus
*Spinoclymenia*Bogoslovsky, 1962 - Genus
*Trigonoclymenia*Schindewolf, 1934 - Genus
*Varioclymenia*Wedekind, 1908

- Genus
- Subfamily Pleuroclymeniinae Korn, 2002
- Genus
*Borisiclymenia*Korn, 2002 - Genus
*Nanoclymenia*Korn, 2002 - Genus
*Pleuroclymenia*Schindewolf, 1934 - Genus
*Trochoclymenia*Schindewolf, 1926

- Genus

- Subfamily Nodosoclymeniinae Korn, 2002

- Family Glatziellidae Schindewolf, 1928
- Superfamily Wocklumeriaceae Schindewolf, 1937
- Family Parawocklumeriidae Schindewolf, 1937
- Genus
*Kamptoclymenia*Schindewolf, 1937 - Genus
*Parawocklumeria*Schindewolf, 1926 - Genus
*Tardewocklumeria*Becker, 2000 - Genus
*Triaclymenia*Schindewolf, 1937

- Genus
- Family Wocklumeriidae Schindewolf, 1937
- Genus
*Epiwocklumeria*Schindewolf, 1937 - Genus
*Kielcensia*Czarnocki, 1989 - Genus
*Synwocklumeria*Librovitch, 1957 - Genus
*Wocklumeria*Wedekind, 1918

- Genus

- Family Parawocklumeriidae Schindewolf, 1937

- Superfamily Clymeniaceae Edwards, 1849
- Suborder Cyrtoclymeniina Korn, 2002
- Superfamily Biloclymeniaceae Bogoslovsky, 1955
- Family Biloclymeniidae Bogoslovsky, 1955
- Genus
*Biloclymenia*Schindewolf, 1923 - Genus
*Dimeroclymenia*Czarnocki, 1989 - Genus
*Kiaclymenia*Bogoslovsky, 1965 - Genus
*Rhiphaeoclymenia*Bogoslovsky, 1965

- Genus
- Family Pachyclymeniidae Korn, 1992
- Genus
*Pachyclymenia*Schindewolf, 1937 - Genus
*Uraloclymenia*Bogoslovsky, 1977

- Genus

- Family Biloclymeniidae Bogoslovsky, 1955
- Superfamily Cyrtoclymeniaceae Hyatt, 1884
- Family Carinoclymeniidae Bogoslovsky, 1975
- Genus
*Acriclymenia*Bogoslovsky, 1975 - Genus
*Carinoclymenia*Bogoslovsky, 1965 - Genus
*Karaclymenia*Bogoslovsky, 1983 - Genus
*Pinacoclymenia*Bogoslovsky, 1975

- Genus
- Family Cymaclymeniidae Hyatt 1884
- Subfamily Cymaclymeniinae Hyatt, 1884
- Genus
*Cymaclymenia*Sepkoski, Jr., 2002 - Genus
*Laganoclymenia*Bogoslovsky, 1979 - Genus
*Procymaclymenia*Korn, 2002 - Genus
*Rodachia*Korn, 2002

- Genus
- Subfamily Genuclymeniinae Korn, 2002
- Genus
*Flexiclymenia*Czarnocki, 1989 - Genus
*Genuclymenia*Wedekind, 1908 - Genus
*Siekluckia*Czarnocki, 1989

- Genus

- Subfamily Cymaclymeniinae Hyatt, 1884
- Family Cyrtoclymeniidae Hyatt, 1884
- Genus
*Cyrtoclymenia*Sepkoski, Jr., 2002 - Genus
*Hexaclymenia*Schindewolf, 1923 - Genus
*Praeflexiclymenia*Czarnocki, 1989 - Genus
*Pricella*Korn, 1991 - Genus
*Protactoclymenia*Wedekind, 1908

- Genus
- Family Rectoclymeniidae Schindewolf, 1923
- Genus
*Cteroclymenia*Bogoslovsky, 1979 - Genus
*Falciclymenia*Schindewolf, 1923 - Genus
*Karadzharia*Korn, 2002 - Genus
*Rectoclymenia*Wedekind, 1908

- Genus

- Family Carinoclymeniidae Bogoslovsky, 1975

- Superfamily Biloclymeniaceae Bogoslovsky, 1955
- Suborder
*Incertae sedis*- Genus
*Borkinia* - Genus
*Gyroclymenia* - Genus
*Kazakhoclymenia* - Genus
*Miroclymenia* - Genus
*Schizoclymenia*

- Genus

## References[edit]

**^**Miller, Furnish, and Schindewolf, 1957; Paleozoic Ammonoidea; Treatise on Invertebrate Paleontology, Part-L, Ammonoidea, Geological Society of America & Univ of Kansas.**^**Evolution of Complexity in Paleozoic Ammonoid Sutures; Supplementary Materia [1]**^**GONIAT**^**D. Korn. 2006. Paleozoic ammonoid classification**^**Classification of A. A. Shevyrev 2006