Panama the Republic of Panama, is a country in Central America, bordered by Costa Rica to the west, Colombia to the southeast, the Caribbean Sea to the north, the Pacific Ocean to the south. The capital and largest city is Panama City, whose metropolitan area is home to nearly half the country's 4 million people. Panama was inhabited by indigenous tribes before Spanish colonists arrived in the 16th century, it broke away from Spain in 1821 and joined the Republic of Gran Colombia, a union of Nueva Granada and Venezuela. After Gran Colombia dissolved in 1831, Panama and Nueva Granada became the Republic of Colombia. With the backing of the United States, Panama seceded from Colombia in 1903, allowing the construction of the Panama Canal to be completed by the US Army Corps of Engineers between 1904 and 1914; the 1977 Torrijos–Carter Treaties led to the transfer of the Canal from the United States to Panama on December 31, 1999. Revenue from canal tolls continues to represent a significant portion of Panama's GDP, although commerce and tourism are major and growing sectors.
It is regarded as a high-income country. In 2015 Panama ranked 60th in the world in terms of the Human Development Index. In 2018, Panama was ranked seventh-most competitive economy in Latin America, according to the World Economic Forum's Global Competitiveness Index. Covering around 40 percent of its land area, Panama's jungles are home to an abundance of tropical plants and animals – some of them found nowhere else on the planet. Panama is a founding member of the United Nations and other international organizations such as OAS, LAIA, G77, WHO and NAM; the definite origin of the name Panama is unknown. There are several theories. One postulates that the country was named after a found species of tree. Another that the first settlers arrived in Panama in August, when butterflies abound, that the name means "many butterflies" in one or several of indigenous Amerindian languages that were spoken in the territory prior to Spanish colonization. Most scientifically corroborated theory, that by Panamanian linguists, states that the word is a hispanicization of Kuna language word "bannaba" which means "distant" or "far away".
A relayed legend in Panama is that there was a fishing village that bore the name "Panamá", which purportedly meant "an abundance of fish", when the Spanish colonizers first landed in the area. The exact location of the village is unspecified; the legend is corroborated by Captain Antonio Tello de Guzmán's diary entries, who reports landing at an unnamed village while exploring the Pacific coast of Panama in 1515. In 1517, Don Gaspar de Espinosa, a Spanish lieutenant, decided to settle a post in the same location Guzmán described. In 1519, Pedrarias Dávila decided to establish the Spanish Empire's Pacific port at the site; the new settlement replaced Santa María La Antigua del Darién, which had lost its function within the Crown's global plan after the Spanish exploitation of the riches in the Pacific began. The official definition and origin of the name as promoted by Panama's Ministry of Education is the "abundance of fish and butterflies"; this is the usual description given in social studies textbooks.
At the time of the arrival of the Spanish in the 16th century, the known inhabitants of Panama included the Cuevas and the Coclé tribes. These people have nearly disappeared; the Isthmus of Panama was formed about three million years ago when the land bridge between North and South America became complete, plants and animals crossed it in both directions. The existence of the isthmus affected the dispersal of people and technology throughout the American continent from the appearance of the first hunters and collectors to the era of villages and cities; the earliest discovered artifacts of indigenous peoples in Panama include Paleo-Indian projectile points. Central Panama was home to some of the first pottery-making in the Americas, for example the cultures at Monagrillo, which date back to 2500–1700 BC; these evolved into significant populations best known through their spectacular burials at the Monagrillo archaeological site, their beautiful Gran Coclé style polychrome pottery. The monumental monolithic sculptures at the Barriles site are important traces of these ancient isthmian cultures.
Before Europeans arrived Panama was settled by Chibchan and Cueva peoples. The largest group were the Cueva; the size of the indigenous population of the isthmus at the time of European colonization is uncertain. Estimates range as high as two million people, but more recent studies place that number closer to 200,000. Archaeological finds and testimonials by early European explorers describe diverse native isthmian groups exhibiting cultural variety and suggesting people developed by regular regional routes of commerce; when Panama was colonized, the indigenous peoples fled into nearby islands. Scholars believe that infectious disease was the primary cause of the population decline of American natives; the indigenous peoples had no acquired immunity to diseases, chronic in Eurasian populations for centuries. Rodrigo de Bastidas sailed westward from Venezuela in 1501 in search of gold, became the first European to explore the isthmus of Panama. A year Christopher Columbus visited the isthmus, established a short-lived settlement in the Darien.
Vasco Núñez de Balboa's tortuous
Asparagales is an order of plants in modern classification systems such as the Angiosperm Phylogeny Group and the Angiosperm Phylogeny Web. The order takes its name from the type family Asparagaceae and is placed in the monocots amongst the lilioid monocots; the order has only been recognized in classification systems. It was first put forward by Huber in 1977 and taken up in the Dahlgren system of 1985 and the APG in 1998, 2003 and 2009. Before this, many of its families were assigned to the old order Liliales, a large order containing all monocots with colourful tepals and lacking starch in their endosperm. DNA sequence analysis indicated that many of the taxa included in Liliales should be redistributed over three orders, Liliales and Dioscoreales; the boundaries of the Asparagales and of its families have undergone a series of changes in recent years. In the APG circumscription, Asparagales is the largest order of monocots with 14 families, 1,122 genera, about 36,000 species; the order is circumscribed on the basis of molecular phylogenetics, but is difficult to define morphologically, since its members are structurally diverse.
Most species of Asparagales are herbaceous perennials, although some are climbers and some are tree-like. The order contains many geophytes. According to telomere sequence, at least two evolutionary switch-points happened within the order. Basal sequence is formed by TTTAGGG like in majority of higher plants. Basal motif was changed to vertebrate-like TTAGGG and the most divergent motif CTCGGTTATGGG appears in Allium. One of the defining characteristics of the order is the presence of phytomelanin, a black pigment present in the seed coat, creating a dark crust. Phytomelanin is found in most families of the Asparagales; the leaves of all species form a tight rosette, either at the base of the plant or at the end of the stem, but along the stem. The flowers are not distinctive, being'lily type', with six tepals and up to six stamina; the order is thought to have first diverged from other related monocots some 120–130 million years ago, although given the difficulty in classifying the families involved, estimates are to be uncertain.
From an economic point of view, the order Asparagales is second in importance within the monocots to the order Poales. Species are used as food and flavourings, as cut flowers, as garden ornamentals. Although most species in the order are herbaceous, some no more than 15 cm high, there are a number of climbers, as well as several genera forming trees, which can exceed 10 m in height. Succulent genera occur in several families. All species have a tight cluster of leaves, either at the base of the plant or at the end of a more-or-less woody stem as with Yucca. In some cases the leaves are produced along the stem; the flowers are in the main not distinctive, being of a general'lily type', with six tepals, either free or fused from the base and up to six stamina. They are clustered at the end of the plant stem; the Asparagales are distinguished from the Liliales by the lack of markings on the tepals, the presence of septal nectaries in the ovaries, rather than the bases of the tepals or stamen filaments, the presence of secondary growth.
They are geophytes, but with linear leaves, a lack of fine reticular venation. The seeds characteristically have the external epidermis either obliterated, or if present, have a layer of black carbonaceous phytomelanin in species with dry fruits; the inner part of the seed coat is collapsed, in contrast to Liliales whose seeds have a well developed outer epidermis, lack phytomelanin, display a cellular inner layer. The orders which have been separated from the old Liliales are difficult to characterize. No single morphological character appears to be diagnostic of the order Asparagales; the flowers of Asparagales are of a general type among the lilioid monocots. Compared to Liliales, they have plain tepals without markings in the form of dots. If nectaries are present, they are in the septa of the ovaries rather than at the base of the tepals or stamens; those species which have large dry seeds have a dark, crust-like outer layer containing the pigment phytomelan. However, some species with hairy seeds, berries, or reduced seeds lack this dark pigment in their seed coats.
Phytomelan is not unique to Asparagales but it is common within the order and rare outside it. The inner portion of the seed coat is completely collapsed. In contrast, the morphologically similar seeds of Liliales have no phytomelan, retain a cellular structure in the inner portion of the seed coat. Most monocots are unable to thicken their stems once they have formed, since they lack the cylindrical meristem present in other angiosperm groups. Asparagales have a method of secondary thickening, otherwise only found inDioscorea. In a process called'anomalous secondary growth', they are able to create new
The Orchidaceae are a diverse and widespread family of flowering plants, with blooms that are colourful and fragrant known as the orchid family. Along with the Asteraceae, they are one of the two largest families of flowering plants; the Orchidaceae have about 28,000 accepted species, distributed in about 763 genera. The determination of which family is larger is still under debate, because verified data on the members of such enormous families are continually in flux. Regardless, the number of orchid species nearly equals the number of bony fishes and is more than twice the number of bird species, about four times the number of mammal species; the family encompasses about 6–11% of all seed plants. The largest genera are Bulbophyllum, Epidendrum and Pleurothallis, it includes Vanilla–the genus of the vanilla plant, the type genus Orchis, many cultivated plants such as Phalaenopsis and Cattleya. Moreover, since the introduction of tropical species into cultivation in the 19th century, horticulturists have produced more than 100,000 hybrids and cultivars.
Orchids are distinguished from other plants, as they share some evident, shared derived characteristics, or synapomorphies. Among these are: bilateral symmetry of the flower, many resupinate flowers, a nearly always modified petal, fused stamens and carpels, small seeds. All orchids are perennial herbs, they can grow according to two patterns: Monopodial: The stem grows from a single bud, leaves are added from the apex each year and the stem grows longer accordingly. The stem of orchids with a monopodial growth can reach several metres in length, as in Vanda and Vanilla. Sympodial: Sympodial orchids have a front and a back; the plant produces a series of adjacent shoots, which grow to a certain size and stop growing and are replaced. Sympodial orchids grow laterally following the surface of their support; the growth continues by development of new leads, with their own leaves and roots, sprouting from or next to those of the previous year, as in Cattleya. While a new lead is developing, the rhizome may start its growth again from a so-called'eye', an undeveloped bud, thereby branching.
Sympodial orchids may have visible pseudobulbs joined by a rhizome, which creeps along the top or just beneath the soil. Terrestrial orchids may form corms or tubers; the root caps of terrestrial orchids are white. Some sympodial terrestrial orchids, such as Orchis and Ophrys, have two subterranean tuberous roots. One is used as a food reserve for wintry periods, provides for the development of the other one, from which visible growth develops. In warm and humid climates, many terrestrial orchids do not need pseudobulbs. Epiphytic orchids, those that grow upon a support, have modified aerial roots that can sometimes be a few meters long. In the older parts of the roots, a modified spongy epidermis, called velamen, has the function of absorbing humidity, it can have a silvery-grey, white or brown appearance. In some orchids, the velamen includes spongy and fibrous bodies near the passage cells, called tilosomes; the cells of the root epidermis grow at a right angle to the axis of the root to allow them to get a firm grasp on their support.
Nutrients for epiphytic orchids come from mineral dust, organic detritus, animal droppings and other substances collecting among on their supporting surfaces. The base of the stem of sympodial epiphytes, or in some species the entire stem, may be thickened to form a pseudobulb that contains nutrients and water for drier periods; the pseudobulb has a smooth surface with lengthwise grooves, can have different shapes conical or oblong. Its size is variable; some Dendrobium species have long, canelike pseudobulbs with short, rounded leaves over the whole length. With ageing, the pseudobulb becomes dormant. At this stage, it is called a backbulb. Backbulbs still hold nutrition for the plant, but a pseudobulb takes over, exploiting the last reserves accumulated in the backbulb, which dies off, too. A pseudobulb lives for about five years. Orchids without noticeable pseudobulbs are said to have growths, an individual component of a sympodial plant. Like most monocots, orchids have simple leaves with parallel veins, although some Vanilloideae have reticulate venation.
Leaves may be ovate, lanceolate, or orbiculate, variable in size on the individual plant. Their characteristics are diagnostic, they are alternate on the stem folded lengthwise along the centre, have no stipules. Orchid leaves have siliceous bodies called stegmata in the vascular bundle sheaths and are fibrous; the structure of the leaves corresponds to the specific habitat of the plant. Species that bask in sunlight, or grow on sites which can be very dry, have thick, leathery leaves and the laminae are covered by a waxy cuticle to retain their necessary water supply. Shade-loving species, on the other hand, have thin leaves; the leaves of most orchids are perennial, that is, they live for several years, while others those with plicate leaves as in Catasetum, shed them annually and de
The tribe Epidendreae of the Orchidaceae comprises six subtribes: Bletiinae sensu MMIV, which contains only the genera Basiphyllaea and Hexalectris Chysinae Coeliinae Laeliinae Pleurothallidinae Ponerinae
Plants are multicellular, predominantly photosynthetic eukaryotes of the kingdom Plantae. Plants were treated as one of two kingdoms including all living things that were not animals, all algae and fungi were treated as plants. However, all current definitions of Plantae exclude the fungi and some algae, as well as the prokaryotes. By one definition, plants form the clade Viridiplantae, a group that includes the flowering plants and other gymnosperms and their allies, liverworts and the green algae, but excludes the red and brown algae. Green plants obtain most of their energy from sunlight via photosynthesis by primary chloroplasts that are derived from endosymbiosis with cyanobacteria, their chloroplasts contain b, which gives them their green color. Some plants are parasitic or mycotrophic and have lost the ability to produce normal amounts of chlorophyll or to photosynthesize. Plants are characterized by sexual reproduction and alternation of generations, although asexual reproduction is common.
There are about 320 thousand species of plants, of which the great majority, some 260–290 thousand, are seed plants. Green plants provide a substantial proportion of the world's molecular oxygen and are the basis of most of Earth's ecosystems on land. Plants that produce grain and vegetables form humankind's basic foods, have been domesticated for millennia. Plants have many cultural and other uses, as ornaments, building materials, writing material and, in great variety, they have been the source of medicines and psychoactive drugs; the scientific study of plants is known as a branch of biology. All living things were traditionally placed into one of two groups and animals; this classification may date from Aristotle, who made the distincton between plants, which do not move, animals, which are mobile to catch their food. Much when Linnaeus created the basis of the modern system of scientific classification, these two groups became the kingdoms Vegetabilia and Animalia. Since it has become clear that the plant kingdom as defined included several unrelated groups, the fungi and several groups of algae were removed to new kingdoms.
However, these organisms are still considered plants in popular contexts. The term "plant" implies the possession of the following traits multicellularity, possession of cell walls containing cellulose and the ability to carry out photosynthesis with primary chloroplasts; when the name Plantae or plant is applied to a specific group of organisms or taxon, it refers to one of four concepts. From least to most inclusive, these four groupings are: Another way of looking at the relationships between the different groups that have been called "plants" is through a cladogram, which shows their evolutionary relationships; these are not yet settled, but one accepted relationship between the three groups described above is shown below. Those which have been called "plants" are in bold; the way in which the groups of green algae are combined and named varies between authors. Algae comprise several different groups of organisms which produce food by photosynthesis and thus have traditionally been included in the plant kingdom.
The seaweeds range from large multicellular algae to single-celled organisms and are classified into three groups, the green algae, red algae and brown algae. There is good evidence that the brown algae evolved independently from the others, from non-photosynthetic ancestors that formed endosymbiotic relationships with red algae rather than from cyanobacteria, they are no longer classified as plants as defined here; the Viridiplantae, the green plants – green algae and land plants – form a clade, a group consisting of all the descendants of a common ancestor. With a few exceptions, the green plants have the following features in common, they undergo closed mitosis without centrioles, have mitochondria with flat cristae. The chloroplasts of green plants are surrounded by two membranes, suggesting they originated directly from endosymbiotic cyanobacteria. Two additional groups, the Rhodophyta and Glaucophyta have primary chloroplasts that appear to be derived directly from endosymbiotic cyanobacteria, although they differ from Viridiplantae in the pigments which are used in photosynthesis and so are different in colour.
These groups differ from green plants in that the storage polysaccharide is floridean starch and is stored in the cytoplasm rather than in the plastids. They appear to have had a common origin with Viridiplantae and the three groups form the clade Archaeplastida, whose name implies that their chloroplasts were derived from a single ancient endosymbiotic event; this is the broadest modern definition of the term'plant'. In contrast, most other algae not only have different pigments but have chloroplasts with three or four surrounding membranes, they are not close relatives of the Archaeplastida having acquired chloroplasts separately from ingested or symbiotic green and red algae. They are thus not included in the broadest modern definition of the plant kingdom, although they were in the past; the green plants or Viridiplantae were traditionally divided into the green algae (including
Colombia the Republic of Colombia, is a sovereign state situated in the northwest of South America, with territories in Central America. Colombia shares a border to the northwest with Panama, to the east with Venezuela and Brazil and to the south with Ecuador and Peru, it shares its maritime limits with Costa Rica, Honduras, Jamaica and the Dominican Republic. Colombia is a unitary, constitutional republic comprising thirty-two departments, with the capital in Bogota. Colombia has been inhabited by various indigenous peoples since 12,000 BCE, including the Muisca and the Tairona, along with the Inca Empire that expanded to the southwest of the country; the Spanish arrived in 1499 and by the mid-16th century conquered and colonized much of the region, establishing the New Kingdom of Granada, with Santafé de Bogotá as its capital. Independence from Spain was achieved in 1819, but by 1830 the "Gran Colombia" Federation was dissolved, with what is now Colombia and Panama emerging as the Republic of New Granada.
The new nation experimented with federalism as the Granadine Confederation, the United States of Colombia, before the Republic of Colombia was declared in 1886. Panama seceded in 1903. Beginning in the 1960s, the country suffered from an asymmetric low-intensity armed conflict and rampant political violence, both of which escalated in the 1990s. Since 2005, there has been significant improvement in security and rule of law. Colombia is one of the most ethnically and linguistically diverse countries in the world, with its rich cultural heritage reflecting influences by indigenous peoples, European settlement, forced African migration, immigration from Europe and the Middle East. Urban centres are located in the highlands of the Andes mountains and the Caribbean coast. Colombia is among the world's 17 megadiverse countries, the most densely biodiverse per square kilometer. Colombia is a middle power and regional actor in Latin America, it is part of the CIVETS group of six leading emerging markets and a member of the UN, the WTO, the OAS, the Pacific Alliance, other international organizations.
Colombia's diversified economy is the fourth largest in Latin America, with macroeconomic stability and favorable long-term growth prospects. The name "Colombia" is derived from the last name of Christopher Columbus, it was conceived by the Venezuelan revolutionary Francisco de Miranda as a reference to all the New World, but to those portions under Spanish rule. The name was adopted by the Republic of Colombia of 1819, formed from the territories of the old Viceroyalty of New Granada; when Venezuela and Cundinamarca came to exist as independent states, the former Department of Cundinamarca adopted the name "Republic of New Granada". New Granada changed its name in 1858 to the Granadine Confederation. In 1863 the name was again changed, this time to United States of Colombia, before adopting its present name – the Republic of Colombia – in 1886. To refer to this country, the Colombian government uses the terms Colombia and República de Colombia. Owing to its location, the present territory of Colombia was a corridor of early human migration from Mesoamerica and the Caribbean to the Andes and Amazon basin.
The oldest archaeological finds are from the Pubenza and El Totumo sites in the Magdalena Valley 100 kilometres southwest of Bogotá. These sites date from the Paleoindian period. At Puerto Hormiga and other sites, traces from the Archaic Period have been found. Vestiges indicate that there was early occupation in the regions of El Abra and Tequendama in Cundinamarca; the oldest pottery discovered in the Americas, found at San Jacinto, dates to 5000–4000 BCE. Indigenous people inhabited the territory, now Colombia by 12,500 BCE. Nomadic hunter-gatherer tribes at the El Abra, Tibitó and Tequendama sites near present-day Bogotá traded with one another and with other cultures from the Magdalena River Valley. Between 5000 and 1000 BCE, hunter-gatherer tribes transitioned to agrarian societies. Beginning in the 1st millennium BCE, groups of Amerindians including the Muisca, Zenú, Tairona developed the political system of cacicazgos with a pyramidal structure of power headed by caciques; the Muisca inhabited the area of what is now the Departments of Boyacá and Cundinamarca high plateau where they formed the Muisca Confederation.
They farmed maize, potato and cotton, traded gold, blankets, ceramic handicrafts and rock salt with neighboring nations. The Tairona inhabited northern Colombia in the isolated mountain range of Sierra Nevada de Santa Marta; the Quimbaya inhabited regions of the Cauca River Valley between the Western and Central Ranges of the Colombian Andes. Most of the Amerindians practiced agriculture and the social structure of each indigenous community was different; some groups of indigenous people such as the Caribs lived in a state of permanent war, but others had less bellicose attitudes. The Incas expanded their empire onto the southwest part of the country. Alonso de Ojeda reached the Guajira Peninsula in 1499. Spanish explorers, led by Rodrigo de Bastidas, made the first exploration
The flowering plants known as angiosperms, Angiospermae or Magnoliophyta, are the most diverse group of land plants, with 64 orders, 416 families 13,164 known genera and c. 369,000 known species. Like gymnosperms, angiosperms are seed-producing plants. However, they are distinguished from gymnosperms by characteristics including flowers, endosperm within the seeds, the production of fruits that contain the seeds. Etymologically, angiosperm means a plant; the term comes from the Greek words sperma. The ancestors of flowering plants diverged from gymnosperms in the Triassic Period, 245 to 202 million years ago, the first flowering plants are known from 160 mya, they diversified extensively during the Early Cretaceous, became widespread by 120 mya, replaced conifers as the dominant trees from 100 to 60 mya. Angiosperms differ from other seed plants in several ways, described in the table below; these distinguishing characteristics taken together have made the angiosperms the most diverse and numerous land plants and the most commercially important group to humans.
Angiosperm stems are made up of seven layers. The amount and complexity of tissue-formation in flowering plants exceeds that of gymnosperms; the vascular bundles of the stem are arranged such that the phloem form concentric rings. In the dicotyledons, the bundles in the young stem are arranged in an open ring, separating a central pith from an outer cortex. In each bundle, separating the xylem and phloem, is a layer of meristem or active formative tissue known as cambium. By the formation of a layer of cambium between the bundles, a complete ring is formed, a regular periodical increase in thickness results from the development of xylem on the inside and phloem on the outside; the soft phloem becomes crushed, but the hard wood persists and forms the bulk of the stem and branches of the woody perennial. Owing to differences in the character of the elements produced at the beginning and end of the season, the wood is marked out in transverse section into concentric rings, one for each season of growth, called annual rings.
Among the monocotyledons, the bundles are more numerous in the young stem and are scattered through the ground tissue. They once formed the stem increases in diameter only in exceptional cases; the characteristic feature of angiosperms is the flower. Flowers show remarkable variation in form and elaboration, provide the most trustworthy external characteristics for establishing relationships among angiosperm species; the function of the flower is to ensure fertilization of the ovule and development of fruit containing seeds. The floral apparatus may arise terminally from the axil of a leaf; as in violets, a flower arises singly in the axil of an ordinary foliage-leaf. More the flower-bearing portion of the plant is distinguished from the foliage-bearing or vegetative portion, forms a more or less elaborate branch-system called an inflorescence. There are two kinds of reproductive cells produced by flowers. Microspores, which will divide to become pollen grains, are the "male" cells and are borne in the stamens.
The "female" cells called megaspores, which will divide to become the egg cell, are contained in the ovule and enclosed in the carpel. The flower may consist only of these parts, as in willow, where each flower comprises only a few stamens or two carpels. Other structures are present and serve to protect the sporophylls and to form an envelope attractive to pollinators; the individual members of these surrounding structures are known as petals. The outer series is green and leaf-like, functions to protect the rest of the flower the bud; the inner series is, in general, white or brightly colored, is more delicate in structure. It functions to attract bird pollinators. Attraction is effected by color and nectar, which may be secreted in some part of the flower; the characteristics that attract pollinators account for the popularity of flowers and flowering plants among humans. While the majority of flowers are perfect or hermaphrodite, flowering plants have developed numerous morphological and physiological mechanisms to reduce or prevent self-fertilization.
Heteromorphic flowers have short carpels and long stamens, or vice versa, so animal pollinators cannot transfer pollen to the pistil. Homomorphic flowers may employ a biochemical mechanism called self-incompatibility to discriminate between self and non-self pollen grains. In other species, the male and female parts are morphologically separated, developing on different flowers; the botanical term "Angiosperm", from the Ancient Greek αγγείον, angeíon and σπέρμα, was coined in the form Angiospermae by Paul Hermann in 1690, as the name of one of his primary divisions of the plant kingdom. This included flowering plants possessing seeds enclosed in capsules, distinguished from his Gymnospermae, or flowering plants with achenial or schizo-carpic fruits, the whole fruit or each of its pieces being here regarded as a seed and naked; the term and its antonym were maintained by Carl Linnaeus with the same sense, but with restricted application, in the names of the orders of his class Didynamia. Its use with any