Cronartium is a genus of rust fungi in the family Cronartiaceae. They are heteroecious rusts with two alternating hosts a pine and a flowering plant, up to five spore stages. Many of the species are plant diseases of major economic importance. Species and natural distributionCronartium appalachianum: Pinus virginiana, Santalaceae. Eastern North America. Cronartium arizonicum: Pinus ponderosa and related pines, Scrophulariaceae. Western North America. Cronartium comandrae: Pinus subgenus Pinus, Santalaceae. North America. Cronartium comptoniae: Pinus subgenus Pinus, Myricaceae. North America. Cronartium conigenum: Pinus subgenus Pinus, Fagaceae. Southwestern North America. Cronartium flaccidum: Pinus subgenus Pinus, several families. Europe, Asia. Cronartium occidentale: Strobus subgenus Strobus, Saxifragaceae. Southwestern North America. Cronartium orientale: Pinus subgenus Pinus Quercus. Japan. Cronartium quercuum: Pinus subgenus Pinus, Fagaceae. North America, Asia. Cronartium ribicola: Pinus subgenus Strobus, Grossulariaceae.
Europe, Asia. Cronartium stalactiforme: Pinus subgenus Pinus, Scrophulariaceae. North America. Cronartium strobilinum: Pinus subgenus Pinus, Fagaceae. Southeastern North America; some of the species have been introduced accidentally outside of their natural ranges, notably C. ribicola into North America, where it is an invasive species causing heavy mortality in several pines which have little or no resistance to it. Millar, C. I. & Kinloch, B. B.. Taxonomy and coevolution of pines and their stem rusts. Pp. 1–38 in: Hiratsuka, Y. et al.. Rusts of pine. Proceedings of the IUFRO Rusts of Pine Working Party Conference. Forestry Canada Information Report NOR-X-317. Forestry Images: photos of some Cronartium rusts
Western white pine
Western white pine called silver pine, California mountain pine, in the family Pinaceae, is a species of pine that occurs in the mountains of the western United States and Canada the Sierra Nevada, the Cascade Range, the Coast Range, the northern Rocky Mountains. The tree extends down to sea level in many areas in Oregon and Washington, it is the state tree of Idaho, is sometimes known as the Idaho pine. Western white pine is a large tree growing to 30–50 metres and exceptionally up to 70 metres tall, it is a member of the white pine group, Pinus subgenus Strobus, like all members of that group, the leaves are in fascicles of five, with a deciduous sheath. The needles are finely serrated, 5–13 cm long; the cones are long and slender, 12–32 cm long and 3–4 cm broad, opening to 5–8 cm broad. The seeds are small, 4–7 mm long, have a long slender wing 15–22 mm long, it is related to the Eastern white pine, differing from it in having larger cones longer-lasting leaves with more prominent stomatal bands, a somewhat denser and narrower habit.
The branches are borne in regular whorls, produced at the rate of one a year. It is grown as an ornamental tree, but has been logged throughout much of its range in the past. Western white pine has been affected by the white pine blister rust, a fungus, accidentally introduced from Europe in 1909; the United States Forest Service estimates that 90% of the Western white pines have been killed by the blister rust west of the Cascades. Large stands have been succeeded by non-pine species; the rust has killed much of the whitebark pine outside of California. Blister rust is less severe in California, Western white and whitebark pines have survived there in great numbers. Resistance to the blister rust is genetic, due to Western white pine's genetic variability some individuals are unaffected by the rust; the Forest Service has a program for locating and breeding rust-resistant Western white pine and sugar pine. Seedlings of these trees have been introduced into the wild. US Forest Service Dorena Tree Improvement Center Chase, J. Smeaton.
Cone-bearing Trees of the California Mountains. Chicago: A. C. McClurg & Co. p. 99. LCCN 11004975. OCLC 3477527. LCC QK495. C75 C4, with illustrations by Carl Eytel - Kurut, Gary F. "Carl Eytel: Southern California Desert Artist", California State Library Foundation, Bulletin No. 95, pp. 17-20 retrieved Nov. 13, 2011 Jepson Manual treatment - Pinus monticola Pinus monticola - U. C. Photos Gallery
Wikispecies is a wiki-based online project supported by the Wikimedia Foundation. Its aim is to create a comprehensive free content catalogue of all species. Jimmy Wales stated that editors are not required to fax in their degrees, but that submissions will have to pass muster with a technical audience. Wikispecies is available under the GNU Free Documentation License and CC BY-SA 3.0. Started in September 2004, with biologists across the world invited to contribute, the project had grown a framework encompassing the Linnaean taxonomy with links to Wikipedia articles on individual species by April 2005. Benedikt Mandl co-ordinated the efforts of several people who are interested in getting involved with the project and contacted potential supporters in early summer 2004. Databases were evaluated and the administrators contacted, some of them have agreed on providing their data for Wikispecies. Mandl defined two major tasks: Figure out how the contents of the data base would need to be presented—by asking experts, potential non-professional users and comparing that with existing databases Figure out how to do the software, which hardware is required and how to cover the costs—by asking experts, looking for fellow volunteers and potential sponsorsAdvantages and disadvantages were discussed by the wikimedia-I mailing list.
The board of directors of the Wikimedia Foundation voted by 4 to 0 in favor of the establishment of a Wikispecies. The project is hosted at species.wikimedia.org. It was merged to a sister project of Wikimedia Foundation on September 14, 2004. On October 10, 2006, the project exceeded 75,000 articles. On May 20, 2007, the project exceeded 100,000 articles with a total of 5,495 registered users. On September 8, 2008, the project exceeded 150,000 articles with a total of 9,224 registered users. On October 23, 2011, the project reached 300,000 articles. On June 16, 2014, the project reached 400,000 articles. On January 7, 2017, the project reached 500,000 articles. On October 30, 2018, the project reached 600,000 articles, a total of 1.12 million pages. Wikispecies comprises taxon pages, additionally pages about synonyms, taxon authorities, taxonomical publications, institutions or repositories holding type specimen. Wikispecies asks users to use images from Wikimedia Commons. Wikispecies does not allow the use of content.
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The flowering plants known as angiosperms, Angiospermae or Magnoliophyta, are the most diverse group of land plants, with 64 orders, 416 families 13,164 known genera and c. 369,000 known species. Like gymnosperms, angiosperms are seed-producing plants. However, they are distinguished from gymnosperms by characteristics including flowers, endosperm within the seeds, the production of fruits that contain the seeds. Etymologically, angiosperm means a plant; the term comes from the Greek words sperma. The ancestors of flowering plants diverged from gymnosperms in the Triassic Period, 245 to 202 million years ago, the first flowering plants are known from 160 mya, they diversified extensively during the Early Cretaceous, became widespread by 120 mya, replaced conifers as the dominant trees from 100 to 60 mya. Angiosperms differ from other seed plants in several ways, described in the table below; these distinguishing characteristics taken together have made the angiosperms the most diverse and numerous land plants and the most commercially important group to humans.
Angiosperm stems are made up of seven layers. The amount and complexity of tissue-formation in flowering plants exceeds that of gymnosperms; the vascular bundles of the stem are arranged such that the phloem form concentric rings. In the dicotyledons, the bundles in the young stem are arranged in an open ring, separating a central pith from an outer cortex. In each bundle, separating the xylem and phloem, is a layer of meristem or active formative tissue known as cambium. By the formation of a layer of cambium between the bundles, a complete ring is formed, a regular periodical increase in thickness results from the development of xylem on the inside and phloem on the outside; the soft phloem becomes crushed, but the hard wood persists and forms the bulk of the stem and branches of the woody perennial. Owing to differences in the character of the elements produced at the beginning and end of the season, the wood is marked out in transverse section into concentric rings, one for each season of growth, called annual rings.
Among the monocotyledons, the bundles are more numerous in the young stem and are scattered through the ground tissue. They once formed the stem increases in diameter only in exceptional cases; the characteristic feature of angiosperms is the flower. Flowers show remarkable variation in form and elaboration, provide the most trustworthy external characteristics for establishing relationships among angiosperm species; the function of the flower is to ensure fertilization of the ovule and development of fruit containing seeds. The floral apparatus may arise terminally from the axil of a leaf; as in violets, a flower arises singly in the axil of an ordinary foliage-leaf. More the flower-bearing portion of the plant is distinguished from the foliage-bearing or vegetative portion, forms a more or less elaborate branch-system called an inflorescence. There are two kinds of reproductive cells produced by flowers. Microspores, which will divide to become pollen grains, are the "male" cells and are borne in the stamens.
The "female" cells called megaspores, which will divide to become the egg cell, are contained in the ovule and enclosed in the carpel. The flower may consist only of these parts, as in willow, where each flower comprises only a few stamens or two carpels. Other structures are present and serve to protect the sporophylls and to form an envelope attractive to pollinators; the individual members of these surrounding structures are known as petals. The outer series is green and leaf-like, functions to protect the rest of the flower the bud; the inner series is, in general, white or brightly colored, is more delicate in structure. It functions to attract bird pollinators. Attraction is effected by color and nectar, which may be secreted in some part of the flower; the characteristics that attract pollinators account for the popularity of flowers and flowering plants among humans. While the majority of flowers are perfect or hermaphrodite, flowering plants have developed numerous morphological and physiological mechanisms to reduce or prevent self-fertilization.
Heteromorphic flowers have short carpels and long stamens, or vice versa, so animal pollinators cannot transfer pollen to the pistil. Homomorphic flowers may employ a biochemical mechanism called self-incompatibility to discriminate between self and non-self pollen grains. In other species, the male and female parts are morphologically separated, developing on different flowers; the botanical term "Angiosperm", from the Ancient Greek αγγείον, angeíon and σπέρμα, was coined in the form Angiospermae by Paul Hermann in 1690, as the name of one of his primary divisions of the plant kingdom. This included flowering plants possessing seeds enclosed in capsules, distinguished from his Gymnospermae, or flowering plants with achenial or schizo-carpic fruits, the whole fruit or each of its pieces being here regarded as a seed and naked; the term and its antonym were maintained by Carl Linnaeus with the same sense, but with restricted application, in the names of the orders of his class Didynamia. Its use with any
In biological classification zoology, the type genus is the genus which defines a biological family and the root of the family name. According to the International Code of Zoological Nomenclature, "The name-bearing type of a nominal family-group taxon is a nominal genus called the'type genus'; the type genus for a family-group name is the genus that provided the stem to, added the ending -idae. Example: The family name Cricetidae has as its type genus the genus Cricetus Leske, 1779. In botanical nomenclature, the phrase "type genus" is used, unofficially, as a term of convenience. In the ICN this phrase has no status; the code uses type specimens for ranks up to family, types are optional for higher ranks. The Code does not refer to the genus containing that type as a "type genus". Example: "Poa is the type genus of the family Poaceae and of the order Poales" is another way of saying that the names Poaceae and Poales are based on the generic name Poa. Principle of Typification Type Type species
Basidiomycota is one of two large divisions that, together with the Ascomycota, constitute the subkingdom Dikarya within the kingdom Fungi. More Basidiomycota includes these groups: mushrooms, stinkhorns, bracket fungi, other polypores, jelly fungi, chanterelles, earth stars, bunts, mirror yeasts, the human pathogenic yeast Cryptococcus. Basidiomycota are filamentous fungi composed of hyphae and reproduce sexually via the formation of specialized club-shaped end cells called basidia that bear external meiospores; these specialized spores are called basidiospores. However, some Basidiomycota reproduce asexually in exclusively. Basidiomycota that reproduce asexually can be recognized as members of this division by gross similarity to others, by the formation of a distinctive anatomical feature, cell wall components, definitively by phylogenetic molecular analysis of DNA sequence data; the most recent classification adopted by a coalition of 67 mycologists recognizes three subphyla and two other class level taxa outside of these, among the Basidiomycota.
As now classified, the subphyla join and cut across various obsolete taxonomic groups commonly used to describe Basidiomycota. According to a 2008 estimate, Basidiomycota comprise three subphyla 16 classes, 52 orders, 177 families, 1,589 genera, 31,515 species. Traditionally, the Basidiomycota were divided into two classes, now obsolete: Homobasidiomycetes, including true mushrooms Heterobasidiomycetes, including the jelly and smut fungiPreviously the entire Basidiomycota were called Basidiomycetes, an invalid class level name coined in 1959 as a counterpart to the Ascomycetes, when neither of these taxa were recognized as divisions; the terms basidiomycetes and ascomycetes are used loosely to refer to Basidiomycota and Ascomycota. They are abbreviated to "basidios" and "ascos" as mycological slang; the Agaricomycotina include what had been called the Hymenomycetes, the Gasteromycetes, as well as most of the jelly fungi. The three classes in the Agaricomycotina are the Agaricomycetes, the Dacrymycetes, the Tremellomycetes.
The class Wallemiomycetes is not yet placed in a subdivision, but recent genomic evidence suggests that it is a sister group of Agaricomycotina. The Pucciniomycotina include the rust fungi, the insect parasitic/symbiotic genus Septobasidium, a former group of smut fungi, a mixture of odd, infrequently seen, or recognized fungi parasitic on plants; the eight classes in the Pucciniomycotina are Agaricostilbomycetes, Atractiellomycetes, Classiculomycetes, Cryptomycocolacomycetes, Cystobasidiomycetes, Microbotryomycetes and Pucciniomycetes. The Ustilaginomycotina are most of the Exobasidiales; the classes of the Ustilaginomycotina are the Exobasidiomycetes, the Entorrhizomycetes, the Ustilaginomycetes. Unlike animals and plants which have recognizable male and female counterparts, Basidiomycota tend to have mutually indistinguishable, compatible haploids which are mycelia being composed of filamentous hyphae. Haploid Basidiomycota mycelia fuse via plasmogamy and the compatible nuclei migrate into each other's mycelia and pair up with the resident nuclei.
Karyogamy is delayed, called a dikaryon. The hyphae are said to be dikaryotic. Conversely, the haploid mycelia are called monokaryons; the dikaryotic mycelium is more vigorous than the individual monokaryotic mycelia, proceeds to take over the substrate in which they are growing. The dikaryons can be decades, or centuries; the monokaryons are neither female. They have either a tetrapolar mating system; this results in the fact that following meiosis, the resulting haploid basidiospores and resultant monokaryons, have nuclei that are compatible with 50% or 25% of their sister basidiospores because the mating genes must differ for them to be compatible. However, there are sometimes more than two possible alleles for a given locus, in such species, depending on the specifics, over 90% of monokaryons could be compatible with each other; the maintenance of the dikaryotic status in dikaryons in many Basidiomycota is facilitated by the formation of clamp connections that physically appear to help coordinate and re-establish pairs of compatible nuclei following synchronous mitotic nuclear divisions.
Variations are multiple. In a typical Basidiomycota lifecycle the long lasting dikaryons periodically produce basidia, the specialized club-shaped end cells, in which a pair of compatible nuclei fuse to form a diploid cell. Meiosis follows shortly with the production of 4 haploid nuclei that migrate into 4 external apical basidiospores. Variations occur, however; the basidiospores are ballistic, hence they are sometimes called ballistospores. In most species, the basidiospores disperse and each
A pine is any conifer in the genus Pinus of the family Pinaceae. Pinus is the sole genus in the subfamily Pinoideae; the Plant List compiled by the Royal Botanic Gardens and Missouri Botanical Garden accepts 126 species names of pines as current, together with 35 unresolved species and many more synonyms. The modern English name "pine" derives from Latin pinus, which some have traced to the Indo-European base *pīt- ‘resin’. Before the 19th century, pines were referred to as firs. In some European languages, Germanic cognates of the Old Norse name are still in use for pines—in Danish fyr, in Norwegian fura/fure/furu, Swedish fura/furu, Dutch vuren, German Föhre—but in modern English, fir is now restricted to fir and Douglas fir. Pine trees are evergreen, coniferous resinous trees growing 3–80 m tall, with the majority of species reaching 15–45 m tall; the smallest are Siberian dwarf pine and Potosi pinyon, the tallest is an 81.79 m tall ponderosa pine located in southern Oregon's Rogue River-Siskiyou National Forest.
Pines are long lived and reach ages of 100–1,000 years, some more. The longest-lived is Pinus longaeva. One individual of this species, dubbed "Methuselah", is one of the world's oldest living organisms at around 4,600 years old; this tree can be found in the White Mountains of California. An older tree, now cut down, was dated at 4,900 years old, it was discovered in a grove beneath Wheeler Peak and it is now known as "Prometheus" after the Greek immortal. The bark of most pines is thick and scaly; the branches are produced in regular "pseudo whorls" a tight spiral but appearing like a ring of branches arising from the same point. Many pines are uninodal, producing just one such whorl of branches each year, from buds at the tip of the year's new shoot, but others are multinodal, producing two or more whorls of branches per year; the spiral growth of branches and cone scales may be arranged in Fibonacci number ratios. The new spring shoots are sometimes called "candles"; these "candles" offer foresters a means to evaluate fertility of the vigour of the trees.
Pines have four types of leaf: Seed leaves on seedlings are borne in a whorl of 4–24. Juvenile leaves, which follow on seedlings and young plants, are 2–6 cm long, green or blue-green, arranged spirally on the shoot; these are produced for six months to five years longer. Scale leaves, similar to bud scales, are small and not photosynthetic, arranged spirally like the juvenile leaves. Needles, the adult leaves, are green and bundled in clusters called fascicles; the needles can number from one to seven per fascicle, but number from two to five. Each fascicle is produced from a small bud on a dwarf shoot in the axil of a scale leaf; these bud scales remain on the fascicle as a basal sheath. The needles persist depending on species. If a shoot is damaged, the needle fascicles just below the damage will generate a bud which can replace the lost leaves. Pines are monoecious, having the male and female cones on the same tree, though a few species are sub-dioecious, with individuals predominantly, but not wholly, single-sex.
The male cones are small 1–5 cm long, only present for a short period, falling as soon as they have shed their pollen. The female cones take 1.5–3 years to mature after pollination, with actual fertilization delayed one year. At maturity the female cones are 3–60 cm long; each cone has numerous spirally. The seeds are small and winged, are anemophilous, but some are larger and have only a vestigial wing, are bird-dispersed. At maturity, the cones open to release the seeds, but in some of the bird-dispersed species, the seeds are only released by the bird breaking the cones open. In others, the seeds are stored in closed cones for many years until an environmental cue triggers the cones to open, releasing the seeds; the most common form of serotiny is pyriscence, in which a resin binds the cones shut until melted by a forest fire. Pines are gymnosperms; the genus is divided into two subgenera, which can be distinguished by cone and leaf characters: Pinus subg. Pinus, the yellow, or hard pine group with harder wood and two or three needles per fascicle Pinus subg.
Strobus, the white, or soft pine group with softer wood and five needles per fascicle Pines are native to the Northern Hemisphere, in a few parts of the tropics in the Southern Hemisphere. Most regions of the Northern Hemisphere host some native species of pines. One species crosses the equator in Sumatra to 2°S. In North America, various species occur in regions at latitudes from as far north as 66°N to as far south as 12°N. Pines may be found in a large variety of environments, ranging from semi-arid desert to rainforests, from sea level up to 5,200 metres, from the coldest to the hottest environments on Earth, they occur in mountainous areas with favorable soils and at least some water. Various species have been introduced to temperate and subtropical regions of both hemisp