A tooth is a hard, calcified structure found in the jaws of many vertebrates and used to break down food. Some animals carnivores use teeth for hunting or for defensive purposes; the roots of teeth are covered by gums. Teeth hardness; the cellular tissues that become teeth originate from the embryonic germ layer, the ectoderm. The general structure of teeth is similar across the vertebrates, although there is considerable variation in their form and position; the teeth of mammals have deep roots, this pattern is found in some fish, in crocodilians. In most teleost fish, the teeth are attached to the outer surface of the bone, while in lizards they are attached to the inner surface of the jaw by one side. In cartilaginous fish, such as sharks, the teeth are attached by tough ligaments to the hoops of cartilage that form the jaw; some animals develop only one set of teeth. Sharks, for example, grow a new set of teeth. Rodent incisors grow and wear away continually through gnawing, which helps maintain constant length.
The industry of the beaver is due in part to this qualification. Many rodents such as voles and guinea pigs, but not mice, as well as leporidae like rabbits, have continuously growing molars in addition to incisors. Teeth are not always attached to the jaw. In many reptiles and fish, teeth are attached to the palate or to the floor of the mouth, forming additional rows inside those on the jaws proper; some teleosts have teeth in the pharynx. While not true teeth in the usual sense, the dermal denticles of sharks are identical in structure and are to have the same evolutionary origin. Indeed, teeth appear to have first evolved in sharks, are not found in the more primitive jawless fish – while lampreys do have tooth-like structures on the tongue, these are in fact, composed of keratin, not of dentine or enamel, bear no relationship to true teeth. Though "modern" teeth-like structures with dentine and enamel have been found in late conodonts, they are now supposed to have evolved independently of vertebrates' teeth.
Living amphibians have small teeth, or none at all, since they feed only on soft foods. In reptiles, teeth are simple and conical in shape, although there is some variation between species, most notably the venom-injecting fangs of snakes; the pattern of incisors, canines and molars is found only in mammals, to varying extents, in their evolutionary ancestors. The numbers of these types of teeth vary between species; the genes governing tooth development in mammals are homologous to those involved in the development of fish scales. Study of a tooth plate of a fossil of the extinct fish Romundina stellina showed that the teeth and scales were made of the same tissues found in mammal teeth, lending support to the theory that teeth evolved as a modification of scales. Teeth are among the most distinctive features of mammal species. Paleontologists use teeth to determine their relationships; the shape of the animal's teeth are related to its diet. For example, plant matter is hard to digest, so herbivores have many molars for chewing and grinding.
Carnivores, on the other hand, have canine teeth to tear meat. Mammals, in general, are diphyodont. In humans, the first set starts to appear at about six months of age, although some babies are born with one or more visible teeth, known as neonatal teeth. Normal tooth eruption at about six months can be painful. Kangaroos and manatees are unusual among mammals because they are polyphyodonts. In Aardvarks, teeth lack enamel and have many pulp tubules, hence the name of the order Tubulidentata. In dogs, the teeth are less than humans to form dental cavities because of the high pH of dog saliva, which prevents enamel from demineralizing. Sometimes called cuspids, these teeth are shaped like points and are used for tearing and grasping food Like human teeth, whale teeth have polyp-like protrusions located on the root surface of the tooth; these polyps are made of cementum in both species, but in human teeth, the protrusions are located on the outside of the root, while in whales the nodule is located on the inside of the pulp chamber.
While the roots of human teeth are made of cementum on the outer surface, whales have cementum on the entire surface of the tooth with a small layer of enamel at the tip. This small enamel layer is only seen in older whales where the cementum has been worn away to show the underlying enamel; the toothed whale is a suborder of the cetaceans characterized by having teeth. The teeth differ among the species, they may be numerous, with some dolphins bearing over 100 teeth in their jaws. On the other hand, the narwhals have a giant unicorn-like tusk, a tooth containing millions of sensory pathways and used for sensing during feeding and mating, it is the most neurologically complex tooth known. Beaked whales are toothless, with only bizarre teeth found in males; these teeth may be used for feeding but for demonstrating aggression and showmanship. In humans there are 20 primary teeth, 28 to 32 of what's known as permanent teeth, in addition to other four being third molars or wisdom teeth, each of which may or may not g
Permanent teeth or adult teeth are the second set of teeth formed in diphyodont mammals. In humans and old world simians, there are thirty-two permanent teeth, consisting of six maxillary and six mandibular molars, four maxillary and four mandibular premolars, two maxillary and two mandibular canines, four maxillary and four mandibular incisors; the first permanent tooth appears in the mouth at around six years of age, the mouth will be in a transition time with both primary teeth and permanent teeth during the mixed dentition period until the last primary tooth is lost or shed. The first of the permanent teeth to erupt are the permanent first molars, right behind the last'milk' molars of the primary dentition; these first permanent molars are important for the correct development of a permanent dentition. Up to the age of thirteen years, twenty-eight of the thirty-two permanent teeth will appear; the full permanent dentition is completed much during the permanent dentition period. The four last permanent teeth, the third molars appear between the ages of 17 and 38 years.
It is possible to have extra, or "supernumerary," teeth. This phenomenon is called hyperdontia and is erroneously referred to as "a third set of teeth." These teeth may remain impacted in the bone. Hyperdontia is associated with syndromes such as cleft lip and palate, trichorhinophalangeal syndrome, cleidocranial dysplasia, Gardner's syndrome. Deciduous dentition Tooth development Tooth eruption Ash, Major M. and Stanley J. Nelson, 2003. Wheeler’s Dental Anatomy and Occlusion. 8th edition. This article incorporates text in the public domain from the 20th edition of Gray's Anatomy Permanent dentition - Adult teeth chart Morphology permanent teeth
Cusp of Carabelli
The cusp of Carabelli, or Carabelli's tubercle, or tuberculum anomale of Georg Carabelli is a small additional cusp at the mesiopalatal line angle of maxillary first molars. This extra cusp is found on the first molar, becomes progressively less in the second, third molars; this cusp is absent in some individuals and present in others in a variety of forms. In some cases, the cusp of Carabelli may rival the main cusps in size. Other related forms pits, or furrows; this additional cusp was first described in 1842 by the Hungarian Georg Carabelli, the court dentist of the Austrian Emperor Franz. The cusp of Carabelli is an heritable feature. Kraus proposed that homozygosity of a gene is responsible for a pronounced tubercle, whereas the heterozygote shows slight grooves, tubercles or bulge. Studies showed that the development of this trait is affected by multiple genes; the cusp of Carabelli is most common among Europeans and rarest in Pacific Islands, although no study is referenced here to back up that claim.
Although it is sometimes referred to in textbooks as the Cusp of Carabelli, it is a tubercle, as it is may be made only of enamel, may not contain dentin but never has a root underneath. It is in unlikely, but possible for the cusp to have a pulp horn. If the cusp is large, approaching the size of a regular cusp it will be formed of dentin and enamel. Dens evaginatus Talon cusp Tubercle Simon Hillson. 1996. Dental Anthropology. Cambridge University Press
A mamelon is one of three rounded protuberances which are present on the cutting edge of an incisor tooth when it first erupts through the gum. Mamelons can't be removed except by removal of all teeth. Mamelons are present on permanent lateral incisors. Mamelons are easiest to observe on the maxillary central incisors, appear as three small prominences on the incisal edge of the tooth. Mamelons are ordinarily of no clinical importance, they are worn off early in the life of the tooth. However, when an anterior open bite is present — that is, the anterior teeth are not in contact when the bite is closed — mamelons may remain into adulthood
Marsupials are any members of the mammalian infraclass Marsupialia. All extant marsupials are endemic to Australasia and the Americas. A distinctive characteristic common to these species is that most of the young are carried in a pouch. Well-known marsupials include kangaroos, koalas, opossums and Tasmanian devils; some lesser-known marsupials are the dunnarts and cuscuses. Marsupials represent the clade originating from the last common ancestor of extant metatherians. Like other mammals in the Metatheria, they give birth to undeveloped young that reside in a pouch located on their mothers’ abdomen for a certain amount of time. Close to 70% of the 334 extant species occur on the Australian continent; the remaining 100 are found in the Americas — in South America, but thirteen in Central America, one in North America, north of Mexico. The word marsupial comes from the technical term for the abdominal pouch. It, in turn, is borrowed from Latin and from the ancient Greek μάρσιππος mársippos, meaning "pouch".
Marsupials are taxonomically identified as members of the mammalian infraclass Marsupialia, first described as a family under the order Pollicata by German zoologist Johann Karl Wilhelm Illiger in his 1811 work Prodromus Systematis Mammalium et Avium. However, James Rennie, author of The Natural History of Monkeys and Lemurs, pointed out that the placement of five different groups of mammals - monkeys, tarsiers, aye-ayes and marsupials - under a single order did not appear to have a strong justification. In 1816, French zoologist George Cuvier classified all marsupials under the order Marsupialia. In 1997, researcher J. A. W. Kirsch and others accorded infraclass rank to Marsupialia. There are two primary divisions: Australian marsupials. Marsupialia is further divided as follows:† - Extinct Superorder Ameridelphia Order Didelphimorphia Family Didelphidae: opossums Order Paucituberculata Family Caenolestidae: shrew opossums Superorder Australidelphia Order Microbiotheria Family Microbiotheriidae: monito del monte Order †Yalkaparidontia Order Dasyuromorphia Family †Thylacinidae: thylacine Family Dasyuridae: antechinuses, dunnarts, Tasmanian devil, relatives Family Myrmecobiidae: numbat Order Notoryctemorphia Family Notoryctidae: marsupial moles Order Peramelemorphia Family Thylacomyidae: bilbies Family †Chaeropodidae: pig-footed bandicoots Family Peramelidae: bandicoots and allies Order Diprotodontia Suborder Vombatiformes Family Vombatidae: wombats Family Phascolarctidae: koalas Family †Diprotodontidae: Giant wombats Family †Palorchestidae: Marsupial tapirs Family †Thylacoleonidae: marsupial lions Suborder Phalangeriformes Family Acrobatidae: feathertail glider and feather-tailed possum Family Burramyidae: pygmy possums Family †Ektopodontidae: sprite possums Family Petauridae: striped possum, Leadbeater's possum, yellow-bellied glider, sugar glider, mahogany glider, squirrel glider Family Phalangeridae: brushtail possums and cuscuses Family Pseudocheiridae: ringtailed possums and relatives Family Tarsipedidae: honey possum Suborder Macropodiformes Family Macropodidae: kangaroos and relatives Family Potoroidae: potoroos, rat kangaroos, bettongs Family Hypsiprymnodontidae: musky rat-kangaroo Comprising over 300 extant species, several attempts have been made to interpret the phylogenetic relationships among the different marsupial orders.
Studies differ on whether Didelphimorphia or Paucituberculata is the sister group to all other marsupials. Though the order Microbiotheria is found in South America, morphological similarities suggest it is related to Australian marsupials. Molecular analyses in 2010 and 2011 identified Microbiotheria as the sister group to all Australian marsupials. However, the relations among the four Australidelphid orders are not as well understood; the cladogram below, depicting the relationships among the various marsupial orders, is based on a 2015 phylogenetic study. DNA evidence supports a South American origin for marsupials, with Australian marsupials arising from a single Gondwanan migration of marsupials from South America to Australia. There are many small arboreal species in each group; the term "opossum" is used to refer to American species, while similar Australian species are properly called "possums". Marsupials have the typical characteristics of mammals—e.g. Mammary glands, three middle ear bones, true hair.
There are, striking differences as well as a number of anatomical features that separate them from Eutherians. In addition to the front pouch, which contains multiple nipples for the protection and sustenance of their young, marsupials have other common structural features. Ossified patellae are absent in most modern marsupials and epipubic bones are present. Marsupials lack a gross communication between the right and left brain hemispheres; the skull has peculiarities in comparison to placental mammals. In general, the skull is small and tight. Holes are located in the front of the orbit; the cheekbone extends further to the rear. The angular extension of the lower jaw is bent toward the center. Another feature is the hard palate which, in contrast to the placental mammals' foramina, always have more openings. The
In dentistry, crown refers to the anatomical area of teeth covered by enamel. The crown is visible in the mouth after developing below the gingiva and erupting into place. If part of the tooth gets chipped or broken, a dentist can apply an artificial crown. Crowns are used most to cover a damaged tooth or cover an implant. Bridges are used to cover a space if one or more teeth is missing, they are cemented to natural implants surrounding the space where the tooth once stood. There are various materials; the cement crowns look like regular teeth while the stainless steel crowns are gold. Ash, Major M. and Stanley J. Nelson. Wheeler’s Dental Anatomy and Occlusion. 8th edition. 2003. ISBN 0-7216-9382-2
The Hominidae, whose members are known as great apes or hominids, are a taxonomic family of primates that includes eight extant species in four genera: Pongo, the Bornean and Tapanuli orangutan. Several revisions in classifying the great apes have caused the use of the term "hominid" to vary over time, its original meaning referred only to their closest extinct relatives. That restrictive meaning has now been assumed by the term "hominin", which comprises all members of the human clade after the split from the chimpanzees; the current, 21st-century meaning of "hominid" includes all the great apes including humans. Usage still varies and some scientists and laypersons still use "hominid" in the original restrictive sense. Within the taxon Hominidae, a number of extant and known extinct, that is, genera are grouped with the humans and gorillas in the subfamily Homininae; the most recent common ancestor of all Hominidae lived 14 million years ago, when the ancestors of the orangutans speciated from the ancestral line of the other three genera.
Those ancestors of the family Hominidae had speciated from the family Hylobatidae 15 million to 20 million years ago. In the early Miocene, about 22 million years ago, there were many species of arboreally adapted primitive catarrhines from East Africa. Fossils at 20 million years ago include fragments attributed to Victoriapithecus, the earliest Old World monkey. Among the genera thought to be in the ape lineage leading up to 13 million years ago are Proconsul, Dendropithecus, Nacholapithecus, Nyanzapithecus, Afropithecus and Kenyapithecus, all from East Africa. At sites far distant from East Africa, the presence of other generalized non-cercopithecids, that is, non-monkey primates, of middle Miocene age—Otavipithecus from cave deposits in Namibia, Pierolapithecus and Dryopithecus from France and Austria—is further evidence of a wide diversity of ancestral ape forms across Africa and the Mediterranean basin during the warm and equable climatic regimes of the early and middle Miocene; the most recent of these far-flung Miocene apes is Oreopithecus, from the fossil-rich coal beds in northern Italy and dated to 9 million years ago.
Molecular evidence indicates that the lineage of gibbons, the lesser apes, diverged from that of the great apes some 18–12 million years ago, that of orangutans diverged from the other great apes at about 12 million years. There are no fossils that document the ancestry of gibbons, which may have originated in a still-unknown South East Asian hominoid population. Species close to the last common ancestor of gorillas and humans may be represented by Nakalipithecus fossils found in Kenya and Ouranopithecus found in Greece. Molecular evidence suggests that between 8 and 4 million years ago, first the gorillas, the chimpanzees split off from the line leading to the humans. Human DNA is 98.4% identical to that of chimpanzees when comparing single nucleotide polymorphisms. The fossil record, however, of gorillas and chimpanzees is limited. Other hominins adapted to the drier environments outside the African equatorial belt; the wet equatorial belt contracted after about 8 million years ago, there is little fossil evidence for the divergence of the hominin lineage from that of gorillas and chimpanzees—which split was thought to have occurred around that time.
The earliest fossils argued by some to belong to the human lineage are Sahelanthropus tchadensis and Orrorin tugenensis, followed by Ardipithecus, with species Ar. kadabba and Ar. ramidus. The classification of the great apes has been revised several times in the last few decades; the original meaning of the term referred to only humans and their closest relatives—what is now the modern meaning of the term "hominin". The meaning of the taxon Hominidae changed leading to a different usage of "hominid" that today includes all the great apes including humans; the term hominid is confused with a number of similar words: A hominoid called an ape, is a member of the superfamily Hominoidea: extant members are the gibbons and the hominids. A hominid is a member of the family Hominidae, the great apes: orangutans, gorillas and humans. A hominine is a member of the subfamily Homininae: gorillas and humans. A hominin is a member of the tribe Hominini: humans. A homininan, following a suggestion by Wood and Richmond, would be a member of the subtribe Homin