The Carboniferous is a geologic period and system that spans 60 million years from the end of the Devonian Period 358.9 million years ago, to the beginning of the Permian Period, 298.9 Mya. The name Carboniferous means "coal-bearing" and derives from the Latin words carbō and ferō, was coined by geologists William Conybeare and William Phillips in 1822. Based on a study of the British rock succession, it was the first of the modern'system' names to be employed, reflects the fact that many coal beds were formed globally during that time; the Carboniferous is treated in North America as two geological periods, the earlier Mississippian and the Pennsylvanian. Terrestrial animal life was well established by the Carboniferous period. Amphibians were the dominant land vertebrates, of which one branch would evolve into amniotes, the first terrestrial vertebrates. Arthropods were very common, many were much larger than those of today. Vast swaths of forest covered the land, which would be laid down and become the coal beds characteristic of the Carboniferous stratigraphy evident today.
The atmospheric content of oxygen reached its highest levels in geological history during the period, 35% compared with 21% today, allowing terrestrial invertebrates to evolve to great size. The half of the period experienced glaciations, low sea level, mountain building as the continents collided to form Pangaea. A minor marine and terrestrial extinction event, the Carboniferous rainforest collapse, occurred at the end of the period, caused by climate change. In the United States the Carboniferous is broken into Mississippian and Pennsylvanian subperiods; the Mississippian is about twice as long as the Pennsylvanian, but due to the large thickness of coal-bearing deposits with Pennsylvanian ages in Europe and North America, the two subperiods were long thought to have been more or less equal in duration. In Europe the Lower Carboniferous sub-system is known as the Dinantian, comprising the Tournaisian and Visean Series, dated at 362.5-332.9 Ma, the Upper Carboniferous sub-system is known as the Silesian, comprising the Namurian and Stephanian Series, dated at 332.9-298.9 Ma.
The Silesian is contemporaneous with the late Mississippian Serpukhovian plus the Pennsylvanian. In Britain the Dinantian is traditionally known as the Carboniferous Limestone, the Namurian as the Millstone Grit, the Westphalian as the Coal Measures and Pennant Sandstone; the International Commission on Stratigraphy faunal stages from youngest to oldest, together with some of their regional subdivisions, are: A global drop in sea level at the end of the Devonian reversed early in the Carboniferous. There was a drop in south polar temperatures; these conditions had little effect in the deep tropics, where lush swamps to become coal, flourished to within 30 degrees of the northernmost glaciers. Mid-Carboniferous, a drop in sea level precipitated a major marine extinction, one that hit crinoids and ammonites hard; this sea level drop and the associated unconformity in North America separate the Mississippian subperiod from the Pennsylvanian subperiod. This happened about 323 million years ago, at the onset of the Permo-Carboniferous Glaciation.
The Carboniferous was a time of active mountain-building as the supercontinent Pangaea came together. The southern continents remained tied together in the supercontinent Gondwana, which collided with North America–Europe along the present line of eastern North America; this continental collision resulted in the Hercynian orogeny in Europe, the Alleghenian orogeny in North America. In the same time frame, much of present eastern Eurasian plate welded itself to Europe along the line of the Ural Mountains. Most of the Mesozoic supercontinent of Pangea was now assembled, although North China, South China continents were still separated from Laurasia; the Late Carboniferous Pangaea was shaped like an "O." There were two major oceans in the Carboniferous—Panthalassa and Paleo-Tethys, inside the "O" in the Carboniferous Pangaea. Other minor oceans were shrinking and closed - Rheic Ocean, the small, shallow Ural Ocean and Proto-Tethys Ocean. Average global temperatures in the Early Carboniferous Period were high: 20 °C.
However, cooling during the Middle Carboniferous reduced average global temperatures to about 12 °C. Lack of growth rings of fossilized trees suggest a lack of seasons of a tropical climate. Glaciations in Gondwana, triggered by Gondwana's southward movement, continued into the Permian and because of the lack of clear markers and breaks, the deposits of this glacial period are referred to as Permo-Carboniferous in age; the cooling and drying of the climate led to the Carboniferous Rainforest Collapse during the late Carboniferous. Tropical rainforests fragmented and were devastated by climate change. Carboniferous rocks in Europe and eastern North America consist of a repeated sequence of limestone, sandstone and coal beds. In North America, the early Carboniferous is marine
In biology, a species is the basic unit of classification and a taxonomic rank of an organism, as well as a unit of biodiversity. A species is defined as the largest group of organisms in which any two individuals of the appropriate sexes or mating types can produce fertile offspring by sexual reproduction. Other ways of defining species include their karyotype, DNA sequence, behaviour or ecological niche. In addition, paleontologists use the concept of the chronospecies since fossil reproduction cannot be examined. While these definitions may seem adequate, when looked at more they represent problematic species concepts. For example, the boundaries between related species become unclear with hybridisation, in a species complex of hundreds of similar microspecies, in a ring species. Among organisms that reproduce only asexually, the concept of a reproductive species breaks down, each clone is a microspecies. All species are given a two-part name, a "binomial"; the first part of a binomial is the genus.
The second part is called the specific epithet. For example, Boa constrictor is one of four species of the genus Boa. None of these is satisfactory definitions, but scientists and conservationists need a species definition which allows them to work, regardless of the theoretical difficulties. If species were fixed and distinct from one another, there would be no problem, but evolutionary processes cause species to change continually, to grade into one another. Species were seen from the time of Aristotle until the 18th century as fixed kinds that could be arranged in a hierarchy, the great chain of being. In the 19th century, biologists grasped. Charles Darwin's 1859 book The Origin of Species explained how species could arise by natural selection; that understanding was extended in the 20th century through genetics and population ecology. Genetic variability arises from mutations and recombination, while organisms themselves are mobile, leading to geographical isolation and genetic drift with varying selection pressures.
Genes can sometimes be exchanged between species by horizontal gene transfer. Viruses are a special case, driven by a balance of mutation and selection, can be treated as quasispecies. Biologists and taxonomists have made many attempts to define species, beginning from morphology and moving towards genetics. Early taxonomists such as Linnaeus had no option but to describe what they saw: this was formalised as the typological or morphological species concept. Ernst Mayr emphasised reproductive isolation, but this, like other species concepts, is hard or impossible to test. Biologists have tried to refine Mayr's definition with the recognition and cohesion concepts, among others. Many of the concepts are quite similar or overlap, so they are not easy to count: the biologist R. L. Mayden recorded about 24 concepts, the philosopher of science John Wilkins counted 26. Wilkins further grouped the species concepts into seven basic kinds of concepts: agamospecies for asexual organisms biospecies for reproductively isolated sexual organisms ecospecies based on ecological niches evolutionary species based on lineage genetic species based on gene pool morphospecies based on form or phenotype and taxonomic species, a species as determined by a taxonomist.
A typological species is a group of organisms in which individuals conform to certain fixed properties, so that pre-literate people recognise the same taxon as do modern taxonomists. The clusters of variations or phenotypes within specimens would differentiate the species; this method was used as a "classical" method of determining species, such as with Linnaeus early in evolutionary theory. However, different phenotypes are not different species. Species named in this manner are called morphospecies. In the 1970s, Robert R. Sokal, Theodore J. Crovello and Peter Sneath proposed a variation on this, a phenetic species, defined as a set of organisms with a similar phenotype to each other, but a different phenotype from other sets of organisms, it differs from the morphological species concept in including a numerical measure of distance or similarity to cluster entities based on multivariate comparisons of a reasonably large number of phenotypic traits. A mate-recognition species is a group of sexually reproducing organisms that recognize one another as potential mates.
Expanding on this to allow for post-mating isolation, a cohesion species is the most inclusive population of individuals having the potential for phenotypic cohesion through intrinsic cohesion mechanisms. A further development of the recognition concept is provided by the biosemiotic concept of species. In microbiology, genes can move even between distantly related bacteria extending to the whole bacterial domain; as a rule of thumb, microbiologists have assumed that kinds of Bacteria or Archaea with 16S ribosomal RNA gene sequences more similar than 97% to each other need to be checked by DNA-DNA hybridisation to decide if they belong to the same species or not. This concept was narrowed in 2006 to a similarity of 98.7%. DNA-DNA hybri
Animals are multicellular eukaryotic organisms that form the biological kingdom Animalia. With few exceptions, animals consume organic material, breathe oxygen, are able to move, can reproduce sexually, grow from a hollow sphere of cells, the blastula, during embryonic development. Over 1.5 million living animal species have been described—of which around 1 million are insects—but it has been estimated there are over 7 million animal species in total. Animals range in length from 8.5 millionths of a metre to 33.6 metres and have complex interactions with each other and their environments, forming intricate food webs. The category includes humans, but in colloquial use the term animal refers only to non-human animals; the study of non-human animals is known as zoology. Most living animal species are in the Bilateria, a clade whose members have a bilaterally symmetric body plan; the Bilateria include the protostomes—in which many groups of invertebrates are found, such as nematodes and molluscs—and the deuterostomes, containing the echinoderms and chordates.
Life forms interpreted. Many modern animal phyla became established in the fossil record as marine species during the Cambrian explosion which began around 542 million years ago. 6,331 groups of genes common to all living animals have been identified. Aristotle divided animals into those with those without. Carl Linnaeus created the first hierarchical biological classification for animals in 1758 with his Systema Naturae, which Jean-Baptiste Lamarck expanded into 14 phyla by 1809. In 1874, Ernst Haeckel divided the animal kingdom into the multicellular Metazoa and the Protozoa, single-celled organisms no longer considered animals. In modern times, the biological classification of animals relies on advanced techniques, such as molecular phylogenetics, which are effective at demonstrating the evolutionary relationships between animal taxa. Humans make use of many other animal species for food, including meat and eggs. Dogs have been used in hunting, while many aquatic animals are hunted for sport.
Non-human animals have appeared in art from the earliest times and are featured in mythology and religion. The word "animal" comes from the Latin animalis, having soul or living being; the biological definition includes all members of the kingdom Animalia. In colloquial usage, as a consequence of anthropocentrism, the term animal is sometimes used nonscientifically to refer only to non-human animals. Animals have several characteristics. Animals are eukaryotic and multicellular, unlike bacteria, which are prokaryotic, unlike protists, which are eukaryotic but unicellular. Unlike plants and algae, which produce their own nutrients animals are heterotrophic, feeding on organic material and digesting it internally. With few exceptions, animals breathe oxygen and respire aerobically. All animals are motile during at least part of their life cycle, but some animals, such as sponges, corals and barnacles become sessile; the blastula is a stage in embryonic development, unique to most animals, allowing cells to be differentiated into specialised tissues and organs.
All animals are composed of cells, surrounded by a characteristic extracellular matrix composed of collagen and elastic glycoproteins. During development, the animal extracellular matrix forms a flexible framework upon which cells can move about and be reorganised, making the formation of complex structures possible; this may be calcified, forming structures such as shells and spicules. In contrast, the cells of other multicellular organisms are held in place by cell walls, so develop by progressive growth. Animal cells uniquely possess the cell junctions called tight junctions, gap junctions, desmosomes. With few exceptions—in particular, the sponges and placozoans—animal bodies are differentiated into tissues; these include muscles, which enable locomotion, nerve tissues, which transmit signals and coordinate the body. There is an internal digestive chamber with either one opening or two openings. Nearly all animals make use of some form of sexual reproduction, they produce haploid gametes by meiosis.
These fuse to form zygotes, which develop via mitosis into a hollow sphere, called a blastula. In sponges, blastula larvae swim to a new location, attach to the seabed, develop into a new sponge. In most other groups, the blastula undergoes more complicated rearrangement, it first invaginates to form a gastrula with a digestive chamber and two separate germ layers, an external ectoderm and an internal endoderm. In most cases, a third germ layer, the mesoderm develops between them; these germ layers differentiate to form tissues and organs. Repeated instances of mating with a close relative during sexual reproduction leads to inbreeding depression within a population due to the increased prevalence of harmful recessive traits. Animals have evolved numerous mechanisms for avoiding close inbreeding. In some species, such as the splendid fairywren, females benefit by mating with multiple males, thus producing more offspring of higher genetic quality; some animals are capable of asexual reproduction, which results
Productida is an extinct order of brachiopods in the extinct class Strophomenata. Following the Treatise Suborder Chonetidina Superfamily Chonetoidea Family Strophochonetidae Family Chonostrophiidae Family Anopliidae Family Eodevonariidae Family Chonetidae Family Rugosochonetidae Family Daviesiellidae Suborder Productidina Superfamily Productoidea Family Productellidae Family Productidae Superfamily Echinoconchoidea Family Echinoconchidae Family Sentosiidae Family Uncertain Superfamily Linoproductoidea Family Linoproductidae Family Monticuliferidae Superfamily Uncertain Suborder Strophalosiidina Superfamily Strophalosioidea Family Strophalosiidae Family Chonopectidae Family Araksalosiidae Superfamily Aulostegoidea Family Aulostegidae Family Cooperinidae Family Scacchinellidae Superfamily Richthofenioidea Family Richthofeniidae Family Hercosiidae Family Cyclacanthariidae Family Gemmellaroiidae Suborder Lyttoniidina Superfamily Lyttonioidea Family Lyttoniidae Family Rigbyellidae Superfamily Permianelloidea Family Permianellidae Data related to Productida at Wikispecies Media related to Productida at Wikimedia Commons Productida at fossilworks.org
Kelvingrove Art Gallery and Museum
The Kelvingrove Art Gallery and Museum is a museum and art gallery in Glasgow, Scotland. It reopened in 2006 after a three-year refurbishment and since has been one of Scotland's most popular visitor attractions; the gallery is located on Argyle Street, in the West End of the city, on the banks of the River Kelvin. It is adjacent to Kelvingrove Park and is situated near the main campus of the University of Glasgow on Gilmorehill; the construction of Kelvingrove was financed by the proceeds of the 1888 International Exhibition held in Kelvingrove Park. The gallery was designed by Sir John W. Simpson and E. J. Milner Allen and opened in 1901, as the Palace of Fine Arts for the Glasgow International Exhibition held in that year, it is built in a Spanish Baroque style, follows the Glaswegian tradition of using Locharbriggs red sandstone, includes an entire program of architectural sculpture by George Frampton, William Shirreffs, Francis Derwent Wood and other sculptors. The centrepiece of the Centre Hall is a concert pipe organ installed by Lewis & Co..
The organ was commissioned as part of the Glasgow International Exhibition, held in Kelvingrove Park in 1901. The organ was installed in the concert hall of the exhibition, capable of seating 3,000 people; the Centre Hall of the newly completed Art Gallery and Museum was intended from the beginning to be a space in which to hold concerts. When the 1901 exhibition ended, a Councillor urged the Glasgow Corporation to purchase the organ, stating that without it, "the art gallery would be a body without a soul". Purchase price and installation costs were met from the surplus exhibition proceeds, the organ was installed in the Centre Hall by Lewis and Co; the present case front in walnut with non-functional display pipes was commissioned at this time from John W. Simpson. Simpson was the senior partner of architects of the gallery building. There is an urban myth in Glasgow that the building was accidentally built back-to-front, the architect jumped from one of the towers in despair upon realising his mistake.
In reality, the grand entrance was always intended to face into Kelvingrove Park. The museum's collections came from the McLellan Galleries and from the old Kelvingrove House Museum in Kelvingrove Park, it has one of the finest collections of arms and armour in the world and a vast natural history collection. The art collection includes many outstanding European artworks, including works by the Old Masters, French Impressionists, Dutch Renaissance, Scottish Colourists and exponents of the Glasgow School; the museum houses Christ of Saint John of the Cross by Salvador Dalí. The copyright of this painting was bought by the curator at the time after a meeting with Dalí himself. For a period between 1993 and 2006, the painting was moved to the St Mungo Museum of Religious Life and Art; the museum contains a large gift of the decorative arts from Anne Hull Grundy, an art collector and philanthropist, covering the history of European jewellery in the 18th and 19th centuries. Kelvingrove was reopened by Her Majesty Queen Elizabeth II on 11 July 2006 after a three-year closure for major refurbishment and restoration.
The work cost around £28 million and includes a new restaurant and a large basement extension to its display space to accommodate the 8,000 exhibits now on display. A new display layout and wayfinding scheme was introduced to make the building more visitor-friendly. After its 2003–06 refurbishment, the museum was the most popular free-to-enter visitor attraction in Scotland, recording 2.23 million visitors in 2007. These numbers made it the most visited museum in the United Kingdom outside London. In 2015 there were 1,261,552 visitors; the Kelvingrove Museum is mentioned in the lyrics of the Irish ballad "Hot Asphalt", a song about Irish navvies laying asphalt in Britain. During the course of the song, the singer'catches his death of cold', is taxidermied and displayed in the museum'as a monument to the Irish making hot asphalt'. Museum website
The United Kingdom the United Kingdom of Great Britain and Northern Ireland, sometimes referred to as Britain, is a sovereign country located off the north-western coast of the European mainland. The United Kingdom includes the island of Great Britain, the north-eastern part of the island of Ireland, many smaller islands. Northern Ireland is the only part of the United Kingdom that shares a land border with another sovereign state, the Republic of Ireland. Apart from this land border, the United Kingdom is surrounded by the Atlantic Ocean, with the North Sea to the east, the English Channel to the south and the Celtic Sea to the south-west, giving it the 12th-longest coastline in the world; the Irish Sea lies between Great Ireland. With an area of 242,500 square kilometres, the United Kingdom is the 78th-largest sovereign state in the world, it is the 22nd-most populous country, with an estimated 66.0 million inhabitants in 2017. The UK is constitutional monarchy; the current monarch is Queen Elizabeth II, who has reigned since 1952, making her the longest-serving current head of state.
The United Kingdom's capital and largest city is London, a global city and financial centre with an urban area population of 10.3 million. Other major urban areas in the UK include Greater Manchester, the West Midlands and West Yorkshire conurbations, Greater Glasgow and the Liverpool Built-up Area; the United Kingdom consists of four constituent countries: England, Scotland and Northern Ireland. Their capitals are London, Edinburgh and Belfast, respectively. Apart from England, the countries have their own devolved governments, each with varying powers, but such power is delegated by the Parliament of the United Kingdom, which may enact laws unilaterally altering or abolishing devolution; the nearby Isle of Man, Bailiwick of Guernsey and Bailiwick of Jersey are not part of the UK, being Crown dependencies with the British Government responsible for defence and international representation. The medieval conquest and subsequent annexation of Wales by the Kingdom of England, followed by the union between England and Scotland in 1707 to form the Kingdom of Great Britain, the union in 1801 of Great Britain with the Kingdom of Ireland created the United Kingdom of Great Britain and Ireland.
Five-sixths of Ireland seceded from the UK in 1922, leaving the present formulation of the United Kingdom of Great Britain and Northern Ireland. There are fourteen British Overseas Territories, the remnants of the British Empire which, at its height in the 1920s, encompassed a quarter of the world's land mass and was the largest empire in history. British influence can be observed in the language and political systems of many of its former colonies; the United Kingdom is a developed country and has the world's fifth-largest economy by nominal GDP and ninth-largest economy by purchasing power parity. It has a high-income economy and has a high Human Development Index rating, ranking 14th in the world, it was the world's first industrialised country and the world's foremost power during the 19th and early 20th centuries. The UK remains a great power, with considerable economic, military and political influence internationally, it is sixth in military expenditure in the world. It has been a permanent member of the United Nations Security Council since its first session in 1946.
It has been a leading member state of the European Union and its predecessor, the European Economic Community, since 1973. The United Kingdom is a member of the Commonwealth of Nations, the Council of Europe, the G7, the G20, NATO, the Organisation for Economic Co-operation and Development and the World Trade Organization; the 1707 Acts of Union declared that the kingdoms of England and Scotland were "United into One Kingdom by the Name of Great Britain". The term "United Kingdom" has been used as a description for the former kingdom of Great Britain, although its official name from 1707 to 1800 was "Great Britain"; the Acts of Union 1800 united the kingdom of Great Britain and the kingdom of Ireland in 1801, forming the United Kingdom of Great Britain and Ireland. Following the partition of Ireland and the independence of the Irish Free State in 1922, which left Northern Ireland as the only part of the island of Ireland within the United Kingdom, the name was changed to the "United Kingdom of Great Britain and Northern Ireland".
Although the United Kingdom is a sovereign country, Scotland and Northern Ireland are widely referred to as countries. The UK Prime Minister's website has used the phrase "countries within a country" to describe the United Kingdom; some statistical summaries, such as those for the twelve NUTS 1 regions of the United Kingdom refer to Scotland and Northern Ireland as "regions". Northern Ireland is referred to as a "province". With regard to Northern Ireland, the descriptive name used "can be controversial, with the choice revealing one's political preferences"; the term "Great Britain" conventionally refers to the island of Great Britain, or politically to England and Wales in combination. However, it is sometimes used as a loose synonym for the United Kingdom as a whole; the term "Britain" is used both as a synonym for Great Britain, as a synonym for the United Kingdom. Usage is mixed, with the BBC preferring to use Britain as shorthand only for Great Britain and the UK Government, while accepting that both terms refer to the United K