Japan is an island country in East Asia. Located in the Pacific Ocean, it lies off the eastern coast of the Asian continent and stretches from the Sea of Okhotsk in the north to the East China Sea and the Philippine Sea in the south; the kanji that make up Japan's name mean "sun origin", it is called the "Land of the Rising Sun". Japan is a stratovolcanic archipelago consisting of about 6,852 islands; the four largest are Honshu, Hokkaido and Shikoku, which make up about ninety-seven percent of Japan's land area and are referred to as home islands. The country is divided into 47 prefectures in eight regions, with Hokkaido being the northernmost prefecture and Okinawa being the southernmost one; the population of 127 million is the world's tenth largest. 90.7 % of people live in cities. About 13.8 million people live in the capital of Japan. The Greater Tokyo Area is the most populous metropolitan area in the world with over 38 million people. Archaeological research indicates; the first written mention of Japan is in Chinese history texts from the 1st century AD.
Influence from other regions China, followed by periods of isolation from Western Europe, has characterized Japan's history. From the 12th century until 1868, Japan was ruled by successive feudal military shōguns who ruled in the name of the Emperor. Japan entered into a long period of isolation in the early 17th century, ended in 1853 when a United States fleet pressured Japan to open to the West. After nearly two decades of internal conflict and insurrection, the Imperial Court regained its political power in 1868 through the help of several clans from Chōshū and Satsuma – and the Empire of Japan was established. In the late 19th and early 20th centuries, victories in the First Sino-Japanese War, the Russo-Japanese War and World War I allowed Japan to expand its empire during a period of increasing militarism; the Second Sino-Japanese War of 1937 expanded into part of World War II in 1941, which came to an end in 1945 following the Japanese surrender. Since adopting its revised constitution on May 3, 1947, during the occupation led by SCAP, the sovereign state of Japan has maintained a unitary parliamentary constitutional monarchy with an Emperor and an elected legislature called the National Diet.
Japan is a member of the ASEAN Plus mechanism, UN, the OECD, the G7, the G8, the G20, is considered a great power. Its economy is the world's third-largest by nominal GDP and the fourth-largest by purchasing power parity, it is the world's fourth-largest exporter and fourth-largest importer. Japan benefits from a skilled and educated workforce. Although it has renounced its right to declare war, Japan maintains a modern military with the world's eighth-largest military budget, used for self-defense and peacekeeping roles. Japan is a developed country with a high standard of living and Human Development Index, its population enjoys the highest life expectancy and third lowest infant mortality rate in the world, but is experiencing issues due to an aging population and low birthrate. Japan is renowned for its historical and extensive cinema, influential music industry, video gaming, rich cuisine and its major contributions to science and modern technology; the Japanese word for Japan is 日本, pronounced Nihon or Nippon and means "the origin of the sun".
The character nichi means "sun" or "day". The compound therefore means "origin of the sun" and is the source of the popular Western epithet "Land of the Rising Sun"; the earliest record of the name Nihon appears in the Chinese historical records of the Tang dynasty, the Old Book of Tang. At the end of the seventh century, a delegation from Japan requested that Nihon be used as the name of their country; this name may have its origin in a letter sent in 607 and recorded in the official history of the Sui dynasty. Prince Shōtoku, the Regent of Japan, sent a mission to China with a letter in which he called himself "the Emperor of the Land where the Sun rises"; the message said: "Here, I, the emperor of the country where the sun rises, send a letter to the emperor of the country where the sun sets. How are you". Prior to the adoption of Nihon, other terms such as Yamato and Wakoku were used; the term Wa is a homophone of Wo 倭, used by the Chinese as a designation for the Japanese as early as the third century Three Kingdoms period.
Another form of Wa, Wei in Chinese) was used for an early state in Japan called Nakoku during the Han dynasty. However, the Japanese disliked some connotation of Wa 倭, it was therefore replaced with the substitute character Wa, meaning "togetherness, harmony"; the English word Japan derives from the historical Chinese pronunciation of 日本. The Old Mandarin or early Wu Chinese pronunciation of Japan was recorded by Marco Polo as Cipangu. In modern Shanghainese, a Wu dialect, the pronunciation of characters 日本; the old Malay word for Japan, Japun or Japang, was borrowed from a southern coastal Chinese dialect Fukienese or Ningpo – and this Malay word was encountered by Portuguese traders in Southeast Asia in the 16th century. These Early Portuguese traders brought the word
A herbarium is a collection of preserved plant specimens and associated data used for scientific study.. The specimens may be whole plants or plant parts; the specimens in a herbarium are used as reference material in describing plant taxa. The same term is used in mycology to describe an equivalent collection of preserved fungi, otherwise known as a fungarium. A xylarium is a herbarium specialising in specimens of wood; the term hortorium has been applied to a herbarium specialising in preserving material of horticultural origin. The oldest traditions of making herbarium collection or Hortus sicci have been traced to Italy. Luca Ghini and his students created herbaria of which the oldest extant one is that of Gherardo Cibo from around 1532. While most of the early herbaria were prepared with sheets bound into books, Carolus Linnaeus came up with the idea of maintaining them on free sheets that allowed their easy re-ordering within cabinets. Commensurate with the need of wildlife conservation, it is desirable to include in a herbarium sheet as much of the plant as possible, or at least representative parts of them in the case of large specimens.
To preserve their form and colour, plants collected in the field are arranged and spread flat between thin sheets, known as'flimsies', dried in a plant press, between blotters or absorbent paper. During the drying process the specimens are retained within their flimsies at all times to minimise damage, only the thicker, absorbent drying sheets are replaced. For some plants it may prove helpful to allow the fresh specimen to wilt before being arranged for the press. An opportunity to check and further lay out the specimen to best reveal the required features of the plant occurs when the damp absorbent sheets are changed during the drying/pressing process; the specimens, which are mounted on sheets of stiff white paper, are labelled with all essential data, such as date and place found, description of the plant and special habitat conditions. The sheet is placed in a protective case; as a precaution against insect attack, the pressed plant is frozen or poisoned, the case disinfected. Certain groups of plants are soft, bulky, or otherwise not amenable to drying and mounting on sheets.
For these plants, other methods of preparation and storage may be used. For example, conifer cones and palm fronds may be stored in labelled boxes. Representative flowers or fruits may be pickled in formaldehyde to preserve their three-dimensional structure. Small specimens, such as mosses and lichens, are air-dried and packaged in small paper envelopes. No matter the method of preservation, detailed information on where and when the plant was collected, habitat and the name of the collector is included; the value of a herbarium is much enhanced by the possession of “types”, that is, the original specimens on which the study of a species was founded. Thus the herbarium at the British Museum, rich in the earlier collections made in the eighteenth and early nineteenth centuries, contains the types of many species founded by the earlier workers in botany, it is rich in types of Australian plants from the collections of Sir Joseph Banks and Robert Brown, contains in addition many valuable modern collections.
Most herbaria utilize a standard system of organizing their specimens into herbarium cases. Specimen sheets are stacked in groups by the species to which they belong and placed into a large lightweight folder, labelled on the bottom edge. Groups of species folders are placed together into larger, heavier folders by genus; the genus folders are sorted by taxonomic family according to the standard system selected for use by the herbarium and placed into pigeonholes in herbarium cabinets. Locating a specimen filed in the herbarium requires knowing the nomenclature and classification used by the herbarium, it requires familiarity with possible name changes that have occurred since the specimen was collected, since the specimen may be filed under an older name. Modern herbaria maintain electronic databases of their collections. Many herbaria have initiatives to digitize specimens to produce a virtual herbarium; these records and images are made publicly accessible via the Internet. Herbarium collections can have great significance and value to science, have a large number of uses.
Herbaria are essential for the study of plant taxonomy, the study of geographic distributions, the stabilizing of nomenclature. Linnaeus's herbarium now belongs to the Linnean Society in England. Specimens housed in herbaria may identify the flora of an area. A large collection from a single area is used in writing a field guide or manual to aid in the identification of plants that grow there. With more specimens available, the author of the guide will better understand the variability of form in the plants and the natural distribution over which the plants grow. Herbaria preserve a historical record of change in vegetation over time. In some cases, plants may become extinct altogether. In such cases, specimens preserved in a herbarium can represent the only record of the plant's original distribution. Environmental scientists make use of such data to track changes in human impact. Herbaria have proven useful as source
Aquitaine, archaic Guyenne/Guienne, is a historical region of France and a former administrative region of the country. Since 1 January 2016 it has been part of the region Nouvelle-Aquitaine, it is situated in the south-western part of Metropolitan France, along the Atlantic Ocean and the Pyrenees mountain range on the border with Spain. It is composed of the five departments of Dordogne, Lot-et-Garonne, Pyrénées-Atlantiques and Gironde. In the Middle Ages, Aquitaine was a duchy, whose boundaries fluctuated considerably. There are traces of human settlement by prehistoric peoples in the Périgord, but the earliest attested inhabitants in the south-west were the Aquitani, who were not proper Celtic people, but more akin to the Iberians. Although a number of different languages and dialects were in use in the area during ancient times, it is most that the prevailing language of Aquitaine during the late pre-historic to Roman period was an early form of the Basque language; this has been demonstrated by various Aquitanian names and words that were recorded by the Romans, which are easily readable as Basque.
Whether this Aquitanian language was a remnant of a Vasconic language group that once extended much farther, or it was limited to the Aquitaine/Basque region is not known. One reason the language of Aquitaine is important is because Basque is the last surviving non-Indo-European language in western Europe and it has had some effect on the languages around it, including Spanish and, to a lesser extent, French; the original Aquitania at the time of Caesar's conquest of Gaul included the area bounded by the Garonne River, the Pyrenees and the Atlantic Ocean. The name may stem from Latin'aqua', maybe derived from the town "Aquae Augustae", "Aquae Tarbellicae" or just "Aquis" or as a more general geographical feature. Under Augustus' Roman rule, since 27 BC the province of Aquitania was further stretched to the north to the River Loire, thus including proper Gaul tribes along with old Aquitani south of the Garonne within the same region. In 392, the Roman imperial provinces were restructured as Aquitania Prima, Aquitania Secunda and Aquitania Tertia, better known as Novempopulania in the south-west.
Accounts of Aquitania during the Early Middle Ages are a blur, lacking precision, but there was much unrest. The Visigoths were called into Gaul as foederati, they established themselves as the de facto rulers in south-west Gaul as central Roman rule collapsed. Visigoths established their capital in Toulouse. In 507, they were expelled south to Hispania after their defeat in the Battle of Vouillé by the Franks, who became the new rulers in the area to the south of the Loire; the Roman Aquitania Tertia remained in place as Novempopulania, where a duke was appointed to hold a grip over the Basques. These dukes were quite detached from central Frankish overlordship, sometimes governing as independent rulers with strong ties to their kinsmen south of the Pyrenees; as of 660, the foundations for an independent Aquitaine/Vasconia polity were established by the duke Felix of Aquitaine, a magnate from Toulouse of Gallo-Roman stock. Despite its nominal submission to the Merovingians, the ethnic make-up of new realm Aquitaine wasn't Frankish, but Gallo-Roman north of the Garonne and main towns and Basque south of the Garonne.
A united Basque-Aquitanian realm reached its heyday under Odo the Great's rule. In 721, the Aquitanian duke fended Umayyad troops off at Toulouse, but in 732, an Umayyad expedition commanded by Abdul Rahman Al Ghafiqi defeated Odo next to Bordeaux, went on to loot its way up to Poitiers. Odo was required to pledge allegiance to the Frankish Charles Martel in exchange for help against the advancing Arabic forces. Basque-Aquitanian self-rule temporarily came to a halt in 768 after the assassination of Waifer. In 781, Charlemagne decided to proclaim his son Louis King of Aquitaine within the Carolingian Empire, ruling over a realm comprising the Duchy of Aquitaine and the Duchy of Vasconia He suppressed various Basque uprisings venturing into the lands of Pamplona past the Pyrenees after ravaging Gascony, with a view to imposing his authority in the Vasconia to south of Pyrenees. According to his biography, he achieved everything he wanted and after staying overnight in Pamplona, on his way back his army was attacked in Roncevaux in 812, but narrowly escaped an engagement at the Pyrenean passes.
Seguin, count of Bordeaux and Duke of Vasconia, seemed to have attempted a detachment from the Frankish central authority on Charlemagne's death. The new emperor Louis the Pious reacted by removing him from his capacity, which stirred the Basques into rebellion; the king in turn sent his troops to the territory, obtaining their submission in two campaigns and killing the duke, while his family crossed the Pyrenees and continued to foment risings against Frankish power. In 824, the 2nd Battle of Roncevaux took place, in which counts Aeblus and Aznar, Frankish vassals from the Duchy of Vasconia sent by the new King of Aquitaine, were captured by the joint forces of Iñigo Arista and the Banu Qasi. Before Pepin's death, emperor Louis had appointed a new king in 832, his son Charles the Bald, while the Aquitanian lords elected Pepin II as king; this struggle for control of the kingdom led to
France the French Republic, is a country whose territory consists of metropolitan France in Western Europe and several overseas regions and territories. The metropolitan area of France extends from the Mediterranean Sea to the English Channel and the North Sea, from the Rhine to the Atlantic Ocean, it is bordered by Belgium and Germany to the northeast and Italy to the east, Andorra and Spain to the south. The overseas territories include French Guiana in South America and several islands in the Atlantic and Indian oceans; the country's 18 integral regions span a combined area of 643,801 square kilometres and a total population of 67.3 million. France, a sovereign state, is a unitary semi-presidential republic with its capital in Paris, the country's largest city and main cultural and commercial centre. Other major urban areas include Lyon, Toulouse, Bordeaux and Nice. During the Iron Age, what is now metropolitan France was inhabited by a Celtic people. Rome annexed the area in 51 BC, holding it until the arrival of Germanic Franks in 476, who formed the Kingdom of Francia.
The Treaty of Verdun of 843 partitioned Francia into Middle Francia and West Francia. West Francia which became the Kingdom of France in 987 emerged as a major European power in the Late Middle Ages following its victory in the Hundred Years' War. During the Renaissance, French culture flourished and a global colonial empire was established, which by the 20th century would become the second largest in the world; the 16th century was dominated by religious civil wars between Protestants. France became Europe's dominant cultural and military power in the 17th century under Louis XIV. In the late 18th century, the French Revolution overthrew the absolute monarchy, established one of modern history's earliest republics, saw the drafting of the Declaration of the Rights of Man and of the Citizen, which expresses the nation's ideals to this day. In the 19th century, Napoleon established the First French Empire, his subsequent Napoleonic Wars shaped the course of continental Europe. Following the collapse of the Empire, France endured a tumultuous succession of governments culminating with the establishment of the French Third Republic in 1870.
France was a major participant in World War I, from which it emerged victorious, was one of the Allies in World War II, but came under occupation by the Axis powers in 1940. Following liberation in 1944, a Fourth Republic was established and dissolved in the course of the Algerian War; the Fifth Republic, led by Charles de Gaulle, remains today. Algeria and nearly all the other colonies became independent in the 1960s and retained close economic and military connections with France. France has long been a global centre of art and philosophy, it hosts the world's fourth-largest number of UNESCO World Heritage Sites and is the leading tourist destination, receiving around 83 million foreign visitors annually. France is a developed country with the world's sixth-largest economy by nominal GDP, tenth-largest by purchasing power parity. In terms of aggregate household wealth, it ranks fourth in the world. France performs well in international rankings of education, health care, life expectancy, human development.
France is considered a great power in global affairs, being one of the five permanent members of the United Nations Security Council with the power to veto and an official nuclear-weapon state. It is a leading member state of the European Union and the Eurozone, a member of the Group of 7, North Atlantic Treaty Organization, Organisation for Economic Co-operation and Development, the World Trade Organization, La Francophonie. Applied to the whole Frankish Empire, the name "France" comes from the Latin "Francia", or "country of the Franks". Modern France is still named today "Francia" in Italian and Spanish, "Frankreich" in German and "Frankrijk" in Dutch, all of which have more or less the same historical meaning. There are various theories as to the origin of the name Frank. Following the precedents of Edward Gibbon and Jacob Grimm, the name of the Franks has been linked with the word frank in English, it has been suggested that the meaning of "free" was adopted because, after the conquest of Gaul, only Franks were free of taxation.
Another theory is that it is derived from the Proto-Germanic word frankon, which translates as javelin or lance as the throwing axe of the Franks was known as a francisca. However, it has been determined that these weapons were named because of their use by the Franks, not the other way around; the oldest traces of human life in what is now France date from 1.8 million years ago. Over the ensuing millennia, Humans were confronted by a harsh and variable climate, marked by several glacial eras. Early hominids led a nomadic hunter-gatherer life. France has a large number of decorated caves from the upper Palaeolithic era, including one of the most famous and best preserved, Lascaux. At the end of the last glacial period, the climate became milder. After strong demographic and agricultural development between the 4th and 3rd millennia, metallurgy appeared at the end of the 3rd millennium working gold and bronze, iron. France has numerous megalithic sites from the Neolithic period, including the exceptiona
Bordeaux is a port city on the Garonne in the Gironde department in Southwestern France. The municipality of Bordeaux proper has a population of 252,040. Together with its suburbs and satellite towns, Bordeaux is the centre of the Bordeaux Métropole. With 1,195,335 in the metropolitan area, it is the sixth-largest in France, after Paris, Lyon and Lille, it is the capital of the Nouvelle-Aquitaine region, as well as the prefecture of the Gironde department. Its inhabitants are called "Bordelais" or "Bordelaises"; the term "Bordelais" may refer to the city and its surrounding region. Being at the center of a major wine-growing and wine-producing region, Bordeaux remains a prominent powerhouse and exercises significant influence on the world wine industry although no wine production is conducted within the city limits, it is home to the world's main wine fair and the wine economy in the metro area takes in 14.5 billion euros each year. Bordeaux wine has been produced in the region since the 8th century.
The historic part of the city is on the UNESCO World Heritage List as "an outstanding urban and architectural ensemble" of the 18th century. After Paris, Bordeaux has the highest number of preserved historical buildings of any city in France. In historical times, around 567 BC it was the settlement of a Celtic tribe, the Bituriges Vivisci, who named the town Burdigala of Aquitanian origin; the name Bourde is still the name of a river south of the city. In 107 BC, the Battle of Burdigala was fought by the Romans who were defending the Allobroges, a Gallic tribe allied to Rome, the Tigurini led by Divico; the Romans were defeated and their commander, the consul Lucius Cassius Longinus, was killed in the action. The city fell under Roman rule around its importance lying in the commerce of tin and lead, it became capital of Roman Aquitaine, flourishing during the Severan dynasty. In 276 it was sacked by the Vandals. Further ravage was brought by the same Vandals in 409, the Visigoths in 414, the Franks in 498, beginning a period of obscurity for the city.
In the late 6th century, the city re-emerged as the seat of a county and an archdiocese within the Merovingian kingdom of the Franks, but royal Frankish power was never strong. The city started to play a regional role as a major urban center on the fringes of the newly founded Frankish Duchy of Vasconia. Around 585, Gallactorius is fighting the Basque people; the city was plundered by the troops of Abd er Rahman in 732 after they stormed the fortified city and overwhelmed the Aquitanian garrison. Duke Eudes mustered a force ready to engage the Umayyads outside Bordeaux taking them on in the Battle of the River Garonne somewhere near the river Dordogne; the battle had a high death toll. Although Eudes was defeated here, he saved part of his troops and kept his grip on Aquitaine after the Battle of Poitiers. In 735, the Aquitanian duke Hunald led a rebellion after his father Eudes's death, at which Charles responded by sending an expedition that captured and plundered Bordeaux again, but did not retain it for long.
The following year, the Frankish commander descended again to Aquitaine, but clashed in battle with the Aquitanians and left to take on hostile Burgundian authorities and magnates. In 745, Aquitaine faced yet another expedition by Charles's sons Pepin and Carloman, against Hunald, the Aquitanian princeps strong in Bordeaux. Hunald was defeated, his son Waifer replaced him, confirmed Bordeaux as the capital city. During the last stage of the war against Aquitaine, it was one of Waifer's last important strongholds to fall to King Pepin the Short's troops. Next to Bordeaux, Charlemagne built the fortress of Fronsac on a hill across the border with the Basques, where Basque commanders came over to vow loyalty to him. In 778, Seguin was appointed count of Bordeaux undermining the power of the Duke Lupo, leading to the Battle of Roncevaux Pass that year. In 814, Seguin was made Duke of Vasconia, but he was deposed in 816 for failing to suppress or sympathise with a Basque rebellion. Under the Carolingians, sometimes the Counts of Bordeaux held the title concomitantly with that of Duke of Vasconia.
They were meant to keep the Basques in check and defend the mouth of the Garonne from the Vikings when the latter appeared c. 844 in the region of Bordeaux. In Autumn 845, count Seguin II marched on the Vikings, who were assaulting Bordeaux and Saintes, but he was captured and executed. No bishops were mentioned during part of the 9th in Bordeaux. From the 12th to the 15th century, Bordeaux regained importance following the marriage of Duchess Eléonore of Aquitaine with the French-speaking Count Henri Plantagenet, born in Le Mans, who became, within months of their wedding, King Henry II of England; the city flourished due to the wine trade, the cathedral of St. André was built, it was the capital of an independent state under Edward, the Black Prince, but in the end, after the Battle of Castillon, it was annexed by France which extended its territory. The Château Trompette and the Fort du Hâ, built by Charles VII of France, were the symbols of the new domination, which however deprived the city of its wealth by halting the wine commerce with England.
In 1462, Bordeaux obtained a parliament, but regained importance only in the 16th century when it became the centre of the distribution of sugar and slaves from the West Indies along with the traditional wine. Bordeaux adhered to the Fronde
Gironde is a department in the Nouvelle-Aquitaine region of southwest France. It is named after a major waterway; the Bordeaux wine region is in the Gironde. Gironde is one of the original 83 departments created during the French Revolution on 4 March 1790, it was created from parts of the former provinces of Gascony. From 1793 to 1795, the department's name was changed to Bec-d'Ambès to avoid the association with the revolutionary party, the Girondists. Gironde is part of the current region of Nouvelle-Aquitaine and is surrounded by the departments of Landes, Lot-et-Garonne and Charente-Maritime and the Atlantic Ocean on the west. With an area of 10,000 km², Gironde is the largest department in metropolitan France. If overseas departments are included, Gironde's land area is dwarfed by the 83,846 km² of French Guiana. Gironde is well known for the Côte d'Argent beach, Europe's longest, attracting many surfers to Lacanau each year, it is the birthplace of Jacques-Yves Cousteau who studied the sea and all forms of life in water.
The Great Dune of Pyla in Arcachon Bay near Bordeaux is the tallest sand dune in Europe. The President of the General Council is Jean-Luc Gleyze of the Socialist Party. Cantons of the Gironde department Communes of the Gironde department Arrondissements of the Gironde department Bordeaux wine regions General Council website Prefecture website Gironde at Curlie Tourism Office website
In biology, a species is the basic unit of classification and a taxonomic rank of an organism, as well as a unit of biodiversity. A species is defined as the largest group of organisms in which any two individuals of the appropriate sexes or mating types can produce fertile offspring by sexual reproduction. Other ways of defining species include their karyotype, DNA sequence, behaviour or ecological niche. In addition, paleontologists use the concept of the chronospecies since fossil reproduction cannot be examined. While these definitions may seem adequate, when looked at more they represent problematic species concepts. For example, the boundaries between related species become unclear with hybridisation, in a species complex of hundreds of similar microspecies, in a ring species. Among organisms that reproduce only asexually, the concept of a reproductive species breaks down, each clone is a microspecies. All species are given a two-part name, a "binomial"; the first part of a binomial is the genus.
The second part is called the specific epithet. For example, Boa constrictor is one of four species of the genus Boa. None of these is satisfactory definitions, but scientists and conservationists need a species definition which allows them to work, regardless of the theoretical difficulties. If species were fixed and distinct from one another, there would be no problem, but evolutionary processes cause species to change continually, to grade into one another. Species were seen from the time of Aristotle until the 18th century as fixed kinds that could be arranged in a hierarchy, the great chain of being. In the 19th century, biologists grasped. Charles Darwin's 1859 book The Origin of Species explained how species could arise by natural selection; that understanding was extended in the 20th century through genetics and population ecology. Genetic variability arises from mutations and recombination, while organisms themselves are mobile, leading to geographical isolation and genetic drift with varying selection pressures.
Genes can sometimes be exchanged between species by horizontal gene transfer. Viruses are a special case, driven by a balance of mutation and selection, can be treated as quasispecies. Biologists and taxonomists have made many attempts to define species, beginning from morphology and moving towards genetics. Early taxonomists such as Linnaeus had no option but to describe what they saw: this was formalised as the typological or morphological species concept. Ernst Mayr emphasised reproductive isolation, but this, like other species concepts, is hard or impossible to test. Biologists have tried to refine Mayr's definition with the recognition and cohesion concepts, among others. Many of the concepts are quite similar or overlap, so they are not easy to count: the biologist R. L. Mayden recorded about 24 concepts, the philosopher of science John Wilkins counted 26. Wilkins further grouped the species concepts into seven basic kinds of concepts: agamospecies for asexual organisms biospecies for reproductively isolated sexual organisms ecospecies based on ecological niches evolutionary species based on lineage genetic species based on gene pool morphospecies based on form or phenotype and taxonomic species, a species as determined by a taxonomist.
A typological species is a group of organisms in which individuals conform to certain fixed properties, so that pre-literate people recognise the same taxon as do modern taxonomists. The clusters of variations or phenotypes within specimens would differentiate the species; this method was used as a "classical" method of determining species, such as with Linnaeus early in evolutionary theory. However, different phenotypes are not different species. Species named in this manner are called morphospecies. In the 1970s, Robert R. Sokal, Theodore J. Crovello and Peter Sneath proposed a variation on this, a phenetic species, defined as a set of organisms with a similar phenotype to each other, but a different phenotype from other sets of organisms, it differs from the morphological species concept in including a numerical measure of distance or similarity to cluster entities based on multivariate comparisons of a reasonably large number of phenotypic traits. A mate-recognition species is a group of sexually reproducing organisms that recognize one another as potential mates.
Expanding on this to allow for post-mating isolation, a cohesion species is the most inclusive population of individuals having the potential for phenotypic cohesion through intrinsic cohesion mechanisms. A further development of the recognition concept is provided by the biosemiotic concept of species. In microbiology, genes can move even between distantly related bacteria extending to the whole bacterial domain; as a rule of thumb, microbiologists have assumed that kinds of Bacteria or Archaea with 16S ribosomal RNA gene sequences more similar than 97% to each other need to be checked by DNA-DNA hybridisation to decide if they belong to the same species or not. This concept was narrowed in 2006 to a similarity of 98.7%. DNA-DNA hybri