The flowering plants known as angiosperms, Angiospermae or Magnoliophyta, are the most diverse group of land plants, with 64 orders, 416 families 13,164 known genera and c. 369,000 known species. Like gymnosperms, angiosperms are seed-producing plants. However, they are distinguished from gymnosperms by characteristics including flowers, endosperm within the seeds, the production of fruits that contain the seeds. Etymologically, angiosperm means a plant; the term comes from the Greek words sperma. The ancestors of flowering plants diverged from gymnosperms in the Triassic Period, 245 to 202 million years ago, the first flowering plants are known from 160 mya, they diversified extensively during the Early Cretaceous, became widespread by 120 mya, replaced conifers as the dominant trees from 100 to 60 mya. Angiosperms differ from other seed plants in several ways, described in the table below; these distinguishing characteristics taken together have made the angiosperms the most diverse and numerous land plants and the most commercially important group to humans.
Angiosperm stems are made up of seven layers. The amount and complexity of tissue-formation in flowering plants exceeds that of gymnosperms; the vascular bundles of the stem are arranged such that the phloem form concentric rings. In the dicotyledons, the bundles in the young stem are arranged in an open ring, separating a central pith from an outer cortex. In each bundle, separating the xylem and phloem, is a layer of meristem or active formative tissue known as cambium. By the formation of a layer of cambium between the bundles, a complete ring is formed, a regular periodical increase in thickness results from the development of xylem on the inside and phloem on the outside; the soft phloem becomes crushed, but the hard wood persists and forms the bulk of the stem and branches of the woody perennial. Owing to differences in the character of the elements produced at the beginning and end of the season, the wood is marked out in transverse section into concentric rings, one for each season of growth, called annual rings.
Among the monocotyledons, the bundles are more numerous in the young stem and are scattered through the ground tissue. They once formed the stem increases in diameter only in exceptional cases; the characteristic feature of angiosperms is the flower. Flowers show remarkable variation in form and elaboration, provide the most trustworthy external characteristics for establishing relationships among angiosperm species; the function of the flower is to ensure fertilization of the ovule and development of fruit containing seeds. The floral apparatus may arise terminally from the axil of a leaf; as in violets, a flower arises singly in the axil of an ordinary foliage-leaf. More the flower-bearing portion of the plant is distinguished from the foliage-bearing or vegetative portion, forms a more or less elaborate branch-system called an inflorescence. There are two kinds of reproductive cells produced by flowers. Microspores, which will divide to become pollen grains, are the "male" cells and are borne in the stamens.
The "female" cells called megaspores, which will divide to become the egg cell, are contained in the ovule and enclosed in the carpel. The flower may consist only of these parts, as in willow, where each flower comprises only a few stamens or two carpels. Other structures are present and serve to protect the sporophylls and to form an envelope attractive to pollinators; the individual members of these surrounding structures are known as petals. The outer series is green and leaf-like, functions to protect the rest of the flower the bud; the inner series is, in general, white or brightly colored, is more delicate in structure. It functions to attract bird pollinators. Attraction is effected by color and nectar, which may be secreted in some part of the flower; the characteristics that attract pollinators account for the popularity of flowers and flowering plants among humans. While the majority of flowers are perfect or hermaphrodite, flowering plants have developed numerous morphological and physiological mechanisms to reduce or prevent self-fertilization.
Heteromorphic flowers have short carpels and long stamens, or vice versa, so animal pollinators cannot transfer pollen to the pistil. Homomorphic flowers may employ a biochemical mechanism called self-incompatibility to discriminate between self and non-self pollen grains. In other species, the male and female parts are morphologically separated, developing on different flowers; the botanical term "Angiosperm", from the Ancient Greek αγγείον, angeíon and σπέρμα, was coined in the form Angiospermae by Paul Hermann in 1690, as the name of one of his primary divisions of the plant kingdom. This included flowering plants possessing seeds enclosed in capsules, distinguished from his Gymnospermae, or flowering plants with achenial or schizo-carpic fruits, the whole fruit or each of its pieces being here regarded as a seed and naked; the term and its antonym were maintained by Carl Linnaeus with the same sense, but with restricted application, in the names of the orders of his class Didynamia. Its use with any
Krameria lappacea, the para rhatany and Peruvian rhatany, is a plant species in the genus Krameria, in Peru. "Krameria lappacea". Germplasm Resources Information Network. Agricultural Research Service, United States Department of Agriculture. Krameria lappacea on www.liberherbarum.com
The Zygophyllales are an order of dicotyledonous plants, comprising the following two families: Family Zygophyllaceae Family KrameriaceaeAccording to the Angiosperm Phylogeny Group both families are unplaced to order, but included in the Eurosids I. The APG III system of 2009, recognized this order. If the monogeneric family Krameriaceae shares little common traits with the family Zygophyllaceae, researchers see little advantage in keeping it as a separate family; the name Zygophyllales can be used. The order keeps unchanged in the APG IV system. Under the Cronquist system, the Zygophyllaceae were included within the Sapindales, the Krameriaceae within the Polygalales
Wine is an alcoholic drink made from fermented grapes. Yeast consumes the sugar in the grapes and converts it to ethanol, carbon dioxide, heat. Different varieties of grapes and strains of yeasts produce different styles of wine; these variations result from the complex interactions between the biochemical development of the grape, the reactions involved in fermentation, the terroir, the production process. Many countries enact legal appellations intended to define qualities of wine; these restrict the geographical origin and permitted varieties of grapes, as well as other aspects of wine production. Wines not made from grapes include rice wine and fruit wines such as plum, pomegranate and elderberry. Wine has been produced for thousands of years; the earliest known traces of wine are from Georgia and Sicily although there is evidence of a similar alcoholic drink being consumed earlier in China. The earliest known winery is the 6,100-year-old Areni-1 winery in Armenia. Wine reached the Balkans by 4500 BC and was consumed and celebrated in ancient Greece and Rome.
Throughout history, wine has been consumed for its intoxicating effects. Wine has long played an important role in religion. Red wine was associated with blood by the ancient Egyptians and was used by both the Greek cult of Dionysus and the Romans in their Bacchanalia; the earliest archaeological and archaeobotanical evidence for grape wine and viniculture, dating to 6000–5800 BC was found on the territory of modern Georgia. Both archaeological and genetic evidence suggest that the earliest production of wine elsewhere was later having taken place in the Southern Caucasus, or the West Asian region between Eastern Turkey, northern Iran; the earliest evidence of a grape-based fermented drink was found in China, Georgia from 6000 BC, Iran from 5000 BC, Sicily from 4000 BC. The earliest evidence of a wine production facility is the Areni-1 winery in Armenia and is at least 6100 years old. A 2003 report by archaeologists indicates a possibility that grapes were mixed with rice to produce mixed fermented drinks in China in the early years of the seventh millennium BC.
Pottery jars from the Neolithic site of Jiahu, contained traces of tartaric acid and other organic compounds found in wine. However, other fruits indigenous to the region, such as hawthorn, cannot be ruled out. If these drinks, which seem to be the precursors of rice wine, included grapes rather than other fruits, they would have been any of the several dozen indigenous wild species in China, rather than Vitis vinifera, introduced there 6000 years later; the spread of wine culture westwards was most due to the Phoenicians who spread outward from a base of city-states along the Mediterranean coast of what are today Syria, Lebanon and Palestine. The wines of Byblos were exported to Egypt during the Old Kingdom and throughout the Mediterranean. Evidence includes two Phoenician shipwrecks from 750 BC discovered by Robert Ballard, whose cargo of wine was still intact; as the first great traders in wine, the Phoenicians seem to have protected it from oxidation with a layer of olive oil, followed by a seal of pinewood and resin, similar to retsina.
Although the nuragic Sardinians consumed wine before the arrival of the Phoenicians The earliest remains of Apadana Palace in Persepolis dating back to 515 BC include carvings depicting soldiers from Achaemenid Empire subject nations bringing gifts to the Achaemenid king, among them Armenians bringing their famous wine. Literary references to wine are abundant in Homer and others. In ancient Egypt, six of 36 wine amphoras were found in the tomb of King Tutankhamun bearing the name "Kha'y", a royal chief vintner. Five of these amphoras were designated as originating from the king's personal estate, with the sixth from the estate of the royal house of Aten. Traces of wine have been found in central Asian Xinjiang in modern-day China, dating from the second and first millennia BC; the first known mention of grape-based wines in India is from the late 4th-century BC writings of Chanakya, the chief minister of Emperor Chandragupta Maurya. In his writings, Chanakya condemns the use of alcohol while chronicling the emperor and his court's frequent indulgence of a style of wine known as madhu.
The ancient Romans planted vineyards near garrison towns so wine could be produced locally rather than shipped over long distances. Some of these areas are now world-renowned for wine production; the Romans discovered that burning sulfur candles inside empty wine vessels kept them fresh and free from a vinegar smell. In medieval Europe, the Roman Catholic Church supported wine because the clergy required it for the Mass. Monks in France made wine for years. An old English recipe that survived in various forms until the 19th century calls for refining white wine from bastard—bad or tainted bastardo wine; the English word "wine" comes from the Proto-Germanic *winam, an early borrowing from the Latin vinum, "wine" or " vine", itself derived from the Proto-Indo-European stem *win-o-. The earliest attested terms referring to wine are the Mycenaean Greek me-tu-wo ne-wo, meaning "in" or " of the new wine", wo-no-wa-ti-si, meaning "wine garden", written in Linear B inscriptions. Linear B includes, inter alia, an ideogram for wine
Encyclopedia Americana is one of the largest general encyclopedias in the English language. Following the acquisition of Grolier in 2000, the encyclopedia has been produced by Scholastic; the encyclopedia has more than 45,000 articles, most of them more than 500 words and many running to considerable length. The work's coverage of American and Canadian geography and history has been a traditional strength. Written by 6,500 contributors, the Encyclopedia Americana includes over 9,000 bibliographies, 150,000 cross-references, 1,000+ tables, 1,200 maps, 4,500 black-and-white line art and color images, it has 680 factboxes. Most articles are signed by their contributors. Long available as a 30-volume print set, the Encyclopedia Americana is now marketed as an online encyclopedia requiring a subscription. In March 2008, Scholastic said that print sales remained good but that the company was still deciding on the future of the print edition; the company did not produce an edition in 2007, a change from its previous approach of releasing a revised print edition each year.
The most recent print edition of the Encyclopedia Americana was published in 2006. The online version of the Encyclopedia Americana, first introduced in 1997, continues to be updated and sold; this work, like the print set from which it is derived, is designed for high school and first-year college students along with public library users. It is available to libraries as one of the options in the Grolier Online reference service, which includes the Grolier Multimedia Encyclopedia, intended for middle and high school students, The New Book of Knowledge, an encyclopedia for elementary and middle school students. Grolier Online is not available to individual subscribers. There have been three separate works using the title Encyclopedia Americana; the first began publishing in the 1820s by the German exile Francis Lieber. The 13 volumes of the first edition were completed in 1833, other editions and printings followed in 1835, 1836, 1847-1848, 1849 and 1858. Lieber's work was based upon and was in no small part a translation of the 7th edition of the well established Konversations-Lexikon of Brockhaus.
Some material from this set was carried over into the modern version, as well as the Brockhaus short article method. A separate Encyclopedia Americana was published by J. M. Stoddart between 1883 and 1889, as a supplement to American reprintings of the 9th edition of the Encyclopædia Britannica, it was four quarto volumes meant to "extend and complete the articles in Britannica". Stoddart's work, however, is not connected to the earlier work by Lieber. In 1902 a new version in 16 volumes was published under the title Encyclopedia Americana, under the editorial supervision of Scientific American magazine; the magazine's editor, Frederick Converse Beach, was editor-in-chief, was said to be assisted by hundreds of eminent scholars and authorities who served as consulting editors or authors. The first publisher was R. S. Peale & Co; the relationship with Scientific American was terminated in 1911. From 1907 to 1912, the work was published as The Americana. A major new edition appeared with George Edwin Rines as editor-in-chief.
An Annual or Yearbook was published each year beginning in 1923 and continuing until 2000. The encyclopedia was purchased by Grolier in 1945. By the 1960s, sales of the Americana and its sister publications under Grolier—The Book of Knowledge, the Book of Popular Science, Lands and Peoples—were strong enough to support the company's occupancy of a large building in Midtown Manhattan, at 575 Lexington Avenue. Sales during this period were accomplished through mail-order and door-to-door operations. Telemarketing and third-party distribution through their Lexicon division added to sales volumes in the 1970s. By the late 1970s, Grolier had moved its operations to Connecticut. In 1988 Grolier was purchased by the French media company Hachette, which owned a well-known French-language encyclopedia, the Hachette Encyclopedia. Hachette was absorbed by the French conglomerate the Lagardère Group. A CD-ROM version of the encyclopedia was published in 1995. Although the text and images were stored on separate disks, it was in keeping with standards current at the time.
More the work had been digitized, allowing for release of an online version in 1997. Over the next few years the product was augmented with additional features, supplementary references, Internet links, current events journal. A redesigned interface and reengineered product, featuring enhanced search capabilities and a first-ever ADA-compliant, text-only version for users with disabilities, was presented in 2002; the acquisition of Grolier by Scholastic for US$400 million, took place in 2000. The new owners projected a 30% increase in operating income, although Grolier had experienced earnings of 7% to 8% on income. Staff reductions as a means of controlling costs followed soon thereafter while an effort was made to augment the sales force. Cuts occurred every year between 2000 and 2007, leaving a much-depleted work force to carry out the duties of maintaining a large encyclopedia database. Today, Encyclopedia Americana lives on as an integral database within the Grolier Online product. Frederick Converse Beach, 1902–1917.
Engineer and editor of Scientific American magazine. George Edwin Rines, 1917–1920. Author and editor. A. H. McDannald, 1920–1948. Reporter and author
The eudicots, Eudicotidae or eudicotyledons are a clade of flowering plants, called tricolpates or non-magnoliid dicots by previous authors. The botanical terms were introduced in 1991 by evolutionary botanist James A. Doyle and paleobotanist Carol L. Hotton to emphasize the evolutionary divergence of tricolpate dicots from earlier, less specialized, dicots; the close relationships among flowering plants with tricolpate pollen grains was seen in morphological studies of shared derived characters. These plants have a distinct trait in their pollen grains of exhibiting three colpi or grooves paralleling the polar axis. Molecular evidence confirmed the genetic basis for the evolutionary relationships among flowering plants with tricolpate pollen grains and dicotyledonous traits; the term means "true dicotyledons", as it contains the majority of plants that have been considered dicots and have characteristics of the dicots. The term "eudicots" has subsequently been adopted in botany to refer to one of the two largest clades of angiosperms, monocots being the other.
The remaining angiosperms include magnoliids and what are sometimes referred to as basal angiosperms or paleodicots, but these terms have not been or adopted, as they do not refer to a monophyletic group. The other name for the eudicots is tricolpates, a name which refers to the grooved structure of the pollen. Members of the group have tricolpate pollen; these pollens have three or more pores set in furrows called colpi. In contrast, most of the other seed plants produce monosulcate pollen, with a single pore set in a differently oriented groove called the sulcus; the name "tricolpates" is preferred by some botanists to avoid confusion with the dicots, a nonmonophyletic group. Numerous familiar plants are eudicots, including many common food plants and ornamentals; some common and familiar eudicots include members of the sunflower family such as the common dandelion, the forget-me-not and other members of its family, buttercup and macadamia. Most leafy trees of midlatitudes belong to eudicots, with notable exceptions being magnolias and tulip trees which belong to magnoliids, Ginkgo biloba, not an angiosperm.
The name "eudicots" is used in the APG system, of 1998, APG II system, of 2003, for classification of angiosperms. It is applied to a monophyletic group, which includes most of the dicots. "Tricolpate" is a synonym for the "Eudicot" monophyletic group, the "true dicotyledons". The number of pollen grain furrows or pores helps classify the flowering plants, with eudicots having three colpi, other groups having one sulcus. Pollen apertures are any modification of the wall of the pollen grain; these modifications include thinning and pores, they serve as an exit for the pollen contents and allow shrinking and swelling of the grain caused by changes in moisture content. The elongated apertures/ furrows in the pollen grain are called colpi, along with pores, are a chief criterion for identifying the pollen classes; the eudicots can be divided into two groups: the basal eudicots and the core eudicots. Basal eudicot is an informal name for a paraphyletic group; the core eudicots are a monophyletic group.
A 2010 study suggested the core eudicots can be divided into two clades, Gunnerales and a clade called "Pentapetalae", comprising all the remaining core eudicots. The Pentapetalae can be divided into three clades: Dilleniales superrosids consisting of Saxifragales and rosids superasterids consisting of Santalales, Berberidopsidales and asteridsThis division of the eudicots is shown in the following cladogram: The following is a more detailed breakdown according to APG IV, showing within each clade and orders: clade Eudicots order Ranunculales order Proteales order Trochodendrales order Buxales clade Core eudicots order Gunnerales order Dilleniales clade Superrosids order Saxifragales clade Rosids order Vitales clade Fabids order Fabales order Rosales order Fagales order Cucurbitales order Oxalidales order Malpighiales order Celastrales order Zygophyllales clade Malvids order Geraniales order Myrtales order Crossosomatales order Picramniales order Malvales order Brassicales order Huerteales order Sapindales clade Superasterids order Berberidopsidales order Santalales order Caryophyllales clade Asterids order Cornales order Ericales clade Campanulids order Aquifoliales order Asterales order Escalloniales order Bruniales order Apiales order Dipsacales order Paracryphiales clade Lamiids order Solanales order Lamiales order Vahliales order Gentianales order Boraginales order Garryales order Metteniusales order Icacinales Eudicots at the Encyclopedia of Life Eudicots, Tree of Life Web Project Dicots Plant Life Forms
Wikispecies is a wiki-based online project supported by the Wikimedia Foundation. Its aim is to create a comprehensive free content catalogue of all species. Jimmy Wales stated that editors are not required to fax in their degrees, but that submissions will have to pass muster with a technical audience. Wikispecies is available under the GNU Free Documentation License and CC BY-SA 3.0. Started in September 2004, with biologists across the world invited to contribute, the project had grown a framework encompassing the Linnaean taxonomy with links to Wikipedia articles on individual species by April 2005. Benedikt Mandl co-ordinated the efforts of several people who are interested in getting involved with the project and contacted potential supporters in early summer 2004. Databases were evaluated and the administrators contacted, some of them have agreed on providing their data for Wikispecies. Mandl defined two major tasks: Figure out how the contents of the data base would need to be presented—by asking experts, potential non-professional users and comparing that with existing databases Figure out how to do the software, which hardware is required and how to cover the costs—by asking experts, looking for fellow volunteers and potential sponsorsAdvantages and disadvantages were discussed by the wikimedia-I mailing list.
The board of directors of the Wikimedia Foundation voted by 4 to 0 in favor of the establishment of a Wikispecies. The project is hosted at species.wikimedia.org. It was merged to a sister project of Wikimedia Foundation on September 14, 2004. On October 10, 2006, the project exceeded 75,000 articles. On May 20, 2007, the project exceeded 100,000 articles with a total of 5,495 registered users. On September 8, 2008, the project exceeded 150,000 articles with a total of 9,224 registered users. On October 23, 2011, the project reached 300,000 articles. On June 16, 2014, the project reached 400,000 articles. On January 7, 2017, the project reached 500,000 articles. On October 30, 2018, the project reached 600,000 articles, a total of 1.12 million pages. Wikispecies comprises taxon pages, additionally pages about synonyms, taxon authorities, taxonomical publications, institutions or repositories holding type specimen. Wikispecies asks users to use images from Wikimedia Commons. Wikispecies does not allow the use of content.
All Species Foundation Catalogue of Life Encyclopedia of Life Tree of Life Web Project List of online encyclopedias The Plant List Wikispecies, The free species directory that anyone can edit Species Community Portal The Wikispecies Charter, written by Wales