A chordate is an animal constituting the phylum Chordata. During some period of their life cycle, chordates possess a notochord, a dorsal nerve cord, pharyngeal slits, an endostyle, a post-anal tail: these five anatomical features define this phylum. Chordates are bilaterally symmetric; the Chordata and Ambulacraria together form the superphylum Deuterostomia. Chordates are divided into three subphyla: Vertebrata. There are extinct taxa such as the Vetulicolia. Hemichordata has been presented as a fourth chordate subphylum, but now is treated as a separate phylum: hemichordates and Echinodermata form the Ambulacraria, the sister phylum of the Chordates. Of the more than 65,000 living species of chordates, about half are bony fish that are members of the superclass Osteichthyes. Chordate fossils have been found from as early as the Cambrian explosion, 541 million years ago. Cladistically, vertebrates - chordates with the notochord replaced by a vertebral column during development - are considered to be a subgroup of the clade Craniata, which consists of chordates with a skull.
The Craniata and Tunicata compose the clade Olfactores. Chordates form a phylum of animals that are defined by having at some stage in their lives all of the following anatomical features: A notochord, a stiff rod of cartilage that extends along the inside of the body. Among the vertebrate sub-group of chordates the notochord develops into the spine, in wholly aquatic species this helps the animal to swim by flexing its tail. A dorsal neural tube. In fish and other vertebrates, this develops into the spinal cord, the main communications trunk of the nervous system. Pharyngeal slits; the pharynx is the part of the throat behind the mouth. In fish, the slits are modified to form gills, but in some other chordates they are part of a filter-feeding system that extracts particles of food from the water in which the animals live. Post-anal tail. A muscular tail that extends backwards behind the anus. An endostyle; this is a groove in the ventral wall of the pharynx. In filter-feeding species it produces mucus to gather food particles, which helps in transporting food to the esophagus.
It stores iodine, may be a precursor of the vertebrate thyroid gland. There are soft constraints that separate chordates from certain other biological lineages, but are not part of the formal definition: All chordates are deuterostomes; this means. All chordates are based on a bilateral body plan. All chordates are coelomates, have a fluid filled body cavity called a coelom with a complete lining called peritoneum derived from mesoderm; the following schema is from the third edition of Vertebrate Palaeontology. The invertebrate chordate classes are from Fishes of the World. While it is structured so as to reflect evolutionary relationships, it retains the traditional ranks used in Linnaean taxonomy. Phylum Chordata †Vetulicolia? Subphylum Cephalochordata – Class Leptocardii Clade Olfactores Subphylum Tunicata – Class Ascidiacea Class Thaliacea Class Appendicularia Class Sorberacea Subphylum Vertebrata Infraphylum incertae sedis Cyclostomata Superclass'Agnatha' paraphyletic Class Myxini Class Petromyzontida or Hyperoartia Class †Conodonta Class †Myllokunmingiida Class †Pteraspidomorphi Class †Thelodonti Class †Anaspida Class †Cephalaspidomorphi Infraphylum Gnathostomata Class †Placodermi Class Chondrichthyes Class †Acanthodii Superclass Osteichthyes Class Actinopterygii Class Sarcopterygii Superclass Tetrapoda Class Amphibia Class Sauropsida Class Synapsida Craniates, one of the three subdivisions of chordates, all have distinct skulls.
They include the hagfish. Michael J. Benton commented that "craniates are characterized by their heads, just as chordates, or all deuterostomes, are by their tails". Most craniates are vertebrates; these consist of a series of bony or cartilaginous cylindrical vertebrae with neural arches that protect the spinal cord, with projections that link the vertebrae. However hagfish have incomplete braincases and no vertebrae, are therefore not regarded as vertebrates, but as members of the craniates, the group from which vertebrates are thought to have evolved; however the cladistic exclusion of hagfish from the vertebrates is controversial, as they ma
Paul Gervais full name François Louis Paul Gervaise was a French palaeontologist and entomologist. Gervais was born in Paris, where he obtained the diplomas of doctor of science and of medicine, in 1835 he began palaeontological research as assistant in the laboratory of comparative anatomy at the Muséum national d'Histoire naturelle. In 1841 he obtained the chair of zoology and comparative anatomy at the Faculty of Sciences in Montpellier, of which he was in 1856 appointed dean. In 1848-1852 appeared his important work Zoologie et paléontologie françaises, supplementary to the palaeontological publications of Georges Cuvier and Henri Marie Ducrotay de Blainville. In 1865 he accepted the professorship of zoology at the Sorbonne, vacant through the death of Louis Pierre Gratiolet, he died in Paris on 10 February 1879. According to Florentino Ameghino, Paul Gervais studied a fossil collection obtained from Juan Manuel de Rosas the governor of Buenos Aires. Earlier on, this collection would have been donated or donated-by-force to the Buenos Aires Province by Francisco Javier Muñiz.
Gervais is commemorated in the vernacular and scientific names of the following taxa: a species of snake, Calamaria gervaisii. A species of cetacean, Gervais' beaked whale. Apart from the works mentioned he wrote: with Charles Athanase Walckenaer Histoire naturelle des insectes Zoologie et paléontologie françaises Histoire naturelle des Mammifères Zoologie médicale, mit Pierre-Joseph van Beneden Recherches sur l'ancienneté de l'homme et la période quaternaire Zoologie et Paléontologie générales Ostéographie des cétacés vivants et fossiles,with Pierre-Joseph van Beneden Brachistosternus ehrenbergii Gervais' beaked whale This article incorporates text from a publication now in the public domain: Chisholm, Hugh, ed.. "Gervais, Paul". Encyclopædia Britannica. 11. Cambridge University Press. P. 907
International Union for Conservation of Nature
The International Union for Conservation of Nature is an international organization working in the field of nature conservation and sustainable use of natural resources. It is involved in data gathering and analysis, field projects and education. IUCN's mission is to "influence and assist societies throughout the world to conserve nature and to ensure that any use of natural resources is equitable and ecologically sustainable". Over the past decades, IUCN has widened its focus beyond conservation ecology and now incorporates issues related to sustainable development in its projects. Unlike many other international environmental organisations, IUCN does not itself aim to mobilize the public in support of nature conservation, it tries to influence the actions of governments and other stakeholders by providing information and advice, through building partnerships. The organization is best known to the wider public for compiling and publishing the IUCN Red List of Threatened Species, which assesses the conservation status of species worldwide.
IUCN has a membership of over 1400 non-governmental organizations. Some 16,000 scientists and experts participate in the work of IUCN commissions on a voluntary basis, it employs 1000 full-time staff in more than 50 countries. Its headquarters are in Switzerland. IUCN has observer and consultative status at the United Nations, plays a role in the implementation of several international conventions on nature conservation and biodiversity, it was involved in establishing the World Wide Fund for Nature and the World Conservation Monitoring Centre. In the past, IUCN has been criticized for placing the interests of nature over those of indigenous peoples. In recent years, its closer relations with the business sector have caused controversy. IUCN was established in 1948, it was called the International Union for the Protection of Nature and the World Conservation Union. Establishment IUCN was established on 5 October 1948, in Fontainebleau, when representatives of governments and conservation organizations signed a formal act constituting the International Union for the Protection of Nature.
The initiative to set up the new organisation came from UNESCO and from its first Director General, the British biologist Julian Huxley. The objectives of the new Union were to encourage international cooperation in the protection of nature, to promote national and international action and to compile and distribute information. At the time of its founding IUPN was the only international organisation focusing on the entire spectrum of nature conservation Early years: 1948–1956 IUPN started out with 65 members, its secretariat was located in Brussels. Its first work program focused on saving species and habitats and applying knowledge, advancing education, promoting international agreements and promoting conservation. Providing a solid scientific base for conservation action was the heart of all activities. IUPN and UNESCO were associated, they jointly organized the 1949 Conference on Protection of Nature. In preparation for this conference a list of gravely endangered species was drawn up for the first time, a precursor of the IUCN Red List of Threatened Species.
In the early years of its existence IUCN depended entirely on UNESCO funding and was forced to temporarily scale down activities when this ended unexpectedly in 1954. IUPN was successful in engaging prominent scientists and identifying important issues such as the harmful effects of pesticides on wildlife but not many of the ideas it developed were turned into action; this was caused by unwillingness to act on the part of governments, uncertainty about the IUPN mandate and lack of resources. In 1956, IUPN changed its name to International Union for Conservation of Nature and Natural Resources. Increased profile and recognition: 1956–1965 In the 1950s and 1960s Europe entered a period of economic growth and formal colonies became independent. Both developments had impact on the work of IUCN. Through the voluntary involvement of experts in its Commissions IUCN was able to get a lot of work done while still operating on a low budget, it established links with the Council of Europe. In 1961, at the request of United Nations Economic and Social Council, the United Nations Economic and Social Council, IUCN published the first global list of national parks and protected areas which it has updated since.
IUCN's best known publication, the Red Data Book on the conservation status of species, was first published in 1964. IUCN began to play a part in the development of international treaties and conventions, starting with the African Convention on the Conservation of Nature and Natural Resources. Environmental law and policy making became a new area of expertise. Africa was the focus of many of the early IUCN conservation field projects. IUCN supported the ‘Yellowstone model’ of protected area management, which restricted human presence and activity in order to protect nature. IUCN and other conservation organisations were criticized for protecting nature against people rather than with people; this model was also applied in Africa and played a role in the decision to remove the Maasai people from Serengeti National Park and the Ngorongoro Conservation Area. To establish a stable financial basis for its work, IUCN participated in setting up the World Wildlife Fund
John Edward Gray
John Edward Gray, FRS was a British zoologist. He was the elder brother of zoologist George Robert Gray and son of the pharmacologist and botanist Samuel Frederick Gray; the standard author abbreviation J. E. Gray is used to indicate this person as the author. Or zoological name. Gray was Keeper of Zoology at the British Museum in London from 1840 until Christmas 1874, before the Natural History holdings were split off to the Natural History Museum, he published several catalogues of the museum collections that included comprehensive discussions of animal groups as well as descriptions of new species. He improved the zoological collections to make them amongst the best in the world. Gray was born in Walsall, he assisted his father in writing The Natural Arrangement of British Plants. After being blackballed by the Linnean Society of London, Gray shifted his interest from botany to zoology, he began his zoological career by volunteering to collect insects for the British Museum at age 15. He joined the Zoological Department in 1824 to help John George Children catalog the reptile collection.
In some of his early articles, Gray adopted William Sharp Macleay's quinarian system for classifications of molluscs, echinoderms and mammals. In 1840 he took over Children's position as Keeper of Zoology, which he held for 35 years, publishing well over 1,000 papers, he named many cetacean species, genera and families. During this period he collaborated with Benjamin Waterhouse Hawkins, the noted natural history artist, in producing Gleanings from the Menagerie at Knowsley; the menagerie at Knowsley Hall, near Liverpool, founded by Edward Smith-Stanley, 13th Earl of Derby, at the Stanley ancestral seat, was one of the largest private menageries in Victorian England. Gray married Maria Emma Smith in 1826, she helped him with his scientific work with her drawings. In 1833, Gray was a founder of. Gray was a friend of the coleopterist Hamlet Clark, in 1856-57 they voyaged on Gray's yacht Miranda to Spain and Brazil. Gray was an accomplished watercolourist, his landscape paintings illustrate Clark's account of their journeys.
Gray was interested in postage stamps. On 1 May 1840, the day the Penny Black first went on sale, he purchased several with the intent to save them. During his fifty years employed at the British Museum Gray wrote nearly 500 papers, including many descriptions of species new to science; these had been presented to the Museum by collectors from around the world, included all branches of zoology, although Gray left the descriptions of new birds to his younger brother and colleague George. Gray was active in malacology, the study of molluscs. John Edward Gray was buried at Lewisham. Gray was one of the most prolific taxonomists in the history of zoology, he described more than 300 species and subspecies of reptiles, only surpassed by his successors at the British Museum, George A. Boulenger and Albert Günther and American zoologist Edward D. Cope. Gray described and named numerous marine snails including: The genus Lithopoma Gray, 1850 The genus Euthria Gray, 1850Genera named in his honour include: The snake genus Grayia Günther, 1858Species and subspecies named in his honour include: Ardeola grayii – Indian pond heron Mesoplodon grayi von Haast, 1876 – Gray's beaked whale Crocidura grayi Dobson, 1890 – Luzon shrew Ablepharus grayanus Delma grayii A. Smith, 1849 Microlophus grayii Naultinus grayii Bell, 1843 Salvelinus grayi Günther, 1862 Tropidophorus grayi Günther, 1861 Trachemys venusta grayi 1821: "A natural arrangement of Mollusca, according to their internal structure."
London Medical Repository 15: 229–239. 1821: "On the natural arrangement of Vertebrose Animals." London Medical Repository 15: 296–310. 1824: "A revision of the family Equidae." Zool. J. Lond. 1: 241-248 pl. 9. 1824: "On the natural arrangement of the pulmonobranchous Mollusca." Annals of Philosophy, 8: 107–109. 1824: "On the arrangement of the Papilionidae." Annals of Philosophy 8: 119-120. 1825: "A list and description of some species of shells not taken notice of by Lamarck." Annals of Philosophy 9: 407-415. 1825: "A synopsis of the genera of reptiles and Amphibia, with a description of some new species." Annals of Philosophy 10: 193-217. 1825: "An outline of an attempt at the disposition of the Mammalia into tribes and families with a list of the genera appertaining to each tribe." Annals of Philosophy 10: 337-344. 1825: "An attempt to divide the Echinida, or sea eggs, into natural families." Annals of Philosophy 10: 423-431. 1826: "Vertebrata. Mammalia.". P. 412-415 in King, P. P. Narrative of a Survey of the Intertropical and Western Coasts of Australia.
Performed between the years 1818 and 1822. With an Appendix, containing various subjects relating to hydrography and natural history. London: J. Murray Vol. 2. 1827: "Synopsis of the species of the class Mammalia." P. 1-391 in Baron Cuvier The Animal Kingdom Arranged in Conformity with its Organization, by the Baron Cuvier, with additional descriptions by Edward Griffith and others.. London: George B. Whittaker Vol. 5. 1828: "Spicilegia Zoologica, or original figures and short systematic descriptions of new and unfigured animals." Pt 1. London: Treuttel, Würtz & Co. 1829: "An attempt to improve the natural arrangement of the genera of bat, from actual examination. Phil. Mag
Binomial nomenclature called binominal nomenclature or binary nomenclature, is a formal system of naming species of living things by giving each a name composed of two parts, both of which use Latin grammatical forms, although they can be based on words from other languages. Such a name is called a binomen, binominal name or a scientific name; the first part of the name – the generic name – identifies the genus to which the species belongs, while the second part – the specific name or specific epithet – identifies the species within the genus. For example, humans belong within this genus to the species Homo sapiens. Tyrannosaurus rex is the most known binomial; the formal introduction of this system of naming species is credited to Carl Linnaeus beginning with his work Species Plantarum in 1753. But Gaspard Bauhin, in as early as 1623, had introduced in his book Pinax theatri botanici many names of genera that were adopted by Linnaeus; the application of binomial nomenclature is now governed by various internationally agreed codes of rules, of which the two most important are the International Code of Zoological Nomenclature for animals and the International Code of Nomenclature for algae and plants.
Although the general principles underlying binomial nomenclature are common to these two codes, there are some differences, both in the terminology they use and in their precise rules. In modern usage, the first letter of the first part of the name, the genus, is always capitalized in writing, while that of the second part is not when derived from a proper noun such as the name of a person or place. Both parts are italicized when a binomial name occurs in normal text, thus the binomial name of the annual phlox is now written as Phlox drummondii. In scientific works, the authority for a binomial name is given, at least when it is first mentioned, the date of publication may be specified. In zoology "Patella vulgata Linnaeus, 1758"; the name "Linnaeus" tells the reader who it was that first published a description and name for this species of limpet. "Passer domesticus". The original name given by Linnaeus was Fringilla domestica; the ICZN does not require that the name of the person who changed the genus be given, nor the date on which the change was made, although nomenclatorial catalogs include such information.
In botany "Amaranthus retroflexus L." – "L." is the standard abbreviation used in botany for "Linnaeus". "Hyacinthoides italica Rothm. – Linnaeus first named this bluebell species Scilla italica. The name is composed of two word-forming elements: "bi", a Latin prefix for two, "-nomial", relating to a term or terms; the word "binomium" was used in Medieval Latin to mean a two-term expression in mathematics. Prior to the adoption of the modern binomial system of naming species, a scientific name consisted of a generic name combined with a specific name, from one to several words long. Together they formed a system of polynomial nomenclature; these names had two separate functions. First, to designate or label the species, second, to be a diagnosis or description. In a simple genus, containing only two species, it was easy to tell them apart with a one-word genus and a one-word specific name; such "polynomial names" may sometimes look like binomials, but are different. For example, Gerard's herbal describes various kinds of spiderwort: "The first is called Phalangium ramosum, Branched Spiderwort.
The other... is aptly termed Phalangium Ephemerum Virginianum, Soon-Fading Spiderwort of Virginia". The Latin phrases are short descriptions, rather than identifying labels; the Bauhins, in particular Caspar Bauhin, took some important steps towards the binomial system, by pruning the Latin descriptions, in many cases to two words. The adoption by biologists of a system of binomial nomenclature is due to Swedish botanist and physician Carl von Linné, more known by his Latinized name Carl Linnaeus, it was in his 1753 Species Plantarum that he first began using a one-word "trivial name" together with a generic name in a system of binomial nomenclature. This trivial name is what is now known as specific name; the Bauhins' genus names were retained in many of these, but the descriptive part was reduced to a single word. Linnaeus's trivial names introduced an important new idea, namely that the function of a name could be to give a species a unique label; this meant. Thus Gerard's Phalangium ephemerum virginianum became Tradescantia virgi
South America is a continent in the Western Hemisphere in the Southern Hemisphere, with a small portion in the Northern Hemisphere. It may be considered a subcontinent of the Americas, how it is viewed in the Spanish and Portuguese-speaking regions of the Americas; the reference to South America instead of other regions has increased in the last decades due to changing geopolitical dynamics. It is bordered on the west on the north and east by the Atlantic Ocean, it includes twelve sovereign states, a part of France, a non-sovereign area. In addition to this, the ABC islands of the Kingdom of the Netherlands and Tobago, Panama may be considered part of South America. South America has an area of 17,840,000 square kilometers, its population as of 2016 has been estimated at more than 420 million. South America ranks fourth in fifth in population. Brazil is by far the most populous South American country, with more than half of the continent's population, followed by Colombia, Argentina and Peru. In recent decades Brazil has concentrated half of the region's GDP and has become a first regional power.
Most of the population lives near the continent's western or eastern coasts while the interior and the far south are sparsely populated. The geography of western South America is dominated by the Andes mountains. Most of the continent lies in the tropics; the continent's cultural and ethnic outlook has its origin with the interaction of indigenous peoples with European conquerors and immigrants and, more locally, with African slaves. Given a long history of colonialism, the overwhelming majority of South Americans speak Portuguese or Spanish, societies and states reflect Western traditions. South America occupies the southern portion of the Americas; the continent is delimited on the northwest by the Darién watershed along the Colombia–Panama border, although some may consider the border instead to be the Panama Canal. Geopolitically and geographically all of Panama – including the segment east of the Panama Canal in the isthmus – is included in North America alone and among the countries of Central America.
All of mainland South America sits on the South American Plate. South America is home to Angel Falls in Venezuela. South America's major mineral resources are gold, copper, iron ore and petroleum; these resources found in South America have brought high income to its countries in times of war or of rapid economic growth by industrialized countries elsewhere. However, the concentration in producing one major export commodity has hindered the development of diversified economies; the fluctuation in the price of commodities in the international markets has led to major highs and lows in the economies of South American states causing extreme political instability. This is leading to efforts to diversify production to drive away from staying as economies dedicated to one major export. South America is one of the most biodiverse continents on earth. South America is home to many interesting and unique species of animals including the llama, piranha, vicuña, tapir; the Amazon rainforests possess high biodiversity, containing a major proportion of the Earth's species.
Brazil is the largest country in South America, encompassing around half of the continent's land area and population. The remaining countries and territories are divided among three regions: The Andean States, the Guianas and the Southern Cone. Traditionally, South America includes some of the nearby islands. Aruba, Curaçao, Trinidad and the federal dependencies of Venezuela sit on the northerly South American continental shelf and are considered part of the continent. Geo-politically, the island states and overseas territories of the Caribbean are grouped as a part or subregion of North America, since they are more distant on the Caribbean Plate though San Andres and Providencia are politically part of Colombia and Aves Island is controlled by Venezuela. Other islands that are included with South America are the Galápagos Islands that belong to Ecuador and Easter Island, Robinson Crusoe Island, Chiloé and Tierra del Fuego. In the Atlantic, Brazil owns Fernando de Noronha and Martim Vaz, the Saint Peter and Saint Paul Archipelago, while the Falkland Islands are governed by the United Kingdom, whose sovereignty over the islands is disputed by Argentina.
South Georgia and the South Sandwich Islands may be associate
Mammals are vertebrate animals constituting the class Mammalia, characterized by the presence of mammary glands which in females produce milk for feeding their young, a neocortex, fur or hair, three middle ear bones. These characteristics distinguish them from reptiles and birds, from which they diverged in the late Triassic, 201–227 million years ago. There are around 5,450 species of mammals; the largest orders are the rodents and Soricomorpha. The next three are the Primates, the Cetartiodactyla, the Carnivora. In cladistics, which reflect evolution, mammals are classified as endothermic amniotes, they are the only living Synapsida. The early synapsid mammalian ancestors were sphenacodont pelycosaurs, a group that produced the non-mammalian Dimetrodon. At the end of the Carboniferous period around 300 million years ago, this group diverged from the sauropsid line that led to today's reptiles and birds; the line following the stem group Sphenacodontia split off several diverse groups of non-mammalian synapsids—sometimes referred to as mammal-like reptiles—before giving rise to the proto-mammals in the early Mesozoic era.
The modern mammalian orders arose in the Paleogene and Neogene periods of the Cenozoic era, after the extinction of non-avian dinosaurs, have been among the dominant terrestrial animal groups from 66 million years ago to the present. The basic body type is quadruped, most mammals use their four extremities for terrestrial locomotion. Mammals range in size from the 30–40 mm bumblebee bat to the 30-meter blue whale—the largest animal on the planet. Maximum lifespan varies from two years for the shrew to 211 years for the bowhead whale. All modern mammals give birth to live young, except the five species of monotremes, which are egg-laying mammals; the most species-rich group of mammals, the cohort called placentals, have a placenta, which enables the feeding of the fetus during gestation. Most mammals are intelligent, with some possessing large brains, self-awareness, tool use. Mammals can communicate and vocalize in several different ways, including the production of ultrasound, scent-marking, alarm signals and echolocation.
Mammals can organize themselves into fission-fusion societies and hierarchies—but can be solitary and territorial. Most mammals are polygynous. Domestication of many types of mammals by humans played a major role in the Neolithic revolution, resulted in farming replacing hunting and gathering as the primary source of food for humans; this led to a major restructuring of human societies from nomadic to sedentary, with more co-operation among larger and larger groups, the development of the first civilizations. Domesticated mammals provided, continue to provide, power for transport and agriculture, as well as food and leather. Mammals are hunted and raced for sport, are used as model organisms in science. Mammals have been depicted in art since Palaeolithic times, appear in literature, film and religion. Decline in numbers and extinction of many mammals is driven by human poaching and habitat destruction deforestation. Mammal classification has been through several iterations since Carl Linnaeus defined the class.
No classification system is universally accepted. George Gaylord Simpson's "Principles of Classification and a Classification of Mammals" provides systematics of mammal origins and relationships that were universally taught until the end of the 20th century. Since Simpson's classification, the paleontological record has been recalibrated, the intervening years have seen much debate and progress concerning the theoretical underpinnings of systematization itself through the new concept of cladistics. Though field work made Simpson's classification outdated, it remains the closest thing to an official classification of mammals. Most mammals, including the six most species-rich orders, belong to the placental group; the three largest orders in numbers of species are Rodentia: mice, porcupines, beavers and other gnawing mammals. The next three biggest orders, depending on the biological classification scheme used, are the Primates including the apes and lemurs. According to Mammal Species of the World, 5,416 species were identified in 2006.
These were grouped into 153 families and 29 orders. In 2008, the International Union for Conservation of Nature completed a five-year Global Mammal Assessment for its IUCN Red List, which counted 5,488 species. According to a research published in the Journal of Mammalogy in 2018, the number of recognized mammal species is 6,495 species included 96 extinct; the word "mammal" is modern, from the scientific name Mammalia coined by Carl Linnaeus in 1758, derived from the Latin mamma. In an influential 1988 paper, Timothy Rowe defined Mammalia phylogenetically as the crown group of mammals, the clade consisting of the most recent common ancestor of living monotremes and therian m