In zoological nomenclature, a type species is the species name with which the name of a genus or subgenus is considered to be permanently taxonomically associated, i.e. the species that contains the biological type specimen. A similar concept is used for suprageneric groups called a type genus. In botanical nomenclature, these terms have no formal standing under the code of nomenclature, but are sometimes borrowed from zoological nomenclature. In botany, the type of a genus name is a specimen, the type of a species name; the species name that has that type can be referred to as the type of the genus name. Names of genus and family ranks, the various subdivisions of those ranks, some higher-rank names based on genus names, have such types. In bacteriology, a type species is assigned for each genus; every named genus or subgenus in zoology, whether or not recognized as valid, is theoretically associated with a type species. In practice, there is a backlog of untypified names defined in older publications when it was not required to specify a type.
A type species is both a concept and a practical system, used in the classification and nomenclature of animals. The "type species" represents the reference species and thus "definition" for a particular genus name. Whenever a taxon containing multiple species must be divided into more than one genus, the type species automatically assigns the name of the original taxon to one of the resulting new taxa, the one that includes the type species; the term "type species" is regulated in zoological nomenclature by article 42.3 of the International Code of Zoological Nomenclature, which defines a type species as the name-bearing type of the name of a genus or subgenus. In the Glossary, type species is defined as The nominal species, the name-bearing type of a nominal genus or subgenus; the type species permanently attaches a formal name to a genus by providing just one species within that genus to which the genus name is permanently linked. The species name in turn is fixed, to a type specimen. For example, the type species for the land snail genus Monacha is Helix cartusiana, the name under which the species was first described, known as Monacha cartusiana when placed in the genus Monacha.
That genus is placed within the family Hygromiidae. The type genus for that family is the genus Hygromia; the concept of the type species in zoology was introduced by Pierre André Latreille. The International Code of Zoological Nomenclature states that the original name of the type species should always be cited, it gives an example in Article 67.1. Astacus marinus Fabricius, 1775 was designated as the type species of the genus Homarus, thus giving it the name Homarus marinus. However, the type species of Homarus should always be cited using its original name, i.e. Astacus marinus Fabricius, 1775. Although the International Code of Nomenclature for algae and plants does not contain the same explicit statement, examples make it clear that the original name is used, so that the "type species" of a genus name need not have a name within that genus, thus in Article 10, Ex. 3, the type of the genus name Elodes is quoted as the type of the species name Hypericum aegypticum, not as the type of the species name Elodes aegyptica.
Glossary of scientific naming Genetypes – genetic sequence data from type specimens. Holotype Paratype Principle of Typification Type Type genus
Paralititan was a giant titanosaurian sauropod dinosaur genus discovered in coastal deposits in the Upper Cretaceous Bahariya Formation of Egypt. It lived between 93.5 million years ago. Joshua Smith, who informally led the research team that found the dinosaur fossils, told an interviewer, "It was a enormous dinosaur by any reckoning."Little of Paralititan is known, so its exact size is difficult to estimate. However, the limited material the long humeri, suggested that it is one of the most massive dinosaurs discovered, with an estimated weight of 59 t; the complete right humerus measured 1.69 meters long which at the time of discovery was the longest known in a Cretaceous sauropod. Using Saltasaurus as a guide, Carpenter estimated its length at around 26 m. Scott Hartman estimates an animal, massive, but still smaller than the biggest titanosaurs such as Puertasaurus and Argentinosaurus. In 2010, Gregory S. Paul estimated the length at the weight at twenty tonnes. From the formation another sauropod had been known, Aegyptosaurus.
Paralititan differs from Aegyptosaurus in its larger size, the latter genus weighing only fifteen tons in not having pleurocoels in its front tail vertebrae, in possessing a longer deltopectoral crest on its humerus. Joshua Smith in 1999 in the Bahariya Oasis rediscovered the Gebel el Dist site where Richard Markgraf in 1912, 1913 and 1914 had excavated fossils for Ernst Stromer. In 2000, an American expedition was mounted to revisit the site; however Markgraf had removed all more complete skeletons, leaving only limited remains behind. At a new site, the nearby Gebel Fagga, the expedition succeeded in locating a partial sauropod skeleton, it was identified by Lacovara as a species new to science. It was named and described by Joshua B. Smith, Matthew C. Lamanna, Kenneth J. Lacovara, Peter Dodson, Jennifer R. Smith, Jason Charles Poole, Robert Giegengack and Yousri Attia in 2001 as the type species Paralititan stromeri; the generic name means "Stromer's tidal titan" or "Stromer's tidal giant", in reference to the "paralic" tidal flats the animal lived on.
The specific name honors Ernst Stromer von Reichenbach, a German paleontologist and geologist who first established the presence of dinosaur fossils in this area in 1911. Paralititan represents the first tetrapod reported from the Bahariya Formation since Romer's publication of 1935; the holotype specimen of Paralititan, CGM 81119, was found in a layer of the Bahariya Formation, dating from the Cenomanian. It consists of a partial skeleton lacking the skull, it is incomplete, apart from bone fragments containing two fused posterior sacral vertebrae, two anterior caudal vertebrae, both incomplete scapulae, two humeri and a metacarpal. The Paralititan type specimen shows evidence of having been scavenged by a carnivorous dinosaur as it was disarticulated within an oval of eight metres length with the various bones being clustered. A Carcharodontosaurus tooth was discovered in between the clusters; the holotype is part of the collection of the Cairo Geological Museum. The large anterior dorsal vertebra 1912V11164, in 1932 by Stromer referred to an undetermined "Giant Sauropod", was in 2001 tentatively referred to Paralititan.
The autochthonous, scavenged skeleton was preserved in tidal flat deposits containing in the form of fossil leaves and root systems, a mangrove vegetation of seed ferns, Weichselia reticulata. The mangrove ecosystem it inhabited was situated along the southern shore of the Tethys Sea. Paralititan is the first dinosaur demonstrated to have inhabited a mangrove habitat, it lived at the same time and place as giant predators Carcharodontosaurus and the sauropod Aegyptosaurus. Paralititan in the Dino Directory
Sauropodomorpha is an extinct clade of long-necked, saurischian dinosaurs that includes the sauropods and their ancestral relatives. Sauropods grew to large sizes, had long necks and tails, were quadrupedal, became the largest animals to walk the Earth; the "prosauropods", which preceded the sauropods, were smaller and were able to walk on two legs. The sauropodomorphs were the dominant terrestrial herbivores throughout much of the Mesozoic Era, from their origins in the mid-Triassic until their decline and extinction at the end of the Cretaceous. Sauropodomorphs were adapted to browsing higher than any other contemporary herbivore, giving them access to high tree foliage; this feeding strategy is supported by many of their defining characteristics, such as: a light, tiny skull on the end of a long neck and a counterbalancing long tail. Their teeth were weak, shaped like leaves or spoons. Instead of grinding teeth, they had stomach stones, similar to the gizzard stones of modern birds and crocodiles, to help digest tough plant fibers.
The front of the upper mouth bends down in. One of the earliest known sauropodomorphs, was small and slender; the largest sauropods, like Supersaurus, Diplodocus hallorum and Argentinosaurus, reached 30–40 metres in length, 60,000–100,000 kilograms or more in mass. Bipedal, as their size increased they evolved a four-legged graviportal gait adapted only to walking on land, like elephants; the early sauropodomorphs were most omnivores as their shared common ancestor with the other saurischian lineage was a carnivore. Therefore, their evolution to herbivory went hand in hand with their increasing size and neck length, they had large nostrils, retained a thumb with a big claw, which may have been used for defense — though their primary defensive adaptation was their extreme size. Sauropodomorphs can be distinguished as a group on the basis of some of the following synapomorphies: The presence of large nares; the distal part of the tibia is covered by an ascending process of the astragalus. Their hind limbs are short.
The presence of three or more sacral vertebrae. The teeth are thin and are spatula-like, with bladed and serrated crowns; the presence of a minimum of 10 cervical vertebrae that are elongated The presence of 25 presacral vertebrae The manus had a large digit I Among the first dinosaurs to evolve in the Late Triassic Period, about 230 million years ago, they became the dominant herbivores by halfway through the late Triassic. Their perceived decline in the early Cretaceous is most a bias in fossil sampling, as most fossils are known from Europe and North America. Sauropods were still the dominant herbivores in the Gondwanan landmasses, however; the spread of flowering plants and "advanced" ornithischians, another major group of herbivorous dinosaurs, are most not a major factor in sauropod decline in the northern continents. Like all non-avian dinosaurs, the sauropodomorphs became extinct 66 Mya, during the Cretaceous–Paleogene extinction event; the earliest and most basal sauropodomorphs known are Chromogisaurus novasi and Panphagia protos, both from the Ischigualasto Formation, dated to 231.4 million years ago.
Some studies have found Eoraptor lunensis, traditionally considered a theropod, to be an early member of the sauropodomorph lineage, which would make it the most basal sauropodomorph known. Sauropodomorpha is one of the two major clades within the order Saurischia; the sauropodomorphs' sister group, the Theropoda, includes bipedal carnivores like Velociraptor and Tyrannosaurus. However, sauropodomorphs share a number of characteristics with the Ornithischia, so a small minority of palaeontologists, like Bakker, have placed both sets of herbivores within a group called "Phytodinosauria" or "Ornithischiformes". In Linnaean taxonomy, Sauropodomorpha is left unranked, it was established by Friedrich von Huene in 1932, who broke it into two groups: the basal forms within Prosauropoda, their descendants, the giant Sauropoda. Phylogenetic analyses by Adam Yates and others placed Sauropoda within a paraphyletic "Prosauropoda". Recent cladistic analyses suggest that the clade Prosauropoda, named by Huene in 1920 and was defined by Sereno, in 1998, as all animals more related to Plateosaurus engelhardti than to Saltasaurus loricatus, is a junior synonym of Plateosauridae as both contain the same taxa.
Most modern classification schemes break the prosauropods into a half-dozen groups that evolved separately from one common lineage. While they have a number of shared characteristics, the evolutionary requirements for giraffe-like browsing high in the trees may have caused convergent evolution, where similar traits evolve separately because they faced the same evolutionary pressure, instead of trait
Spain the Kingdom of Spain, is a country located in Europe. Its continental European territory is situated on the Iberian Peninsula, its territory includes two archipelagoes: the Canary Islands off the coast of Africa, the Balearic Islands in the Mediterranean Sea. The African enclaves of Ceuta, Peñón de Vélez de la Gomera make Spain the only European country to have a physical border with an African country. Several small islands in the Alboran Sea are part of Spanish territory; the country's mainland is bordered to the south and east by the Mediterranean Sea except for a small land boundary with Gibraltar. With an area of 505,990 km2, Spain is the largest country in Southern Europe, the second largest country in Western Europe and the European Union, the fourth largest country in the European continent. By population, Spain is the fifth in the European Union. Spain's capital and largest city is Madrid. Modern humans first arrived in the Iberian Peninsula around 35,000 years ago. Iberian cultures along with ancient Phoenician, Greek and Carthaginian settlements developed on the peninsula until it came under Roman rule around 200 BCE, after which the region was named Hispania, based on the earlier Phoenician name Spn or Spania.
At the end of the Western Roman Empire the Germanic tribal confederations migrated from Central Europe, invaded the Iberian peninsula and established independent realms in its western provinces, including the Suebi and Vandals. The Visigoths would forcibly integrate all remaining independent territories in the peninsula, including Byzantine provinces, into the Kingdom of Toledo, which more or less unified politically and all the former Roman provinces or successor kingdoms of what was documented as Hispania. In the early eighth century the Visigothic Kingdom fell to the Moors of the Umayyad Islamic Caliphate, who arrived to rule most of the peninsula in the year 726, leaving only a handful of small Christian realms in the north and lasting up to seven centuries in the Kingdom of Granada; this led to many wars during a long reconquering period across the Iberian Peninsula, which led to the creation of the Kingdom of Leon, Kingdom of Castile, Kingdom of Aragon and Kingdom of Navarre as the main Christian kingdoms to face the invasion.
Following the Moorish conquest, Europeans began a gradual process of retaking the region known as the Reconquista, which by the late 15th century culminated in the emergence of Spain as a unified country under the Catholic Monarchs. Until Aragon had been an independent kingdom, which had expanded toward the eastern Mediterranean, incorporating Sicily and Naples, had competed with Genoa and Venice. In the early modern period, Spain became the world's first global empire and the most powerful country in the world, leaving a large cultural and linguistic legacy that includes more than 570 million Hispanophones, making Spanish the world's second-most spoken native language, after Mandarin Chinese. During the Golden Age there were many advancements in the arts, with world-famous painters such as Diego Velázquez; the most famous Spanish literary work, Don Quixote, was published during the Golden Age. Spain hosts the world's third-largest number of UNESCO World Heritage Sites. Spain is a secular parliamentary democracy and a parliamentary monarchy, with King Felipe VI as head of state.
It is a major developed country and a high income country, with the world's fourteenth largest economy by nominal GDP and sixteenth largest by purchasing power parity. It is a member of the United Nations, the European Union, the Eurozone, the Council of Europe, the Organization of Ibero-American States, the Union for the Mediterranean, the North Atlantic Treaty Organization, the Organisation for Economic Co-operation and Development, Organization for Security and Co-operation in Europe, the Schengen Area, the World Trade Organization and many other international organisations. While not an official member, Spain has a "Permanent Invitation" to the G20 summits, participating in every summit, which makes Spain a de facto member of the group; the origins of the Roman name Hispania, from which the modern name España was derived, are uncertain due to inadequate evidence, although it is documented that the Phoenicians and Carthaginians referred to the region as Spania, therefore the most accepted etymology is a Semitic-Phoenician one.
Down the centuries there have been a number of accounts and hypotheses: The Renaissance scholar Antonio de Nebrija proposed that the word Hispania evolved from the Iberian word Hispalis, meaning "city of the western world". Jesús Luis Cunchillos argues that the root of the term span is the Phoenician word spy, meaning "to forge metals". Therefore, i-spn-ya would mean "the land where metals are forged", it may be a derivation of the Phoenician I-Shpania, meaning "island of rabbits", "land of rabbits" or "edge", a reference to Spain's location at the end of the Mediterranean. The word in question means "Hyrax" due to Phoenicians confusing the two animals. Hispania may derive from the poetic use of the term Hesperia, reflecting the Greek perception of Italy as a "western land" or "land of the setting sun" (Hesperia
Sanpasaurus is a poorly known sauropod dinosaur from the Early Jurassic of Sichuan, China. The type species, S. yaoi, was described by Chung Chien Young, in 1944. The type remains, IVPP V.156, consists of 20 vertebrae, scapulae and some hindlimb bones. Reported by Young as an ornithopod ornithischian, this specimen was unambiguously referred to Sauropoda in 2016 by McPhee et al. Sansapasaurus is known from remains recovered from the Maanshan Member of the Ziliujing Formation
Isanosaurus was one of the first sauropod dinosaurs. It lived 210 million years ago during the Late Triassic in Thailand; the only species is Isanosaurus attavipachi. Though important for the understanding of sauropod origin and early evolution, Isanosaurus is poorly known. Exact relationships to other early sauropods remain unresolved; the only specimen includes a neck vertebra, a back vertebra and part of a second, six tail vertebra, two chevrons, fragmentary ribs, the right sternal plate, the right shoulder blade, the left thigh bone. This individual may have measured 6.5 metres. However, the vertebral neural arches have been found separated from the vertebral centra, indicating that these elements were not fused with each other. Early sauropodomorphs were primitively bipedal. Isanosaurus, being one of the first sauropods known shows a quadrupedal locomotion; the legs were column-like, as indicated by the straight thigh bone. In prosauropods, but in the basal sauropod Antetonitrus, the thigh bone was sigmoidal.
Like in other sauropods, bony processes of the femur were reduced in Isanosaurus. Additional important features can be found in the vertebrae; the neck vertebrae were distinctly opisthocoelous, forming ball-and-socket joints with neighbouring vertebrae. The tail vertebrae, on the other hand, were amphicoelous; the dorsal neural spines were high, like those of some sauropods, unlike the low prosauropod neural spines. The lateral sides of the vertebrae were concave, but not excavated as in sauropods; the specimen was found in dark red sandstone of the Nam Phong Formation near the village of Ban Non Thaworn, in Chaiyaphum Province. When discovered in 1998, the skeleton had been eroded away. With regard to vertebrate fossils, the Nam Phong Formation is poorly explored: besides Isanosaurus, only two articulated ischia were found. Whether these ischia belong to Isanosaurus is unclear, because no pelvic bones are preserved in the holotype specimen. Isanosaurus was described by French palaeontologist Éric Buffetaut and colleagues in 2000.
The name is derived from Isan. Restoration and discussion of Isanosaurus, by noted paleoartist Luis Rey
Reptiles are tetrapod animals in the class Reptilia, comprising today's turtles, snakes, lizards and their extinct relatives. The study of these traditional reptile orders combined with that of modern amphibians, is called herpetology; because some reptiles are more related to birds than they are to other reptiles, the traditional groups of "reptiles" listed above do not together constitute a monophyletic grouping or clade. For this reason, many modern scientists prefer to consider the birds part of Reptilia as well, thereby making Reptilia a monophyletic class, including all living Diapsids; the earliest known proto-reptiles originated around 312 million years ago during the Carboniferous period, having evolved from advanced reptiliomorph tetrapods that became adapted to life on dry land. Some early examples include Casineria. In addition to the living reptiles, there are many diverse groups that are now extinct, in some cases due to mass extinction events. In particular, the Cretaceous–Paleogene extinction event wiped out the pterosaurs, plesiosaurs and sauropods, as well as many species of theropods, including troodontids, dromaeosaurids and abelisaurids, along with many Crocodyliformes, squamates.
Modern non-avian reptiles inhabit all the continents except Antarctica, although some birds are found on the periphery of Antarctica. Several living subgroups are recognized: Testudines, 350 species. Reptiles are tetrapod vertebrates, creatures that either have four limbs or, like snakes, are descended from four-limbed ancestors. Unlike amphibians, reptiles do not have an aquatic larval stage. Most reptiles are oviparous, although several species of squamates are viviparous, as were some extinct aquatic clades – the fetus develops within the mother, contained in a placenta rather than an eggshell; as amniotes, reptile eggs are surrounded by membranes for protection and transport, which adapt them to reproduction on dry land. Many of the viviparous species feed their fetuses through various forms of placenta analogous to those of mammals, with some providing initial care for their hatchlings. Extant reptiles range in size from a tiny gecko, Sphaerodactylus ariasae, which can grow up to 17 mm to the saltwater crocodile, Crocodylus porosus, which can reach 6 m in length and weigh over 1,000 kg.
In the 13th century the category of reptile was recognized in Europe as consisting of a miscellany of egg-laying creatures, including "snakes, various fantastic monsters, assorted amphibians, worms", as recorded by Vincent of Beauvais in his Mirror of Nature. In the 18th century, the reptiles were, from the outset of classification, grouped with the amphibians. Linnaeus, working from species-poor Sweden, where the common adder and grass snake are found hunting in water, included all reptiles and amphibians in class "III – Amphibia" in his Systema Naturæ; the terms "reptile" and "amphibian" were interchangeable, "reptile" being preferred by the French. Josephus Nicolaus Laurenti was the first to formally use the term "Reptilia" for an expanded selection of reptiles and amphibians similar to that of Linnaeus. Today, the two groups are still treated under the same heading as herptiles, it was not until the beginning of the 19th century that it became clear that reptiles and amphibians are, in fact, quite different animals, Pierre André Latreille erected the class Batracia for the latter, dividing the tetrapods into the four familiar classes of reptiles, amphibians and mammals.
The British anatomist Thomas Henry Huxley made Latreille's definition popular and, together with Richard Owen, expanded Reptilia to include the various fossil "antediluvian monsters", including dinosaurs and the mammal-like Dicynodon he helped describe. This was not the only possible classification scheme: In the Hunterian lectures delivered at the Royal College of Surgeons in 1863, Huxley grouped the vertebrates into mammals and ichthyoids, he subsequently proposed the names of Ichthyopsida for the latter two groups. In 1866, Haeckel demonstrated that vertebrates could be divided based on their reproductive strategies, that reptiles and mammals were united by the amniotic egg; the terms "Sauropsida" and "Theropsida" were used again in 1916 by E. S. Goodrich to distinguish between lizards and their relatives on the one hand and mammals and their extinct relatives on the other. Goodrich supported this division by the nature of the hearts and blood vessels in each group, other features, such as the structure of the forebrain.
According to Goodrich, both lineages evolved from an earlier stem group, Protosauria in which he included some animals today considered reptile-like amphibians, as well as early reptiles. In 1956, D. M. S. Watson observed that the first two groups diverged early in reptilian history, so he divided Goodrich's Protosauria between them, he reinterpreted Sauropsida and Theropsida to exclude birds and mammals, respectively. Thus his Sauropsida included Procolophonia, Millerosauria, Squamata, Rhynchocephalia