The Hylidae are a wide-ranging family of frogs referred to as "tree frogs and their allies". However, the hylids include a diversity of frog species, many of which do not live in trees, but are terrestrial or semiaquatic. Most hylids show adaptations suitable for an arboreal lifestyle, including forward-facing eyes providing binocular vision, adhesive pads on the fingers and toes. In the nonarboreal species, these features may be reduced, or absent; the Cyclorana species are burrowing frogs. Hylids feed on insects and other invertebrates, but some larger species can feed on small vertebrates. Hylids lay their eggs depending on species. Many use ponds, or puddles that collect in the holes of their trees, while others use bromeliads or other water-holding plants. Other species lay their eggs on the leaves of vegetation hanging over water, allowing the tadpoles to drop into the pond when they hatch. A few species use fast-flowing streams, attaching the eggs to the substrate; the tadpoles of these species have suckers enabling them to hold on to rocks.
Another unusual adaptation is found in some South American hylids, which brood the eggs on the back of the female. The tadpoles of most hylid species have laterally placed eyes and broad tails with narrow, filamentous tips; the European tree frog, Hyla arborea, is common in the middle and south of Europe, ranges into Asia and North Africa. The species becomes noisy on the approach of rain and is sometimes kept in confinement as a kind of barometer. North America has many species of the family Hylidae, including the gray tree frog and the American green tree frog; the spring peeper is widespread in the eastern United States and is heard on spring and summer evenings. "Tree frog" is a popular name for several of the Hylidae. H. versicolor is the Gray tree frog, Trachycephalus lichenatus is the lichened tree frog, Trachycephalus marmoratus is the marbled tree frog. However, the name "treefrog" is not unique to this family being used for many species of the Rhacophoridae. Rabbs' fringe-limbed tree frog, Ecnomiohyla rabborum, was a species of Hydilae that went extinct in September 2016.
The last of its kind, a male named Toughie died on September 30, when it was pronounced that the species had become extinct. The family Hylidae is divided into these subfamilies and genera: Hylidae This article incorporates text from the Collier's New Encyclopedia. "Amero-Australian Treefrogs". William E. Duellman. Grzimek's Animal Life Encyclopedia. Ed. Michael Hutchins, Arthur V. Evans, Jerome A. Jackson, Devra G. Kleiman, James B. Murphy, Dennis A. Thoney, et al. Vol. 6: Amphibians. 2nd ed. Detroit: Gale, 2004. P225-243. Amnh.org: Amphibian Species of the World Data related to Hylidae at Wikispecies
The Hylinae are the largest of three subfamilies of Hylidae, the "tree frogs". It contains nearly 700 species in 41 genera, they are found in North and South America, temperate Asia, Africa north of the Sahara. The 41 recognized genera are: http://www.tolweb.org/Hylinae
Amphibians are ectothermic, tetrapod vertebrates of the class Amphibia. Modern amphibians are all Lissamphibia, they inhabit a wide variety of habitats, with most species living within terrestrial, arboreal or freshwater aquatic ecosystems. Thus amphibians start out as larvae living in water, but some species have developed behavioural adaptations to bypass this; the young undergo metamorphosis from larva with gills to an adult air-breathing form with lungs. Amphibians use their skin as a secondary respiratory surface and some small terrestrial salamanders and frogs lack lungs and rely on their skin, they are superficially similar to lizards but, along with mammals and birds, reptiles are amniotes and do not require water bodies in which to breed. With their complex reproductive needs and permeable skins, amphibians are ecological indicators; the earliest amphibians evolved in the Devonian period from sarcopterygian fish with lungs and bony-limbed fins, features that were helpful in adapting to dry land.
They diversified and became dominant during the Carboniferous and Permian periods, but were displaced by reptiles and other vertebrates. Over time, amphibians shrank in size and decreased in diversity, leaving only the modern subclass Lissamphibia; the three modern orders of amphibians are Anura and Apoda. The number of known amphibian species is 8,000, of which nearly 90% are frogs; the smallest amphibian in the world is a frog from New Guinea with a length of just 7.7 mm. The largest living amphibian is the 1.8 m Chinese giant salamander, but this is dwarfed by the extinct 9 m Prionosuchus from the middle Permian of Brazil. The study of amphibians is called batrachology, while the study of both reptiles and amphibians is called herpetology; the word "amphibian" is derived from the Ancient Greek term ἀμφίβιος, which means "both kinds of life", ἀμφί meaning "of both kinds" and βιος meaning "life". The term was used as a general adjective for animals that could live on land or in water, including seals and otters.
Traditionally, the class Amphibia includes all tetrapod vertebrates. Amphibia in its widest sense was divided into three subclasses, two of which are extinct: Subclass Lepospondyli† Subclass Temnospondyli† Subclass Lissamphibia Salientia: Jurassic to present—6,200 current species in 53 families Caudata: Jurassic to present—652 current species in 9 families Gymnophiona: Jurassic to present—192 current species in 10 families The actual number of species in each group depends on the taxonomic classification followed; the two most common systems are the classification adopted by the website AmphibiaWeb, University of California and the classification by herpetologist Darrel Frost and the American Museum of Natural History, available as the online reference database "Amphibian Species of the World". The numbers of species cited above follows Frost and the total number of known amphibian species as of March 31, 2019 is 8,000, of which nearly 90% are frogs. With the phylogenetic classification, the taxon Labyrinthodontia has been discarded as it is a polyparaphyletic group without unique defining features apart from shared primitive characteristics.
Classification varies according to the preferred phylogeny of the author and whether they use a stem-based or a node-based classification. Traditionally, amphibians as a class are defined as all tetrapods with a larval stage, while the group that includes the common ancestors of all living amphibians and all their descendants is called Lissamphibia; the phylogeny of Paleozoic amphibians is uncertain, Lissamphibia may fall within extinct groups, like the Temnospondyli or the Lepospondyli, in some analyses in the amniotes. This means that advocates of phylogenetic nomenclature have removed a large number of basal Devonian and Carboniferous amphibian-type tetrapod groups that were placed in Amphibia in Linnaean taxonomy, included them elsewhere under cladistic taxonomy. If the common ancestor of amphibians and amniotes is included in Amphibia, it becomes a paraphyletic group. All modern amphibians are included in the subclass Lissamphibia, considered a clade, a group of species that have evolved from a common ancestor.
The three modern orders are Anura and Gymnophiona. It has been suggested that salamanders arose separately from a Temnospondyl-like ancestor, that caecilians are the sister group of the advanced reptiliomorph amphibians, thus of amniotes. Although the fossils of several older proto-frogs with primitive characteristics are known, the oldest "true frog" is Prosalirus bitis, from the Early Jurassic Kayenta Formation of Arizona, it is anatomically similar to modern frogs. The oldest known caecilian is another Early Jurassic species, Eocaecilia micropodia from Arizona; the earliest salamander is Beiyanerpeton jianpingensis from the Late Jurassic of northeastern China. Authorities disagree as to whether Salientia is a superorder that includes the order Anura, or whether
A chordate is an animal constituting the phylum Chordata. During some period of their life cycle, chordates possess a notochord, a dorsal nerve cord, pharyngeal slits, an endostyle, a post-anal tail: these five anatomical features define this phylum. Chordates are bilaterally symmetric; the Chordata and Ambulacraria together form the superphylum Deuterostomia. Chordates are divided into three subphyla: Vertebrata. There are extinct taxa such as the Vetulicolia. Hemichordata has been presented as a fourth chordate subphylum, but now is treated as a separate phylum: hemichordates and Echinodermata form the Ambulacraria, the sister phylum of the Chordates. Of the more than 65,000 living species of chordates, about half are bony fish that are members of the superclass Osteichthyes. Chordate fossils have been found from as early as the Cambrian explosion, 541 million years ago. Cladistically, vertebrates - chordates with the notochord replaced by a vertebral column during development - are considered to be a subgroup of the clade Craniata, which consists of chordates with a skull.
The Craniata and Tunicata compose the clade Olfactores. Chordates form a phylum of animals that are defined by having at some stage in their lives all of the following anatomical features: A notochord, a stiff rod of cartilage that extends along the inside of the body. Among the vertebrate sub-group of chordates the notochord develops into the spine, in wholly aquatic species this helps the animal to swim by flexing its tail. A dorsal neural tube. In fish and other vertebrates, this develops into the spinal cord, the main communications trunk of the nervous system. Pharyngeal slits; the pharynx is the part of the throat behind the mouth. In fish, the slits are modified to form gills, but in some other chordates they are part of a filter-feeding system that extracts particles of food from the water in which the animals live. Post-anal tail. A muscular tail that extends backwards behind the anus. An endostyle; this is a groove in the ventral wall of the pharynx. In filter-feeding species it produces mucus to gather food particles, which helps in transporting food to the esophagus.
It stores iodine, may be a precursor of the vertebrate thyroid gland. There are soft constraints that separate chordates from certain other biological lineages, but are not part of the formal definition: All chordates are deuterostomes; this means. All chordates are based on a bilateral body plan. All chordates are coelomates, have a fluid filled body cavity called a coelom with a complete lining called peritoneum derived from mesoderm; the following schema is from the third edition of Vertebrate Palaeontology. The invertebrate chordate classes are from Fishes of the World. While it is structured so as to reflect evolutionary relationships, it retains the traditional ranks used in Linnaean taxonomy. Phylum Chordata †Vetulicolia? Subphylum Cephalochordata – Class Leptocardii Clade Olfactores Subphylum Tunicata – Class Ascidiacea Class Thaliacea Class Appendicularia Class Sorberacea Subphylum Vertebrata Infraphylum incertae sedis Cyclostomata Superclass'Agnatha' paraphyletic Class Myxini Class Petromyzontida or Hyperoartia Class †Conodonta Class †Myllokunmingiida Class †Pteraspidomorphi Class †Thelodonti Class †Anaspida Class †Cephalaspidomorphi Infraphylum Gnathostomata Class †Placodermi Class Chondrichthyes Class †Acanthodii Superclass Osteichthyes Class Actinopterygii Class Sarcopterygii Superclass Tetrapoda Class Amphibia Class Sauropsida Class Synapsida Craniates, one of the three subdivisions of chordates, all have distinct skulls.
They include the hagfish. Michael J. Benton commented that "craniates are characterized by their heads, just as chordates, or all deuterostomes, are by their tails". Most craniates are vertebrates; these consist of a series of bony or cartilaginous cylindrical vertebrae with neural arches that protect the spinal cord, with projections that link the vertebrae. However hagfish have incomplete braincases and no vertebrae, are therefore not regarded as vertebrates, but as members of the craniates, the group from which vertebrates are thought to have evolved; however the cladistic exclusion of hagfish from the vertebrates is controversial, as they ma
A frog is any member of a diverse and carnivorous group of short-bodied, tailless amphibians composing the order Anura. The oldest fossil "proto-frog" appeared in the early Triassic of Madagascar, but molecular clock dating suggests their origins may extend further back to the Permian, 265 million years ago. Frogs are distributed, ranging from the tropics to subarctic regions, but the greatest concentration of species diversity is in tropical rainforests. There are accounting for over 85 % of extant amphibian species, they are one of the five most diverse vertebrate orders. Warty frog species tend to be called toads, but the distinction between frogs and toads is informal, not from taxonomy or evolutionary history. An adult frog has a stout body, protruding eyes, anteriorly-attached tongue, limbs folded underneath, no tail. Frogs have glandular skin, with secretions ranging from distasteful to toxic, their skin varies in colour from well-camouflaged dappled brown and green to vivid patterns of bright red or yellow and black to show toxicity and ward off predators.
Adult frogs live on dry land. Frogs lay their eggs in water; the eggs hatch into aquatic larvae called tadpoles that have internal gills. They have specialized rasping mouth parts suitable for herbivorous, omnivorous or planktivorous diets; the life cycle is completed. A few species bypass the tadpole stage. Adult frogs have a carnivorous diet consisting of small invertebrates, but omnivorous species exist and a few feed on fruit. Frog skin has a rich microbiome, important to their health. Frogs are efficient at converting what they eat into body mass, they are an important food source for predators and part of the food web dynamics of many of the world's ecosystems. The skin is semi-permeable, making them susceptible to dehydration, so they either live in moist places or have special adaptations to deal with dry habitats. Frogs produce a wide range of vocalizations in their breeding season, exhibit many different kinds of complex behaviours to attract mates, to fend off predators and to survive.
Frogs are valued as food by humans and have many cultural roles in literature and religion. Frog populations have declined since the 1950s. More than one third of species are considered to be threatened with extinction and over 120 are believed to have become extinct since the 1980s; the number of malformations among frogs is on the rise and an emerging fungal disease, has spread around the world. Conservation biologists are working to resolve them; the use of the common names "frog" and "toad" has no taxonomic justification. From a classification perspective, all members of the order Anura are frogs, but only members of the family Bufonidae are considered "true toads"; the use of the term "frog" in common names refers to species that are aquatic or semi-aquatic and have smooth, moist skins. There are numerous exceptions to this rule; the European fire-bellied toad has a warty skin and prefers a watery habitat whereas the Panamanian golden frog is in the toad family Bufonidae and has a smooth skin.
The origin of the order name Anura — and its original spelling Anoures — is the Ancient Greek "alpha privative" prefix ἀν- "without", οὐρά, meaning "animal tail". It refers to the tailless character of these amphibians; the origins of the word frog are debated. The word is first attested in Old English as frogga, but the usual Old English word for the frog was frosc, it is agreed that the word frog is somehow related to this. Old English frosc remained in dialectal use in English as frosh and frosk into the nineteenth century, is paralleled in other Germanic languages, with examples in the modern languages including German Frosch, Icelandic froskur, Dutch vors; these words allow us to reconstruct a Common Germanic ancestor *froskaz. The third edition of the Oxford English Dictionary finds that the etymology of *froskaz is uncertain, but agrees with arguments that it could plausibly derive from a Proto-Indo-European base along the lines of *preu = "jump". How Old English frosc gave rise to frogga is, uncertain, as the development does not involve a regular sound-change.
Instead, it seems that there was a trend in Old English to coin nicknames for animals ending in -g, with examples—themselves all of uncertain etymology—including dog, pig and wig. Frog appears to have been adapted from frosc as part of this trend. Meanwhile, the word toad, first attested as Old English tādige, is unique to English and is of uncertain etymology, it is the basis for the word tadpole, first attested as Middle English taddepol meaning'toad-head'. About 88% of amphibian species are classified in the order Anura; these include over 7,000 species in 56 families, of which the Craugastoridae, Hylidae and Bufonidae are the richest in species. The Anura include any fossil species that fit within the anuran definition; the characteristics of anuran adults include: 9 or fewer presacral vertebrae, the presence of a urostyle formed of fused vertebrae, no tail, a long and forward-sloping ilium, shorter fore limbs than hind limbs and ulna fused and fibula fused, elongated an
Edward Drinker Cope
Edward Drinker Cope was an American paleontologist and comparative anatomist, as well as a noted herpetologist and ichthyologist. He was a founder of the Neo-Lamarckism school of thought. Born to a wealthy Quaker family, Cope distinguished himself as a child prodigy interested in science. Though his father tried to raise Cope as a gentleman farmer, he acquiesced to his son's scientific aspirations. Cope had one child. Cope had little formal scientific training, he eschewed a teaching position for field work, he made regular trips to the American West, prospecting in the 1870s and 1880s as a member of United States Geological Survey teams. A personal feud between Cope and paleontologist Othniel Charles Marsh led to a period of intense fossil-finding competition now known as the Bone Wars. Cope's financial fortunes soured after failed mining ventures in the 1880s, forcing him to sell off much of his fossil collection, he experienced a resurgence in his career toward the end of his life before dying on April 12, 1897.
Though Cope's scientific pursuits nearly bankrupted him, his contributions helped to define the field of American paleontology. He was a prodigious writer, with 1,400 papers published over his lifetime, although his rivals debated the accuracy of his published works, he discovered and named more than 1,000 vertebrate species, including hundreds of fishes and dozens of dinosaurs. His proposal for the origin of mammalian molars is notable among his theoretical contributions. "Cope's rule", the hypothesis that mammalian lineages grow larger over geologic time, while named after him, is "neither explicit nor implicit" in his work. Edward Drinker Cope was born on the eldest son of Alfred and Hanna Cope; the death of his mother when he was three years old seemed to have had little effect on young Edward, as he mentioned in his letters that he had no recollection of her. His stepmother, Rebecca Biddle, filled the maternal role. Alfred, an orthodox member of the Religious Society of Friends or Quakers, operated a lucrative shipping business started by his father, Thomas P. Cope, in 1821.
He was a philanthropist who gave money to the Society of Friends, the Philadelphia Zoological Gardens, the Institute for Colored Youth. Edward was born and raised in a large stone house called "Fairfield", whose location is now within the boundaries of Philadelphia; the 8 acres of pristine and exotic gardens of the house offered a landscape that Edward was able to explore. The Copes began teaching their children to read and write at a young age, took Edward on trips across New England and to museums and gardens. Cope's interest in animals became apparent at a young age. Alfred intended to give his son the same education he himself had received. At age nine, Edward was sent to a day school in Philadelphia and in 1853 at the age of 12, Edward was sent to the Friends' Boarding School at Westtown, near West Chester, Pennsylvania; the school was founded in 1799 with fundraising by members of the Society of Friends, provided much of the Cope family's education. The prestigious school was expensive, costing Alfred $500 in tuition each year, in his first year, Edward studied algebra, scripture, grammar and Latin.
Edward's letters home requesting a larger allowance show he was able to manipulate his father, he was, according to author and Cope biographer Jane Davidson, "a bit of a spoiled brat". His letters suggest he was lonely at the school—it was the first time he had been away from his home for an extended period. Otherwise, Edward's studies progressed much like a typical boy—he had "less than perfect" or "not quite satisfactory" marks for conduct from his teachers, did not work hard on his penmanship lessons, which may have contributed to his illegible handwriting as an adult. Edward returned to Westtown in 1855. Biology began to interest him more that year, he studied natural history texts in his spare time. While at the school, he visited the Academy of Natural Sciences. Edward obtained bad marks due to quarreling and bad conduct, his letters to his father show he chafed at farm work and betrayed flashes of the temper for which he would become well known. After sending Edward back to the farm for summer break in 1854 and 1855, Alfred did not return Edward to school after spring 1856.
Instead, Alfred attempted to turn his son into a gentleman farmer, which he considered a wholesome profession that would yield enough profit to lead a comfortable life, improve the undersized Edward's health. Until 1863, Cope's letters to his father continually expressed his yearning for a more professional scientific career than that of a farmer, which he called "dreadfully boring". While working on farms, Edward continued his education on his own. In 1858, he began working part-time at the Academy of Natural Sciences and cataloguing specimens, published his first series of research results in January 1859. Cope began taking German classes with a former Westtown teacher. Though Alfred resisted his son's pursuit of a science career, he paid for his son's private studies. Instead of working the farm his father bought for him, Edward rented out the land and used the income to further his scientific endeavors. Alfred gave in to Edward's wishes and paid for university cl