|Masdevallia teaguei Luer|
|Masdevallia teaguei Luer|
Monocotyledons referred to as monocots, are flowering plants whose seeds contain only one embryonic leaf, or cotyledon. They constitute one of the major groups into which the flowering plants have traditionally been divided, the rest of the flowering plants having two cotyledons and therefore classified as dicotyledons, or dicots. However, molecular phylogenetic research has shown that while the monocots form a monophyletic group or clade, the dicots do not. Monocots have always been recognized as a group, but with various taxonomic ranks and under several different names; the APG III system of 2009 recognises a clade called "monocots" but does not assign it to a taxonomic rank. The monocots include about 60,000 species; the largest family in this group by number of species are the orchids, with more than 20,000 species. About half as many species belong to the true grasses, which are economically the most important family of monocots. In agriculture the majority of the biomass produced; these include not only major grains, but forage grasses, sugar cane, the bamboos.
Other economically important monocot crops include various palms and plantains, gingers and their relatives and cardamom, pineapple, water chestnut, leeks and garlic. Many houseplants are monocot epiphytes. Additionally most of the horticultural bulbs, plants cultivated for their blooms, such as lilies, irises, cannas and tulips, are monocots; the monocots or monocotyledons have, as the name implies, a single cotyledon, or embryonic leaf, in their seeds. This feature was used to contrast the monocots with the dicotyledons or dicots which have two cotyledons. From a diagnostic point of view the number of cotyledons is neither a useful characteristic, nor is it reliable; the single cotyledon is only one of a number of modifications of the body plan of the ancestral monocotyledons, whose adaptive advantages are poorly understood, but may have been related to adaption to aquatic habitats, prior to radiation to terrestrial habitats. Monocots are sufficiently distinctive that there has been disagreement as to membership of this group, despite considerable diversity in terms of external morphology.
However, morphological features that reliably characterise major clades are rare. Thus monocots are distinguishable from other angiosperms both in terms of their uniformity and diversity. On the one hand the organisation of the shoots, leaf structure and floral configuration are more uniform than in the remaining angiosperms, yet within these constraints a wealth of diversity exists, indicating a high degree of evolutionary success. Monocot diversity includes perennial geophytes such as ornamental flowers including and succulent epiphytes, all in the lilioid monocots, major cereal grains in the grass family and forage grasses as well as woody tree-like palm trees, bamboo and bromeliads, bananas and ginger in the commelinid monocots, as well as both emergent and aroids, as well as floating or submerged aquatic plants such as seagrass. Organisation and life formsThe most important distinction is their growth pattern, lacking a lateral meristem that allows for continual growth in diameter with height, therefore this characteristic is a basic limitation in shoot construction.
Although herbaceous, some arboraceous monocots reach great height and mass. The latter include agaves, palms and bamboos; this creates challenges in water transport. Some, such as species of Yucca, develop anomalous secondary growth, while palm trees utilise an anomalous primary growth form described as establishment growth; the axis undergoes primary thickening, that progresses from internode to internode, resulting in a typical inverted conical shape of the basal primary axis. The limited conductivity contributes to limited branching of the stems. Despite these limitations a wide variety of adaptive growth forms has resulted from epiphytic orchids and bromeliads to submarine Alismatales and mycotrophic Burmanniaceae and Triuridaceae. Other forms of adaptation include the climbing vines of Araceae which use negative phototropism to locate host trees, while some palms such as Calamus manan produce the longest shoots in the plant kingdom, up to 185 m long. Other monocots Poales, have adopted a therophyte life form.
LeavesThe cotyledon, the primordial Angiosperm leaf consists of a proximal leaf base or hypophyll and a distal hyperphyll. In monocots the hypophyll tends to be the dominant part in contrast to other angiosperms. From these, considerable diversity arises. Mature monocot leaves are narrow and linear, forming a sheath
The Pleurothallidinae are a neotropical subtribe of plants of the orchid family including 29 genera in more than 4000 species. Occurring species of this subtribe are among the more popular orchids of horticulturalists the genera Dracula, Dryadella and Restrepia; the following genera are considered monophyletic: Barbosella, Dresslerella, Lepanthes, Platystele, Restrepia, Scaphosepalum and Zootrophion. Many genera in the Pleurothallidinae were found polyphyletic, for example species attributed to the genus Pleurothallis are scattered across five clades. Acianthera – Anathallis – Andinia – Barbosella – Brachionidium – Chamelophyton – Dilomilis – Diodonopsis – Draconanthes – Dracula – Dresslerella – Dryadella – Echinosepala – Frondaria – Kraenzlinella – Lepanthes – Lepanthopsis – Masdevallia – Myoxanthus – Neocogniauxia – Octomeria – Pabstiella – Phloeophila – Platystele – Pleurothallis – Pleurothallopsis – Porroglossum – Restrepia – Restrepiella – Scaphosepalum – Specklinia – Stelis – Teagueia – Tomzanonia – Trichosalpinx – Trisetella – Zootrophion DNA-based reclassification of the Pleurothallidinae Alec M. Pridgeon, Rodolfo Solano and Mark W. Chase - Phylogenetic relationships in Pleurothallidinae: combined evidence from nuclear and plastid DNA sequences.
Plants are multicellular, predominantly photosynthetic eukaryotes of the kingdom Plantae. Plants were treated as one of two kingdoms including all living things that were not animals, all algae and fungi were treated as plants. However, all current definitions of Plantae exclude the fungi and some algae, as well as the prokaryotes. By one definition, plants form the clade Viridiplantae, a group that includes the flowering plants and other gymnosperms and their allies, liverworts and the green algae, but excludes the red and brown algae. Green plants obtain most of their energy from sunlight via photosynthesis by primary chloroplasts that are derived from endosymbiosis with cyanobacteria, their chloroplasts contain b, which gives them their green color. Some plants are parasitic or mycotrophic and have lost the ability to produce normal amounts of chlorophyll or to photosynthesize. Plants are characterized by sexual reproduction and alternation of generations, although asexual reproduction is common.
There are about 320 thousand species of plants, of which the great majority, some 260–290 thousand, are seed plants. Green plants provide a substantial proportion of the world's molecular oxygen and are the basis of most of Earth's ecosystems on land. Plants that produce grain and vegetables form humankind's basic foods, have been domesticated for millennia. Plants have many cultural and other uses, as ornaments, building materials, writing material and, in great variety, they have been the source of medicines and psychoactive drugs; the scientific study of plants is known as a branch of biology. All living things were traditionally placed into one of two groups and animals; this classification may date from Aristotle, who made the distincton between plants, which do not move, animals, which are mobile to catch their food. Much when Linnaeus created the basis of the modern system of scientific classification, these two groups became the kingdoms Vegetabilia and Animalia. Since it has become clear that the plant kingdom as defined included several unrelated groups, the fungi and several groups of algae were removed to new kingdoms.
However, these organisms are still considered plants in popular contexts. The term "plant" implies the possession of the following traits multicellularity, possession of cell walls containing cellulose and the ability to carry out photosynthesis with primary chloroplasts; when the name Plantae or plant is applied to a specific group of organisms or taxon, it refers to one of four concepts. From least to most inclusive, these four groupings are: Another way of looking at the relationships between the different groups that have been called "plants" is through a cladogram, which shows their evolutionary relationships; these are not yet settled, but one accepted relationship between the three groups described above is shown below. Those which have been called "plants" are in bold; the way in which the groups of green algae are combined and named varies between authors. Algae comprise several different groups of organisms which produce food by photosynthesis and thus have traditionally been included in the plant kingdom.
The seaweeds range from large multicellular algae to single-celled organisms and are classified into three groups, the green algae, red algae and brown algae. There is good evidence that the brown algae evolved independently from the others, from non-photosynthetic ancestors that formed endosymbiotic relationships with red algae rather than from cyanobacteria, they are no longer classified as plants as defined here; the Viridiplantae, the green plants – green algae and land plants – form a clade, a group consisting of all the descendants of a common ancestor. With a few exceptions, the green plants have the following features in common, they undergo closed mitosis without centrioles, have mitochondria with flat cristae. The chloroplasts of green plants are surrounded by two membranes, suggesting they originated directly from endosymbiotic cyanobacteria. Two additional groups, the Rhodophyta and Glaucophyta have primary chloroplasts that appear to be derived directly from endosymbiotic cyanobacteria, although they differ from Viridiplantae in the pigments which are used in photosynthesis and so are different in colour.
These groups differ from green plants in that the storage polysaccharide is floridean starch and is stored in the cytoplasm rather than in the plastids. They appear to have had a common origin with Viridiplantae and the three groups form the clade Archaeplastida, whose name implies that their chloroplasts were derived from a single ancient endosymbiotic event; this is the broadest modern definition of the term'plant'. In contrast, most other algae not only have different pigments but have chloroplasts with three or four surrounding membranes, they are not close relatives of the Archaeplastida having acquired chloroplasts separately from ingested or symbiotic green and red algae. They are thus not included in the broadest modern definition of the plant kingdom, although they were in the past; the green plants or Viridiplantae were traditionally divided into the green algae (including
Colombia the Republic of Colombia, is a sovereign state situated in the northwest of South America, with territories in Central America. Colombia shares a border to the northwest with Panama, to the east with Venezuela and Brazil and to the south with Ecuador and Peru, it shares its maritime limits with Costa Rica, Honduras, Jamaica and the Dominican Republic. Colombia is a unitary, constitutional republic comprising thirty-two departments, with the capital in Bogota. Colombia has been inhabited by various indigenous peoples since 12,000 BCE, including the Muisca and the Tairona, along with the Inca Empire that expanded to the southwest of the country; the Spanish arrived in 1499 and by the mid-16th century conquered and colonized much of the region, establishing the New Kingdom of Granada, with Santafé de Bogotá as its capital. Independence from Spain was achieved in 1819, but by 1830 the "Gran Colombia" Federation was dissolved, with what is now Colombia and Panama emerging as the Republic of New Granada.
The new nation experimented with federalism as the Granadine Confederation, the United States of Colombia, before the Republic of Colombia was declared in 1886. Panama seceded in 1903. Beginning in the 1960s, the country suffered from an asymmetric low-intensity armed conflict and rampant political violence, both of which escalated in the 1990s. Since 2005, there has been significant improvement in security and rule of law. Colombia is one of the most ethnically and linguistically diverse countries in the world, with its rich cultural heritage reflecting influences by indigenous peoples, European settlement, forced African migration, immigration from Europe and the Middle East. Urban centres are located in the highlands of the Andes mountains and the Caribbean coast. Colombia is among the world's 17 megadiverse countries, the most densely biodiverse per square kilometer. Colombia is a middle power and regional actor in Latin America, it is part of the CIVETS group of six leading emerging markets and a member of the UN, the WTO, the OAS, the Pacific Alliance, other international organizations.
Colombia's diversified economy is the fourth largest in Latin America, with macroeconomic stability and favorable long-term growth prospects. The name "Colombia" is derived from the last name of Christopher Columbus, it was conceived by the Venezuelan revolutionary Francisco de Miranda as a reference to all the New World, but to those portions under Spanish rule. The name was adopted by the Republic of Colombia of 1819, formed from the territories of the old Viceroyalty of New Granada; when Venezuela and Cundinamarca came to exist as independent states, the former Department of Cundinamarca adopted the name "Republic of New Granada". New Granada changed its name in 1858 to the Granadine Confederation. In 1863 the name was again changed, this time to United States of Colombia, before adopting its present name – the Republic of Colombia – in 1886. To refer to this country, the Colombian government uses the terms Colombia and República de Colombia. Owing to its location, the present territory of Colombia was a corridor of early human migration from Mesoamerica and the Caribbean to the Andes and Amazon basin.
The oldest archaeological finds are from the Pubenza and El Totumo sites in the Magdalena Valley 100 kilometres southwest of Bogotá. These sites date from the Paleoindian period. At Puerto Hormiga and other sites, traces from the Archaic Period have been found. Vestiges indicate that there was early occupation in the regions of El Abra and Tequendama in Cundinamarca; the oldest pottery discovered in the Americas, found at San Jacinto, dates to 5000–4000 BCE. Indigenous people inhabited the territory, now Colombia by 12,500 BCE. Nomadic hunter-gatherer tribes at the El Abra, Tibitó and Tequendama sites near present-day Bogotá traded with one another and with other cultures from the Magdalena River Valley. Between 5000 and 1000 BCE, hunter-gatherer tribes transitioned to agrarian societies. Beginning in the 1st millennium BCE, groups of Amerindians including the Muisca, Zenú, Tairona developed the political system of cacicazgos with a pyramidal structure of power headed by caciques; the Muisca inhabited the area of what is now the Departments of Boyacá and Cundinamarca high plateau where they formed the Muisca Confederation.
They farmed maize, potato and cotton, traded gold, blankets, ceramic handicrafts and rock salt with neighboring nations. The Tairona inhabited northern Colombia in the isolated mountain range of Sierra Nevada de Santa Marta; the Quimbaya inhabited regions of the Cauca River Valley between the Western and Central Ranges of the Colombian Andes. Most of the Amerindians practiced agriculture and the social structure of each indigenous community was different; some groups of indigenous people such as the Caribs lived in a state of permanent war, but others had less bellicose attitudes. The Incas expanded their empire onto the southwest part of the country. Alonso de Ojeda reached the Guajira Peninsula in 1499. Spanish explorers, led by Rodrigo de Bastidas, made the first exploration
The flowering plants known as angiosperms, Angiospermae or Magnoliophyta, are the most diverse group of land plants, with 64 orders, 416 families 13,164 known genera and c. 369,000 known species. Like gymnosperms, angiosperms are seed-producing plants. However, they are distinguished from gymnosperms by characteristics including flowers, endosperm within the seeds, the production of fruits that contain the seeds. Etymologically, angiosperm means a plant; the term comes from the Greek words sperma. The ancestors of flowering plants diverged from gymnosperms in the Triassic Period, 245 to 202 million years ago, the first flowering plants are known from 160 mya, they diversified extensively during the Early Cretaceous, became widespread by 120 mya, replaced conifers as the dominant trees from 100 to 60 mya. Angiosperms differ from other seed plants in several ways, described in the table below; these distinguishing characteristics taken together have made the angiosperms the most diverse and numerous land plants and the most commercially important group to humans.
Angiosperm stems are made up of seven layers. The amount and complexity of tissue-formation in flowering plants exceeds that of gymnosperms; the vascular bundles of the stem are arranged such that the phloem form concentric rings. In the dicotyledons, the bundles in the young stem are arranged in an open ring, separating a central pith from an outer cortex. In each bundle, separating the xylem and phloem, is a layer of meristem or active formative tissue known as cambium. By the formation of a layer of cambium between the bundles, a complete ring is formed, a regular periodical increase in thickness results from the development of xylem on the inside and phloem on the outside; the soft phloem becomes crushed, but the hard wood persists and forms the bulk of the stem and branches of the woody perennial. Owing to differences in the character of the elements produced at the beginning and end of the season, the wood is marked out in transverse section into concentric rings, one for each season of growth, called annual rings.
Among the monocotyledons, the bundles are more numerous in the young stem and are scattered through the ground tissue. They once formed the stem increases in diameter only in exceptional cases; the characteristic feature of angiosperms is the flower. Flowers show remarkable variation in form and elaboration, provide the most trustworthy external characteristics for establishing relationships among angiosperm species; the function of the flower is to ensure fertilization of the ovule and development of fruit containing seeds. The floral apparatus may arise terminally from the axil of a leaf; as in violets, a flower arises singly in the axil of an ordinary foliage-leaf. More the flower-bearing portion of the plant is distinguished from the foliage-bearing or vegetative portion, forms a more or less elaborate branch-system called an inflorescence. There are two kinds of reproductive cells produced by flowers. Microspores, which will divide to become pollen grains, are the "male" cells and are borne in the stamens.
The "female" cells called megaspores, which will divide to become the egg cell, are contained in the ovule and enclosed in the carpel. The flower may consist only of these parts, as in willow, where each flower comprises only a few stamens or two carpels. Other structures are present and serve to protect the sporophylls and to form an envelope attractive to pollinators; the individual members of these surrounding structures are known as petals. The outer series is green and leaf-like, functions to protect the rest of the flower the bud; the inner series is, in general, white or brightly colored, is more delicate in structure. It functions to attract bird pollinators. Attraction is effected by color and nectar, which may be secreted in some part of the flower; the characteristics that attract pollinators account for the popularity of flowers and flowering plants among humans. While the majority of flowers are perfect or hermaphrodite, flowering plants have developed numerous morphological and physiological mechanisms to reduce or prevent self-fertilization.
Heteromorphic flowers have short carpels and long stamens, or vice versa, so animal pollinators cannot transfer pollen to the pistil. Homomorphic flowers may employ a biochemical mechanism called self-incompatibility to discriminate between self and non-self pollen grains. In other species, the male and female parts are morphologically separated, developing on different flowers; the botanical term "Angiosperm", from the Ancient Greek αγγείον, angeíon and σπέρμα, was coined in the form Angiospermae by Paul Hermann in 1690, as the name of one of his primary divisions of the plant kingdom. This included flowering plants possessing seeds enclosed in capsules, distinguished from his Gymnospermae, or flowering plants with achenial or schizo-carpic fruits, the whole fruit or each of its pieces being here regarded as a seed and naked; the term and its antonym were maintained by Carl Linnaeus with the same sense, but with restricted application, in the names of the orders of his class Didynamia. Its use with any
Asparagales is an order of plants in modern classification systems such as the Angiosperm Phylogeny Group and the Angiosperm Phylogeny Web. The order takes its name from the type family Asparagaceae and is placed in the monocots amongst the lilioid monocots; the order has only been recognized in classification systems. It was first put forward by Huber in 1977 and taken up in the Dahlgren system of 1985 and the APG in 1998, 2003 and 2009. Before this, many of its families were assigned to the old order Liliales, a large order containing all monocots with colourful tepals and lacking starch in their endosperm. DNA sequence analysis indicated that many of the taxa included in Liliales should be redistributed over three orders, Liliales and Dioscoreales; the boundaries of the Asparagales and of its families have undergone a series of changes in recent years. In the APG circumscription, Asparagales is the largest order of monocots with 14 families, 1,122 genera, about 36,000 species; the order is circumscribed on the basis of molecular phylogenetics, but is difficult to define morphologically, since its members are structurally diverse.
Most species of Asparagales are herbaceous perennials, although some are climbers and some are tree-like. The order contains many geophytes. According to telomere sequence, at least two evolutionary switch-points happened within the order. Basal sequence is formed by TTTAGGG like in majority of higher plants. Basal motif was changed to vertebrate-like TTAGGG and the most divergent motif CTCGGTTATGGG appears in Allium. One of the defining characteristics of the order is the presence of phytomelanin, a black pigment present in the seed coat, creating a dark crust. Phytomelanin is found in most families of the Asparagales; the leaves of all species form a tight rosette, either at the base of the plant or at the end of the stem, but along the stem. The flowers are not distinctive, being'lily type', with six tepals and up to six stamina; the order is thought to have first diverged from other related monocots some 120–130 million years ago, although given the difficulty in classifying the families involved, estimates are to be uncertain.
From an economic point of view, the order Asparagales is second in importance within the monocots to the order Poales. Species are used as food and flavourings, as cut flowers, as garden ornamentals. Although most species in the order are herbaceous, some no more than 15 cm high, there are a number of climbers, as well as several genera forming trees, which can exceed 10 m in height. Succulent genera occur in several families. All species have a tight cluster of leaves, either at the base of the plant or at the end of a more-or-less woody stem as with Yucca. In some cases the leaves are produced along the stem; the flowers are in the main not distinctive, being of a general'lily type', with six tepals, either free or fused from the base and up to six stamina. They are clustered at the end of the plant stem; the Asparagales are distinguished from the Liliales by the lack of markings on the tepals, the presence of septal nectaries in the ovaries, rather than the bases of the tepals or stamen filaments, the presence of secondary growth.
They are geophytes, but with linear leaves, a lack of fine reticular venation. The seeds characteristically have the external epidermis either obliterated, or if present, have a layer of black carbonaceous phytomelanin in species with dry fruits; the inner part of the seed coat is collapsed, in contrast to Liliales whose seeds have a well developed outer epidermis, lack phytomelanin, display a cellular inner layer. The orders which have been separated from the old Liliales are difficult to characterize. No single morphological character appears to be diagnostic of the order Asparagales; the flowers of Asparagales are of a general type among the lilioid monocots. Compared to Liliales, they have plain tepals without markings in the form of dots. If nectaries are present, they are in the septa of the ovaries rather than at the base of the tepals or stamens; those species which have large dry seeds have a dark, crust-like outer layer containing the pigment phytomelan. However, some species with hairy seeds, berries, or reduced seeds lack this dark pigment in their seed coats.
Phytomelan is not unique to Asparagales but it is common within the order and rare outside it. The inner portion of the seed coat is completely collapsed. In contrast, the morphologically similar seeds of Liliales have no phytomelan, retain a cellular structure in the inner portion of the seed coat. Most monocots are unable to thicken their stems once they have formed, since they lack the cylindrical meristem present in other angiosperm groups. Asparagales have a method of secondary thickening, otherwise only found inDioscorea. In a process called'anomalous secondary growth', they are able to create new
Masdevallia, abbreviated Masd in horticultural trade, is a large genus of flowering plants of the Pleurothallidinae, a subtribe of the orchid family. There are over 500 species, grouped into several subgenera; the genus is named for a physician and botanist in the court of Charles III of Spain. These plants are found from Mexico to southern Brazil, but in the higher regions of the Andes of Ecuador and Colombia and Bolivia, they may be terrestrials or growing as lithophytes on damp rocks. The plants are characterized by an abbreviated to elongate and creeping rhizome that gives rise to stems that lack pseudobulbs; the stem bears a single, erect to pendent, ovate to lanceolate leaf. The flowers occur singly or in racemose inflorescences, they are characterized by reduced corolla. The sepals are fused at the base and caudate; the petals flank the semiterete column and the tongue-shaped lip is flexibly hinged to a free column foot. The species are sensitive to inappropriate cultural conditions and will show signs of stress by leaf spotting or dropping.
They can be grown in pots with sphagnum or seedling grade wood chips although a few species produce descending inflorescences and are best accommodated in baskets. In both cases the rhizome should remain at the surface of the medium. Most of these plants are from high altitude cloud forests and require cool conditions and abundant moisture throughout the year, they cannot tolerate dryness, low humidity, or excessive temperatures and the plants are easy to kill. They will drop all their leaves and collapse if allowed to dry or are exposed to high temperatures. Many members of this genus from high altitude cloud forests defy cultivation. Most of the species from this genus are considered less difficult in cultivation than plants from the genus Dracula, some of them are easy to cultivate and have a'weedy' habit such as Veitch's masdevallia, but the majority of these species are very difficult to maintain in cultivation unless the plants can be kept cool and moist all the time. Low humidity conditions or watering the plants with a water source which contains high levels of dissolved salts will result in the leaves yellowing and dying from the tips back to the rhizome.
The plants should be provided with rain water or distilled water or a pure water source. The medium should always remain moist as the plants do not have any significant storage structures like most orchids. Masdevallia antonii Masdevallia davisii – Davis' masdevallia, orchid of the sun, qoriwaqanki Masdevallia pinocchio Masdevallia unguentum Masdevallia veitchiana – Veitch's masdevallia, king of the masdevallias, gallo-gallo, waqanki Masdevallia was found to be polyphyletic and the taxonomy of the group is unresolved but the accepted taxonomy proceeds as follows: Subgenus Amanda Section Amandae: 28+ species, e.g. M. amanda, M. bulbophyllopsis, M. melanopus, M. polysticta. Separated in Spilotantha. Section Ophioglossae: 1-2 species – M. ophioglossa, M. ophioglossa ssp. grossa Subgenus Cucullatia: 4 species – M. cerastes, M. corniculata, M. cucullata, M. macrura Subgenus Fissia: 3 species – M. mutica, M. picturata, M. pleurothalloides Subgenus Masdevallia Section Amaluzae: 6 species, e.g. M. amaluzae, M. carmenensis, M. patula Section Aphanes: 3 species – M. aphanes, M. capillaris, M. scopaea Section Coriaceae: Subsection Coriaceae: ~35 species, e.g. M. angulata, M. caesia, M. civilis, M. elephanticeps, M. foetens, M. fractiflexa Subsection Durae: 4 species – M. ayabacana, M. dura, M. panguiensis, M. utriculata Section Ligiae: Monotypic – M. ligiae Section Masdevallia: Subsection Caudatae: ~28 species, e.g. M. bottae, M. caudata, M. decumana, M. lychniphora, M. triangularis, M. xanthina Subsection Coccineae: 12 species, e.g. M. amabilis, M. barlaeana, M. coccinea, M. ignea, M. veitchiana Subsection Masdevallia: ~58 species, e.g. M. agaster, M. calocodon, M. mejiana, M. pumila, M. uniflora Subsection Oscillantes: ~11 species, e.g. M. andreettana, M. wageneriana Subsection Saltatrices: 14 species, e.g. M. angulifera, M. constricta, M. limax, M. saltatrix, M. urosalpinx, M. ventricularia Subsection Tubulosae: 7 species, e.g. M. bangii, M. irapana, M. tubulosa Section Mentosae: Monotypic – M. mentosa Section Minutae: ~21 species, e.g. M. floribunda, M. herradurae, M. minuta, M. nicaraguae, M. wendlandiana Section Racemosae: Monotypic – M. racemosa Section Reichenbachianae: Subsection Dentatae: 2 species – M. collina, M. macrogenia Subsection Reichenbachianae: ~11 species, e.g. M. rolfeana, M. schroderiana, M. striatella Subgenus Meleagris: 7 species, e.g. M. anisomorpha, M. heteroptera, M. meleagris Subgenus Nidifica: 4-5 species: e.g. M. dynastes, M. nidifica Subgenus Polyantha Section Alaticaules: 97 species, e.g. M. bicolor, M. infracta, M. scobina, M. stenorrhynchos, M. tovarensis, M. vargasii, M. weberbaueri Section Polyanthae: 7 species, e.g. M. discoidea, M. lata, M. polyantha, M. schlimii Subgenus Scabripes Subgenus Volvula Gerritsen, Mary E. & Parsons, Ron: Masdevallias, Gems of the Orchid World.
Timber Press. Luer, Carlyle A.: Icones Pleurothallidinarum, Systematics of Masdevallia. Missouri Botanical Garden Press. Pridgeon, A. M. Solano, R. & Chase, M. W.: Phylogenetic relationships in Pleurothallidinae: combined evidence from nuclear and plastid DNA sequences. American Journal of Botany 88: 2286-2308. Med