Crenatula is a genus in the family Pteriidae. List of molluscan genera represented in the fossil record Tëmkin I. Molecular phylogeny of pearl oysters and their relatives. BMC Evolutionary Biology 10: 342 Media related to Crenatula at Wikimedia Commons Crenatula at the Encyclopedia of Life Crenatula at Biolib.cz Crenatula at the World Register of Marine Species
Mollusca is the second largest phylum of invertebrate animals. The members are known as mollusks. Around 85,000 extant species of molluscs are recognized; the number of fossil species is estimated between 100,000 additional species. Molluscs are the largest marine phylum, comprising about 23% of all the named marine organisms. Numerous molluscs live in freshwater and terrestrial habitats, they are diverse, not just in size and in anatomical structure, but in behaviour and in habitat. The phylum is divided into 8 or 9 taxonomic classes, of which two are extinct. Cephalopod molluscs, such as squid and octopus, are among the most neurologically advanced of all invertebrates—and either the giant squid or the colossal squid is the largest known invertebrate species; the gastropods are by far the most numerous molluscs and account for 80% of the total classified species. The three most universal features defining modern molluscs are a mantle with a significant cavity used for breathing and excretion, the presence of a radula, the structure of the nervous system.
Other than these common elements, molluscs express great morphological diversity, so many textbooks base their descriptions on a "hypothetical ancestral mollusc". This has a single, "limpet-like" shell on top, made of proteins and chitin reinforced with calcium carbonate, is secreted by a mantle covering the whole upper surface; the underside of the animal consists of a single muscular "foot". Although molluscs are coelomates, the coelom tends to be small; the main body cavity is a hemocoel. The "generalized" mollusc's feeding system consists of a rasping "tongue", the radula, a complex digestive system in which exuded mucus and microscopic, muscle-powered "hairs" called cilia play various important roles; the generalized mollusc has three in bivalves. The brain, in species that have one, encircles the esophagus. Most molluscs have eyes, all have sensors to detect chemicals and touch; the simplest type of molluscan reproductive system relies on external fertilization, but more complex variations occur.
All produce eggs, from which may emerge trochophore larvae, more complex veliger larvae, or miniature adults. The coelomic cavity is reduced, they have kidney-like organs for excretion. Good evidence exists for the appearance of gastropods and bivalves in the Cambrian period, 541 to 485.4 million years ago. However, the evolutionary history both of molluscs' emergence from the ancestral Lophotrochozoa and of their diversification into the well-known living and fossil forms are still subjects of vigorous debate among scientists. Molluscs still are an important food source for anatomically modern humans. There is a risk of food poisoning from toxins which can accumulate in certain molluscs under specific conditions and because of this, many countries have regulations to reduce this risk. Molluscs have, for centuries been the source of important luxury goods, notably pearls, mother of pearl, Tyrian purple dye, sea silk, their shells have been used as money in some preindustrial societies. Mollusc species can represent hazards or pests for human activities.
The bite of the blue-ringed octopus is fatal, that of Octopus apollyon causes inflammation that can last for over a month. Stings from a few species of large tropical cone shells can kill, but their sophisticated, though produced, venoms have become important tools in neurological research. Schistosomiasis is transmitted to humans via water snail hosts, affects about 200 million people. Snails and slugs can be serious agricultural pests, accidental or deliberate introduction of some snail species into new environments has damaged some ecosystems; the words mollusc and mollusk are both derived from the French mollusque, which originated from the Latin molluscus, from mollis, soft. Molluscus was itself an adaptation of Aristotle's τὰ μαλάκια ta malákia, which he applied inter alia to cuttlefish; the scientific study of molluscs is accordingly called malacology. The name Molluscoida was used to denote a division of the animal kingdom containing the brachiopods and tunicates, the members of the three groups having been supposed to somewhat resemble the molluscs.
As it is now known these groups have no relation to molluscs, little to one another, the name Molluscoida has been abandoned. The most universal features of the body structure of molluscs are a mantle with a significant cavity used for breathing and excretion, the organization of the nervous system. Many have a calcareous shell. Molluscs have developed such a varied range of body structures, it is difficult to find synapomorphies to apply to all modern groups; the most general characteristic of molluscs is they are bilaterally symmetrical. The following are present in all modern molluscs: The dorsal part of the body wall is a mantle which secretes calcareous spicules, plates or shells, it overlaps the body with enough spare room to form a mantle cavity. The anus and genitals open into the mantle cavity. There are two pairs of main nerve cords. Other characteristics that appear in textbooks have significant exceptions: Estimates of accepted described living species of molluscs vary from 50,000 to a maximum of 120,000 species.
In 1969 David Nicol estimated the probable total number of living mollusc species at 107,000 of which were ab
An ocean is a body of water that composes much of a planet's hydrosphere. On Earth, an ocean is one of the major conventional divisions of the World Ocean; these are, in descending order by area, the Pacific, Indian and Arctic Oceans. The word "ocean" is used interchangeably with "sea" in American English. Speaking, a sea is a body of water or enclosed by land, though "the sea" refers to the oceans. Saline water covers 361,000,000 km2 and is customarily divided into several principal oceans and smaller seas, with the ocean covering 71% of Earth's surface and 90% of the Earth's biosphere; the ocean contains 97% of Earth's water, oceanographers have stated that less than 5% of the World Ocean has been explored. The total volume is 1.35 billion cubic kilometers with an average depth of nearly 3,700 meters. As the world ocean is the principal component of Earth's hydrosphere, it is integral to life, forms part of the carbon cycle, influences climate and weather patterns; the World Ocean is the habitat of 230,000 known species, but because much of it is unexplored, the number of species that exist in the ocean is much larger over two million.
The origin of Earth's oceans is unknown. Extraterrestrial oceans may be composed of water or other compounds; the only confirmed large stable bodies of extraterrestrial surface liquids are the lakes of Titan, although there is evidence for the existence of oceans elsewhere in the Solar System. Early in their geologic histories and Venus are theorized to have had large water oceans; the Mars ocean hypothesis suggests that nearly a third of the surface of Mars was once covered by water, a runaway greenhouse effect may have boiled away the global ocean of Venus. Compounds such as salts and ammonia dissolved in water lower its freezing point so that water might exist in large quantities in extraterrestrial environments as brine or convecting ice. Unconfirmed oceans are speculated beneath the surface of natural satellites; the Solar System's giant planets are thought to have liquid atmospheric layers of yet to be confirmed compositions. Oceans may exist on exoplanets and exomoons, including surface oceans of liquid water within a circumstellar habitable zone.
Ocean planets are a hypothetical type of planet with a surface covered with liquid. The word ocean comes from the figure in classical antiquity, the elder of the Titans in classical Greek mythology, believed by the ancient Greeks and Romans to be the divine personification of the sea, an enormous river encircling the world; the concept of Ōkeanós has an Indo-European connection. Greek Ōkeanós has been compared to the Vedic epithet ā-śáyāna-, predicated of the dragon Vṛtra-, who captured the cows/rivers. Related to this notion, the Okeanos is represented with a dragon-tail on some early Greek vases. Though described as several separate oceans, the global, interconnected body of salt water is sometimes referred to as the World Ocean or global ocean; the concept of a continuous body of water with free interchange among its parts is of fundamental importance to oceanography. The major oceanic divisions – listed below in descending order of area and volume – are defined in part by the continents, various archipelagos, other criteria.
Oceans are fringed by smaller, adjoining bodies of water such as seas, bays and straits. The mid-ocean ridges of the world are connected and form a single global mid-oceanic ridge system, part of every ocean and the longest mountain range in the world; the continuous mountain range is 65,000 km long. The total mass of the hydrosphere is about 1.4 quintillion metric tons, about 0.023% of Earth's total mass. Less than 3% is freshwater; the area of the World Ocean is about 361.9 million square kilometers, which covers about 70.9% of Earth's surface, its volume is 1.335 billion cubic kilometers. This can be thought of as a cube of water with an edge length of 1,101 kilometers, its average depth is about 3,688 meters, its maximum depth is 10,994 meters at the Mariana Trench. Nearly half of the world's marine waters are over 3,000 meters deep; the vast expanses of deep ocean cover about 66% of Earth's surface. This does not include seas not connected to the World Ocean, such as the Caspian Sea; the bluish ocean color is a composite of several contributing agents.
Prominent contributors include dissolved organic chlorophyll. Mariners and other seafarers have reported that the ocean emits a visible glow which extends for miles at night. In 2005, scientists announced that for the first time, they had obtained photographic evidence of this glow, it is most caused by bioluminescence. Oceanographers divide the ocean into different vertical zones defined by physical and biological conditions; the pelagic zone includes all open ocean regions, can be divided into further regions categorized by depth and light abundance. The photic zone includes the oceans from the surface to a depth of
Animals are multicellular eukaryotic organisms that form the biological kingdom Animalia. With few exceptions, animals consume organic material, breathe oxygen, are able to move, can reproduce sexually, grow from a hollow sphere of cells, the blastula, during embryonic development. Over 1.5 million living animal species have been described—of which around 1 million are insects—but it has been estimated there are over 7 million animal species in total. Animals range in length from 8.5 millionths of a metre to 33.6 metres and have complex interactions with each other and their environments, forming intricate food webs. The category includes humans, but in colloquial use the term animal refers only to non-human animals; the study of non-human animals is known as zoology. Most living animal species are in the Bilateria, a clade whose members have a bilaterally symmetric body plan; the Bilateria include the protostomes—in which many groups of invertebrates are found, such as nematodes and molluscs—and the deuterostomes, containing the echinoderms and chordates.
Life forms interpreted. Many modern animal phyla became established in the fossil record as marine species during the Cambrian explosion which began around 542 million years ago. 6,331 groups of genes common to all living animals have been identified. Aristotle divided animals into those with those without. Carl Linnaeus created the first hierarchical biological classification for animals in 1758 with his Systema Naturae, which Jean-Baptiste Lamarck expanded into 14 phyla by 1809. In 1874, Ernst Haeckel divided the animal kingdom into the multicellular Metazoa and the Protozoa, single-celled organisms no longer considered animals. In modern times, the biological classification of animals relies on advanced techniques, such as molecular phylogenetics, which are effective at demonstrating the evolutionary relationships between animal taxa. Humans make use of many other animal species for food, including meat and eggs. Dogs have been used in hunting, while many aquatic animals are hunted for sport.
Non-human animals have appeared in art from the earliest times and are featured in mythology and religion. The word "animal" comes from the Latin animalis, having soul or living being; the biological definition includes all members of the kingdom Animalia. In colloquial usage, as a consequence of anthropocentrism, the term animal is sometimes used nonscientifically to refer only to non-human animals. Animals have several characteristics. Animals are eukaryotic and multicellular, unlike bacteria, which are prokaryotic, unlike protists, which are eukaryotic but unicellular. Unlike plants and algae, which produce their own nutrients animals are heterotrophic, feeding on organic material and digesting it internally. With few exceptions, animals breathe oxygen and respire aerobically. All animals are motile during at least part of their life cycle, but some animals, such as sponges, corals and barnacles become sessile; the blastula is a stage in embryonic development, unique to most animals, allowing cells to be differentiated into specialised tissues and organs.
All animals are composed of cells, surrounded by a characteristic extracellular matrix composed of collagen and elastic glycoproteins. During development, the animal extracellular matrix forms a flexible framework upon which cells can move about and be reorganised, making the formation of complex structures possible; this may be calcified, forming structures such as shells and spicules. In contrast, the cells of other multicellular organisms are held in place by cell walls, so develop by progressive growth. Animal cells uniquely possess the cell junctions called tight junctions, gap junctions, desmosomes. With few exceptions—in particular, the sponges and placozoans—animal bodies are differentiated into tissues; these include muscles, which enable locomotion, nerve tissues, which transmit signals and coordinate the body. There is an internal digestive chamber with either one opening or two openings. Nearly all animals make use of some form of sexual reproduction, they produce haploid gametes by meiosis.
These fuse to form zygotes, which develop via mitosis into a hollow sphere, called a blastula. In sponges, blastula larvae swim to a new location, attach to the seabed, develop into a new sponge. In most other groups, the blastula undergoes more complicated rearrangement, it first invaginates to form a gastrula with a digestive chamber and two separate germ layers, an external ectoderm and an internal endoderm. In most cases, a third germ layer, the mesoderm develops between them; these germ layers differentiate to form tissues and organs. Repeated instances of mating with a close relative during sexual reproduction leads to inbreeding depression within a population due to the increased prevalence of harmful recessive traits. Animals have evolved numerous mechanisms for avoiding close inbreeding. In some species, such as the splendid fairywren, females benefit by mating with multiple males, thus producing more offspring of higher genetic quality; some animals are capable of asexual reproduction, which results
Pteria is a genus of molluscs in the family Pteriidae. The species of the genus are sometimes referred to as winged oysters; the World Register of Marine Species includes the following species in the genus: Pteria admirabilis Wang, 2002 Pteria aegyptiaca Pteria avicular Pteria bernhardi Pteria broomei Huber, 2010 Pteria brunnea Pteria bulliformis Wang, 2002 Pteria colymbus — Atlantic wing-oyster Pteria cooki Lamprell & Healy, 1997 Pteria dendronephthya Habe, 1960 Pteria fibrosa Pteria formosa Pteria gregata Pteria heteroptera Pteria hirundo Linnaeus, 1758 Pteria howensis Lamprell & Healy, 1997 Pteria lata Pteria maura Pteria peasei ‒ swift wing oyster Pteria penguin — Penguin's wing oyster Pteria physoides Pteria saltata Pteria spectrum Pteria sterna Gould, 1851 — Pacific wing-oyster Pteria straminea Pteria tortirostris Pteria venezuelensis
Pinctada is a genus of saltwater oysters, marine bivalve mollusks in the family Pteriidae, the pearl oysters. These oysters have a strong inner shell layer composed of nacre known as "mother of pearl". Pearl oysters are not related to either the edible oysters of family Ostreidae or the freshwater pearl mussels of the families Unionidae and Margaritiferidae. P. margaritifera and P. maxima are used for culturing South Tahitian pearls. They are cultured primarily in the central and eastern Indo-Pacific. A pearl oyster can be seen on the reverse side of the 1,000-peso bill of the Philippines. All species within the genus produce pearls. Attempts have been made to harvest pearls commercially from many Pinctada species. However, the only species that are of significant commercial interest are: Gulf pearl oyster, Pinctada radiata. Black-lip oyster, Pinctada margaritifera. Black South Sea pearls. Pearls are obtained in commercial quantities from some species of the related winged oyster genus Pteria. Pearls are produced from freshwater mussel species unrelated to pearl oysters.
These freshwater species include Hyriopsis cumingii, Hyriopsis schlegelii, a hybrid of the two species. At danger from the large demand for pearls, the typical lifespan of a pearl oyster is around 3 years to 14 years. Pinctada maxima are seeded at about 2 years of age and take 2 years to develop a pearl, they can be reseeded up to 4 times. Akoya pearls are harvested after about 9 to 16 months. Research carried out by biologist Aldemaro Romero Jr. allowed him to discover that the first animal population depleted by Europeans in the American continent was a pearl oyster species off the coast of Venezuela. He analyzed historical records and used information about the biology of these and other species to explain its rapid disappearance; the World Register of Marine Species includes the following species in the genus: Pinctada albina – Shark Bay pearl oyster Pinctada capensis Pinctada chemnitzii Pinctada cumingii Pinctada fucata – Akoya pearl oyster – now accepted as Pinctada imbricata fucata Pinctada galtsoffi Bartsch, 1931 Pinctada imbricata Röding, 1798 – Gulf pearl oyster Pinctada inflata Pinctada longisquamosa Pinctada maculata Pinctada margaritifera – Black-lip oyster Pinctada maxima – White-lip oyster or gold-lip oyster Pinctada mazatlanica Pinctada nigra Pinctada petersii Pinctada radiata – Atlantic pearl-oyster – now accepted as Pinctada imbricata radiata Pinctada reeveana Pinctada sugillata Pinctada vidua Media related to Pinctada at Wikimedia Commons OBIS Indo-Pacific molluscan database entry for Pinctada
Wikispecies is a wiki-based online project supported by the Wikimedia Foundation. Its aim is to create a comprehensive free content catalogue of all species. Jimmy Wales stated that editors are not required to fax in their degrees, but that submissions will have to pass muster with a technical audience. Wikispecies is available under the GNU Free Documentation License and CC BY-SA 3.0. Started in September 2004, with biologists across the world invited to contribute, the project had grown a framework encompassing the Linnaean taxonomy with links to Wikipedia articles on individual species by April 2005. Benedikt Mandl co-ordinated the efforts of several people who are interested in getting involved with the project and contacted potential supporters in early summer 2004. Databases were evaluated and the administrators contacted, some of them have agreed on providing their data for Wikispecies. Mandl defined two major tasks: Figure out how the contents of the data base would need to be presented—by asking experts, potential non-professional users and comparing that with existing databases Figure out how to do the software, which hardware is required and how to cover the costs—by asking experts, looking for fellow volunteers and potential sponsorsAdvantages and disadvantages were discussed by the wikimedia-I mailing list.
The board of directors of the Wikimedia Foundation voted by 4 to 0 in favor of the establishment of a Wikispecies. The project is hosted at species.wikimedia.org. It was merged to a sister project of Wikimedia Foundation on September 14, 2004. On October 10, 2006, the project exceeded 75,000 articles. On May 20, 2007, the project exceeded 100,000 articles with a total of 5,495 registered users. On September 8, 2008, the project exceeded 150,000 articles with a total of 9,224 registered users. On October 23, 2011, the project reached 300,000 articles. On June 16, 2014, the project reached 400,000 articles. On January 7, 2017, the project reached 500,000 articles. On October 30, 2018, the project reached 600,000 articles, a total of 1.12 million pages. Wikispecies comprises taxon pages, additionally pages about synonyms, taxon authorities, taxonomical publications, institutions or repositories holding type specimen. Wikispecies asks users to use images from Wikimedia Commons. Wikispecies does not allow the use of content.
All Species Foundation Catalogue of Life Encyclopedia of Life Tree of Life Web Project List of online encyclopedias The Plant List Wikispecies, The free species directory that anyone can edit Species Community Portal The Wikispecies Charter, written by Wales