The Precambrian is the earliest part of Earth's history, set before the current Phanerozoic Eon. The Precambrian is so named because it preceded the Cambrian, the first period of the Phanerozoic eon, named after Cambria, the Latinised name for Wales, where rocks from this age were first studied; the Precambrian accounts for 88% of the Earth's geologic time. The Precambrian is an informal unit of geologic time, subdivided into three eons of the geologic time scale, it spans from the formation of Earth about 4.6 billion years ago to the beginning of the Cambrian Period, about 541 million years ago, when hard-shelled creatures first appeared in abundance. Little is known about the Precambrian, despite it making up seven-eighths of the Earth's history, what is known has been discovered from the 1960s onwards; the Precambrian fossil record is poorer than that of the succeeding Phanerozoic, fossils from the Precambrian are of limited biostratigraphic use. This is because many Precambrian rocks have been metamorphosed, obscuring their origins, while others have been destroyed by erosion, or remain buried beneath Phanerozoic strata.
It is thought that the Earth coalesced from material in orbit around the Sun at 4,543 Ma, may have been struck by a large planetesimal shortly after it formed, splitting off material that formed the Moon. A stable crust was in place by 4,433 Ma, since zircon crystals from Western Australia have been dated at 4,404 ± 8 Ma; the term "Precambrian" is recognized by the International Commission on Stratigraphy as the only "supereon" in geologic time. "Precambrian" is still used by geologists and paleontologists for general discussions not requiring the more specific eon names. As of 2010, the United States Geological Survey considers the term informal, lacking a stratigraphic rank. A specific date for the origin of life has not been determined. Carbon found in 3.8 billion-year-old rocks from islands off western Greenland may be of organic origin. Well-preserved microscopic fossils of bacteria older than 3.46 billion years have been found in Western Australia. Probable fossils 100 million years older have been found in the same area.
However, there is evidence. There is a solid record of bacterial life throughout the remainder of the Precambrian. Excluding a few contested reports of much older forms from North America and India, the first complex multicellular life forms seem to have appeared at 1500 Ma, in the Mesoproterozoic era of the Proterozoic eon. Fossil evidence from the Ediacaran period of such complex life comes from the Lantian formation, at least 580 million years ago. A diverse collection of soft-bodied forms is found in a variety of locations worldwide and date to between 635 and 542 Ma; these are referred to as Vendian biota. Hard-shelled creatures appeared toward the end of that time span, marking the beginning of the Phanerozoic eon. By the middle of the following Cambrian period, a diverse fauna is recorded in the Burgess Shale, including some which may represent stem groups of modern taxa; the increase in diversity of lifeforms during the early Cambrian is called the Cambrian explosion of life. While land seems to have been devoid of plants and animals and other microbes formed prokaryotic mats that covered terrestrial areas.
Tracks from an animal with leg like appendages have been found in what was mud 551 million years ago. Evidence of the details of plate motions and other tectonic activity in the Precambrian has been poorly preserved, it is believed that small proto-continents existed prior to 4280 Ma, that most of the Earth's landmasses collected into a single supercontinent around 1130 Ma. The supercontinent, known as Rodinia, broke up around 750 Ma. A number of glacial periods have been identified going as far back as the Huronian epoch 2400–2100 Ma. One of the best studied is the Sturtian-Varangian glaciation, around 850–635 Ma, which may have brought glacial conditions all the way to the equator, resulting in a "Snowball Earth"; the atmosphere of the early Earth is not well understood. Most geologists believe it was composed of nitrogen, carbon dioxide, other inert gases, was lacking in free oxygen. There is, evidence that an oxygen-rich atmosphere existed since the early Archean. At present, it is still believed that molecular oxygen was not a significant fraction of Earth's atmosphere until after photosynthetic life forms evolved and began to produce it in large quantities as a byproduct of their metabolism.
This radical shift from a chemically inert to an oxidizing atmosphere caused an ecological crisis, sometimes called the oxygen catastrophe. At first, oxygen would have combined with other elements in Earth's crust iron, removing it from the atmosphere. After the supply of oxidizable surfaces ran out, oxygen would have begun to accumulate in the atmosphere, the modern high-oxygen atmosphere would have developed. Evidence for this lies in older rocks that contain massive banded iron formations that were laid down as iron oxides. A terminology has evolved covering the early years of the Earth's existence, as radiometric dating has allowed real dates to be assigned to specific formations and features; the Precambrian is divided into
The Silurian is a geologic period and system spanning 24.6 million years from the end of the Ordovician Period, at 443.8 million years ago, to the beginning of the Devonian Period, 419.2 Mya. The Silurian is the shortest period of the Paleozoic Era; as with other geologic periods, the rock beds that define the period's start and end are well identified, but the exact dates are uncertain by several million years. The base of the Silurian is set at a series of major Ordovician–Silurian extinction events when 60% of marine species were wiped out. A significant evolutionary milestone during the Silurian was the diversification of jawed fish and bony fish. Multi-cellular life began to appear on land in the form of small, bryophyte-like and vascular plants that grew beside lakes and coastlines, terrestrial arthropods are first found on land during the Silurian. However, terrestrial life would not diversify and affect the landscape until the Devonian; the Silurian system was first identified by British geologist Roderick Murchison, examining fossil-bearing sedimentary rock strata in south Wales in the early 1830s.
He named the sequences for a Celtic tribe of Wales, the Silures, inspired by his friend Adam Sedgwick, who had named the period of his study the Cambrian, from the Latin name for Wales. This naming does not indicate any correlation between the occurrence of the Silurian rocks and the land inhabited by the Silures. In 1835 the two men presented a joint paper, under the title On the Silurian and Cambrian Systems, Exhibiting the Order in which the Older Sedimentary Strata Succeed each other in England and Wales, the germ of the modern geological time scale; as it was first identified, the "Silurian" series when traced farther afield came to overlap Sedgwick's "Cambrian" sequence, provoking furious disagreements that ended the friendship. Charles Lapworth resolved the conflict by defining a new Ordovician system including the contested beds. An early alternative name for the Silurian was "Gotlandian" after the strata of the Baltic island of Gotland; the French geologist Joachim Barrande, building on Murchison's work, used the term Silurian in a more comprehensive sense than was justified by subsequent knowledge.
He divided the Silurian rocks of Bohemia into eight stages. His interpretation was questioned in 1854 by Edward Forbes, the stages of Barrande, F, G and H, have since been shown to be Devonian. Despite these modifications in the original groupings of the strata, it is recognized that Barrande established Bohemia as a classic ground for the study of the earliest fossils; the Llandovery Epoch lasted from 443.8 ± 1.5 to 433.4 ± 2.8 mya, is subdivided into three stages: the Rhuddanian, lasting until 440.8 million years ago, the Aeronian, lasting to 438.5 million years ago, the Telychian. The epoch is named for the town of Llandovery in Wales; the Wenlock, which lasted from 433.4 ± 1.5 to 427.4 ± 2.8 mya, is subdivided into the Sheinwoodian and Homerian ages. It is named after Wenlock Edge in England. During the Wenlock, the oldest-known tracheophytes of the genus Cooksonia, appear; the complexity of later Gondwana plants like Baragwanathia, which resembled a modern clubmoss, indicates a much longer history for vascular plants, extending into the early Silurian or Ordovician.
The first terrestrial animals appear in the Wenlock, represented by air-breathing millipedes from Scotland. The Ludlow, lasting from 427.4 ± 1.5 to 423 ± 2.8 mya, comprises the Gorstian stage, lasting until 425.6 million years ago, the Ludfordian stage. It is named for the town of Ludlow in England; the Přídolí, lasting from 423 ± 1.5 to 419.2 ± 2.8 mya, is the final and shortest epoch of the Silurian. It is named after one locality at the Homolka a Přídolí nature reserve near the Prague suburb Slivenec in the Czech Republic. Přídolí is the old name of a cadastral field area. In North America a different suite of regional stages is sometimes used: Cayugan Lockportian Tonawandan Ontarian Alexandrian In Estonia the following suite of regional stages is used: Ohessaare stage Kaugatuma stage Kuressaare stage Paadla stage Rootsiküla stage Jaagarahu stage Jaani stage Adavere stage Raikküla stage Juuru stage With the supercontinent Gondwana covering the equator and much of the southern hemisphere, a large ocean occupied most of the northern half of the globe.
The high sea levels of the Silurian and the flat land resulted in a number of island chains, thus a rich diversity of environmental settings. During the Silurian, Gondwana continued a slow southward drift to high southern latitudes, but there is evidence that the Silurian icecaps were less extensive than those of the late-Ordovician glaciation; the southern continents remained united during this period. The melting of icecaps and glaciers contributed to a rise in sea level, recognizable from the fact that Silurian sediments overlie eroded Ordovician sediments, forming an unconformity; the continents of Avalonia and Laurentia drifted together near the equator, starting the formation of a second supercontinent known as Euramerica. When the proto-Europe coll
In terrestrial animals, plantigrade locomotion means walking with the toes and metatarsals flat on the ground. It is one of three forms of locomotion adopted by terrestrial mammals; the other options are digitigrade, walking on the toes with the heel and wrist permanently raised, unguligrade, walking on the nail or nails of the toes with the heel/wrist and the digits permanently raised. The leg of a plantigrade mammal includes the bones of lower leg; the leg of a digitigrade mammal includes the metatarsals/metacarpals, the bones that in a human compose the arch of the foot and the palm of the hand. The leg of an unguligrade mammal includes the phalanges, the finger and toe bones. Among extinct animals, most early mammals such as pantodonts were plantigrade. A plantigrade foot is the primitive condition for mammals. Among archosaurs, the pterosaurs were plantigrade, walking on the whole of the hind foot and the fingers of the hand-wing. Plantigrade mammal species include: primates Carnivora: Procyonidae: raccoons Musteloidea: wolverines, skunks bears red pandas Barbourofelis rodents: mice, rats lagomorphs: rabbits hedgehogs hyraxes Marsupialia: kangaroosThe primary advantages of a plantigrade foot are stability and weight-bearing ability.
The primary disadvantage of a plantigrade foot is speed. With more bones and joints in the foot, the leg is both shorter and heavier at the far end, which makes it difficult to move rapidly. In humans and other great apes, another possible advantage of a plantigrade foot is that it may enhance fighting performance. Plantigrade foot occurs in humans in static postures of standing and sitting, it should occur in gait. Hypertonicity, clonus, limited range of motion, abnormal flexion neural pattern, a plantarflexor muscle contracture, as well as some forms of footwear such as high heeled shoes, may contribute to an individual only standing and/or walking on his or her toes; this would be evident by the observable heel rise
China the People's Republic of China, is a country in East Asia and the world's most populous country, with a population of around 1.404 billion. Covering 9,600,000 square kilometers, it is the third- or fourth-largest country by total area. Governed by the Communist Party of China, the state exercises jurisdiction over 22 provinces, five autonomous regions, four direct-controlled municipalities, the special administrative regions of Hong Kong and Macau. China emerged as one of the world's earliest civilizations, in the fertile basin of the Yellow River in the North China Plain. For millennia, China's political system was based on hereditary monarchies, or dynasties, beginning with the semi-legendary Xia dynasty in 21st century BCE. Since China has expanded, re-unified numerous times. In the 3rd century BCE, the Qin established the first Chinese empire; the succeeding Han dynasty, which ruled from 206 BC until 220 AD, saw some of the most advanced technology at that time, including papermaking and the compass, along with agricultural and medical improvements.
The invention of gunpowder and movable type in the Tang dynasty and Northern Song completed the Four Great Inventions. Tang culture spread in Asia, as the new Silk Route brought traders to as far as Mesopotamia and Horn of Africa. Dynastic rule ended in 1912 with the Xinhai Revolution; the Chinese Civil War resulted in a division of territory in 1949, when the Communist Party of China established the People's Republic of China, a unitary one-party sovereign state on Mainland China, while the Kuomintang-led government retreated to the island of Taiwan. The political status of Taiwan remains disputed. Since the introduction of economic reforms in 1978, China's economy has been one of the world's fastest-growing with annual growth rates above 6 percent. According to the World Bank, China's GDP grew from $150 billion in 1978 to $12.24 trillion by 2017. Since 2010, China has been the world's second-largest economy by nominal GDP and since 2014, the largest economy in the world by purchasing power parity.
China is the world's largest exporter and second-largest importer of goods. China is a recognized nuclear weapons state and has the world's largest standing army and second-largest defense budget; the PRC is a permanent member of the United Nations Security Council as it replaced the ROC in 1971, as well as an active global partner of ASEAN Plus mechanism. China is a leading member of numerous formal and informal multilateral organizations, including the Shanghai Cooperation Organization, WTO, APEC, BRICS, the BCIM, the G20. In recent times, scholars have argued that it will soon be a world superpower, rivaling the United States; the word "China" has been used in English since the 16th century. It is not a word used by the Chinese themselves, it has been traced through Portuguese and Persian back to the Sanskrit word Cīna, used in ancient India."China" appears in Richard Eden's 1555 translation of the 1516 journal of the Portuguese explorer Duarte Barbosa. Barbosa's usage was derived from Persian Chīn, in turn derived from Sanskrit Cīna.
Cīna was first used including the Mahābhārata and the Laws of Manu. In 1655, Martino Martini suggested that the word China is derived from the name of the Qin dynasty. Although this derivation is still given in various sources, it is complicated by the fact that the Sanskrit word appears in pre-Qin literature; the word may have referred to a state such as Yelang. The meaning transferred to China as a whole; the origin of the Sanskrit word is still a matter of debate, according to the Oxford English Dictionary. The official name of the modern state is the "People's Republic of China"; the shorter form is "China" Zhōngguó, from zhōng and guó, a term which developed under the Western Zhou dynasty in reference to its royal demesne. It was applied to the area around Luoyi during the Eastern Zhou and to China's Central Plain before being used as an occasional synonym for the state under the Qing, it was used as a cultural concept to distinguish the Huaxia people from perceived "barbarians". The name Zhongguo is translated as "Middle Kingdom" in English.
Archaeological evidence suggests that early hominids inhabited China between 2.24 million and 250,000 years ago. The hominid fossils of Peking Man, a Homo erectus who used fire, were discovered in a cave at Zhoukoudian near Beijing; the fossilized teeth of Homo sapiens have been discovered in Fuyan Cave in Hunan. Chinese proto-writing existed in Jiahu around 7000 BCE, Damaidi around 6000 BCE, Dadiwan from 5800–5400 BCE, Banpo dating from the 5th millennium BCE; some scholars have suggested. According to Chinese tradition, the first dynasty was the Xia, which emerged around 2100 BCE; the dynasty was considered mythical by historians until scientific excavations found early Bronze Age sites at Erlitou, Henan in 1959. It remains unclear whether these sites are the remains of the Xia dynasty or of another culture from the same period; the succeeding Shang dynasty is the earliest to be confirmed by contemporary records. The Shang ruled the plain of the Yellow River in eastern China from the 17th to the 11th century BCE.
Their oracle bone script
The Carboniferous is a geologic period and system that spans 60 million years from the end of the Devonian Period 358.9 million years ago, to the beginning of the Permian Period, 298.9 Mya. The name Carboniferous means "coal-bearing" and derives from the Latin words carbō and ferō, was coined by geologists William Conybeare and William Phillips in 1822. Based on a study of the British rock succession, it was the first of the modern'system' names to be employed, reflects the fact that many coal beds were formed globally during that time; the Carboniferous is treated in North America as two geological periods, the earlier Mississippian and the Pennsylvanian. Terrestrial animal life was well established by the Carboniferous period. Amphibians were the dominant land vertebrates, of which one branch would evolve into amniotes, the first terrestrial vertebrates. Arthropods were very common, many were much larger than those of today. Vast swaths of forest covered the land, which would be laid down and become the coal beds characteristic of the Carboniferous stratigraphy evident today.
The atmospheric content of oxygen reached its highest levels in geological history during the period, 35% compared with 21% today, allowing terrestrial invertebrates to evolve to great size. The half of the period experienced glaciations, low sea level, mountain building as the continents collided to form Pangaea. A minor marine and terrestrial extinction event, the Carboniferous rainforest collapse, occurred at the end of the period, caused by climate change. In the United States the Carboniferous is broken into Mississippian and Pennsylvanian subperiods; the Mississippian is about twice as long as the Pennsylvanian, but due to the large thickness of coal-bearing deposits with Pennsylvanian ages in Europe and North America, the two subperiods were long thought to have been more or less equal in duration. In Europe the Lower Carboniferous sub-system is known as the Dinantian, comprising the Tournaisian and Visean Series, dated at 362.5-332.9 Ma, the Upper Carboniferous sub-system is known as the Silesian, comprising the Namurian and Stephanian Series, dated at 332.9-298.9 Ma.
The Silesian is contemporaneous with the late Mississippian Serpukhovian plus the Pennsylvanian. In Britain the Dinantian is traditionally known as the Carboniferous Limestone, the Namurian as the Millstone Grit, the Westphalian as the Coal Measures and Pennant Sandstone; the International Commission on Stratigraphy faunal stages from youngest to oldest, together with some of their regional subdivisions, are: A global drop in sea level at the end of the Devonian reversed early in the Carboniferous. There was a drop in south polar temperatures; these conditions had little effect in the deep tropics, where lush swamps to become coal, flourished to within 30 degrees of the northernmost glaciers. Mid-Carboniferous, a drop in sea level precipitated a major marine extinction, one that hit crinoids and ammonites hard; this sea level drop and the associated unconformity in North America separate the Mississippian subperiod from the Pennsylvanian subperiod. This happened about 323 million years ago, at the onset of the Permo-Carboniferous Glaciation.
The Carboniferous was a time of active mountain-building as the supercontinent Pangaea came together. The southern continents remained tied together in the supercontinent Gondwana, which collided with North America–Europe along the present line of eastern North America; this continental collision resulted in the Hercynian orogeny in Europe, the Alleghenian orogeny in North America. In the same time frame, much of present eastern Eurasian plate welded itself to Europe along the line of the Ural Mountains. Most of the Mesozoic supercontinent of Pangea was now assembled, although North China, South China continents were still separated from Laurasia; the Late Carboniferous Pangaea was shaped like an "O." There were two major oceans in the Carboniferous—Panthalassa and Paleo-Tethys, inside the "O" in the Carboniferous Pangaea. Other minor oceans were shrinking and closed - Rheic Ocean, the small, shallow Ural Ocean and Proto-Tethys Ocean. Average global temperatures in the Early Carboniferous Period were high: 20 °C.
However, cooling during the Middle Carboniferous reduced average global temperatures to about 12 °C. Lack of growth rings of fossilized trees suggest a lack of seasons of a tropical climate. Glaciations in Gondwana, triggered by Gondwana's southward movement, continued into the Permian and because of the lack of clear markers and breaks, the deposits of this glacial period are referred to as Permo-Carboniferous in age; the cooling and drying of the climate led to the Carboniferous Rainforest Collapse during the late Carboniferous. Tropical rainforests fragmented and were devastated by climate change. Carboniferous rocks in Europe and eastern North America consist of a repeated sequence of limestone, sandstone and coal beds. In North America, the early Carboniferous is marine
Dentition pertains to the development of teeth and their arrangement in the mouth. In particular, it is the characteristic arrangement and number of teeth in a given species at a given age; that is, the number and morpho-physiology of the teeth of an animal. Animals whose teeth are all of the same type, such as most non-mammalian vertebrates, are said to have homodont dentition, whereas those whose teeth differ morphologically are said to have heterodont dentition; the dentition of animals with two successions of teeth is referred to as diphyodont, while the dentition of animals with only one set of teeth throughout life is monophyodont. The dentition of animals in which the teeth are continuously discarded and replaced throughout life is termed polyphyodont; the dentition of animals in which the teeth are set in sockets in the jawbones is termed thecodont. The evolutionary origin of the vertebrate dentition remains contentious. Current theories suggest either an "outside-in" or "inside-out" evolutionary origin to teeth, with the dentition arising from odontodes on the skin surface moving into the mouth, or vice versa.
Despite this debate, it is accepted that vertebrate teeth are homologous to the dermal denticles found on the skin of basal Gnathostomes. Since the origin of teeth some 450mya, the vertebrate dentition has diversified within the reptiles and fish: however most of these groups continue to possess a long row of pointed or sharp-sided, undifferentiated teeth that are replaceable; the mammalian pattern is different. The teeth in the upper and lower jaws in mammals have evolved a close-fitting relationship such that they operate together as a unit. "They'occlude'", that is, the chewing surfaces of the teeth are so constructed that the upper and lower teeth are able to fit together, crushing, grinding or tearing the food caught between."All mammals except the monotremes, the xenarthrans, the pangolins, the cetaceans have up to four distinct types of teeth, with a maximum number for each. These are the incisor, the canine, the premolar, the molar; the incisors occupy the front of the tooth row in lower jaws.
They are flat, chisel-shaped teeth that meet in an edge-to-edge bite. Their function is cutting, slicing, or gnawing food into manageable pieces that fit into the mouth for further chewing; the canines are behind the incisors. In many mammals, the canines are pointed, tusk-shaped teeth, projecting beyond the level of the other teeth. In carnivores, they are offensive weapons for bringing down prey. In other mammals such as some primates, they are used to split open hard surfaced food; the premolars and molars are at the back of the mouth. Depending on the particular mammal and its diet, these two kinds of teeth prepare pieces of food to be swallowed by grinding, shearing, or crushing; the specialised teeth—incisors, canines and molars—are found in the same order in every mammal. In many mammals the infants have a set of teeth that are replaced by adult teeth; these are called primary teeth, baby teeth or milk teeth. Animals that have two sets of teeth, one followed by the other, are said to be diphyodont.
The dental formula for milk teeth is the same as for adult teeth except that the molars are missing. Because every mammal's teeth are specialised for different functions, many mammal groups have lost teeth not needed in their adaptation. Tooth form has undergone evolutionary modification as a result of natural selection for specialised feeding or other adaptations. Over time, different mammal groups have evolved distinct dental features, both in the number and type of teeth, in the shape and size of the chewing surface; the number of teeth of each type is written as a dental formula for one side of the mouth, or quadrant, with the upper and lower teeth shown on separate rows. The number of teeth in a mouth is twice that listed. In each set, incisors are indicated first, canines second, premolars third, molars, giving I:C:P:M. So for example, the formula 184.108.40.206 for upper teeth indicates 2 incisors, 1 canine, 2 premolars, 3 molars on one side of the upper mouth. The deciduous dental formula is notated in lowercase lettering preceded by the letter d: for example: di:dc:dp.
An animal's dentition for either deciduous or permanent teeth can thus be expressed as a dental formula, written in the form of a fraction, which can be written as I. C. P. MI. C. P. M, or I. C. P. M / I. C. P. M. For example, the following formulae show the deciduous and usual permanent dentition of all catarrhine primates, including humans: Deciduous: / × 2 = 20; this can be written as di2.dc1.dm2di2.dc1.dm2. Superscript and subscript denote upper and lower jaw, i.e. do not indicate mathematical operations. The dashes in the formula are not mathematical operators, but spacers.'d' denotes deciduous teeth. Another annotation is 220.127.116.11.1.0.2, if the fact that it pertains to deciduous teeth is stated, per examples found in some texts such as The Ca
A chordate is an animal constituting the phylum Chordata. During some period of their life cycle, chordates possess a notochord, a dorsal nerve cord, pharyngeal slits, an endostyle, a post-anal tail: these five anatomical features define this phylum. Chordates are bilaterally symmetric; the Chordata and Ambulacraria together form the superphylum Deuterostomia. Chordates are divided into three subphyla: Vertebrata. There are extinct taxa such as the Vetulicolia. Hemichordata has been presented as a fourth chordate subphylum, but now is treated as a separate phylum: hemichordates and Echinodermata form the Ambulacraria, the sister phylum of the Chordates. Of the more than 65,000 living species of chordates, about half are bony fish that are members of the superclass Osteichthyes. Chordate fossils have been found from as early as the Cambrian explosion, 541 million years ago. Cladistically, vertebrates - chordates with the notochord replaced by a vertebral column during development - are considered to be a subgroup of the clade Craniata, which consists of chordates with a skull.
The Craniata and Tunicata compose the clade Olfactores. Chordates form a phylum of animals that are defined by having at some stage in their lives all of the following anatomical features: A notochord, a stiff rod of cartilage that extends along the inside of the body. Among the vertebrate sub-group of chordates the notochord develops into the spine, in wholly aquatic species this helps the animal to swim by flexing its tail. A dorsal neural tube. In fish and other vertebrates, this develops into the spinal cord, the main communications trunk of the nervous system. Pharyngeal slits; the pharynx is the part of the throat behind the mouth. In fish, the slits are modified to form gills, but in some other chordates they are part of a filter-feeding system that extracts particles of food from the water in which the animals live. Post-anal tail. A muscular tail that extends backwards behind the anus. An endostyle; this is a groove in the ventral wall of the pharynx. In filter-feeding species it produces mucus to gather food particles, which helps in transporting food to the esophagus.
It stores iodine, may be a precursor of the vertebrate thyroid gland. There are soft constraints that separate chordates from certain other biological lineages, but are not part of the formal definition: All chordates are deuterostomes; this means. All chordates are based on a bilateral body plan. All chordates are coelomates, have a fluid filled body cavity called a coelom with a complete lining called peritoneum derived from mesoderm; the following schema is from the third edition of Vertebrate Palaeontology. The invertebrate chordate classes are from Fishes of the World. While it is structured so as to reflect evolutionary relationships, it retains the traditional ranks used in Linnaean taxonomy. Phylum Chordata †Vetulicolia? Subphylum Cephalochordata – Class Leptocardii Clade Olfactores Subphylum Tunicata – Class Ascidiacea Class Thaliacea Class Appendicularia Class Sorberacea Subphylum Vertebrata Infraphylum incertae sedis Cyclostomata Superclass'Agnatha' paraphyletic Class Myxini Class Petromyzontida or Hyperoartia Class †Conodonta Class †Myllokunmingiida Class †Pteraspidomorphi Class †Thelodonti Class †Anaspida Class †Cephalaspidomorphi Infraphylum Gnathostomata Class †Placodermi Class Chondrichthyes Class †Acanthodii Superclass Osteichthyes Class Actinopterygii Class Sarcopterygii Superclass Tetrapoda Class Amphibia Class Sauropsida Class Synapsida Craniates, one of the three subdivisions of chordates, all have distinct skulls.
They include the hagfish. Michael J. Benton commented that "craniates are characterized by their heads, just as chordates, or all deuterostomes, are by their tails". Most craniates are vertebrates; these consist of a series of bony or cartilaginous cylindrical vertebrae with neural arches that protect the spinal cord, with projections that link the vertebrae. However hagfish have incomplete braincases and no vertebrae, are therefore not regarded as vertebrates, but as members of the craniates, the group from which vertebrates are thought to have evolved; however the cladistic exclusion of hagfish from the vertebrates is controversial, as they ma