The Phrynosomatidae are a diverse family of lizards, sometimes classified as a subfamily, found from Panama to the extreme south of Canada. Many members of the group are adapted to life in hot, sandy deserts, although the spiny lizards prefer rocky deserts or relatively moist forest edges, the short-horned lizard lives in prairie or sagebrush environments; the group includes both egg-laying and viviparous species, with the latter being more common in species living at high elevations. The 136 species are organised into 9 genera in this subfamily; the earless taxa are sister genera. Family Phrynosomatidae Callisaurus Blainville, 1835 – zebra-tailed lizards Cophosaurus Troschel, 1852 – greater earless lizards Holbrookia Girard, 1851 – earless lizards Petrosaurus Boulenger, 1885 – California rock lizards Phrynosoma Wiegmann, 1828 – horned lizards Sceloporus Wiegmann, 1828 – spiny lizards Uma Baird, 1859 – fringe-toed lizards Urosaurus Hallowell, 1854 – tree and brush lizards Uta Baird & Girard, 1852 – side-blotched lizards Data related to Phrynosomatidae at Wikispecies Media related to Phrynosomatidae at Wikimedia Commons Phrynosomatidae Family
Mexico the United Mexican States, is a country in the southern portion of North America. It is bordered to the north by the United States. Covering 2,000,000 square kilometres, the nation is the fifth largest country in the Americas by total area and the 13th largest independent state in the world. With an estimated population of over 120 million people, the country is the eleventh most populous state and the most populous Spanish-speaking state in the world, while being the second most populous nation in Latin America after Brazil. Mexico is a federation comprising 31 states and Mexico City, a special federal entity, the capital city and its most populous city. Other metropolises in the state include Guadalajara, Puebla, Tijuana and León. Pre-Columbian Mexico dates to about 8000 BC and is identified as one of five cradles of civilization and was home to many advanced Mesoamerican civilizations such as the Olmec, Teotihuacan, Zapotec and Aztec before first contact with Europeans. In 1521, the Spanish Empire conquered and colonized the territory from its politically powerful base in Mexico-Tenochtitlan, administered as the viceroyalty of New Spain.
Three centuries the territory became a nation state following its recognition in 1821 after the Mexican War of Independence. The post-independence period was tumultuous, characterized by economic inequality and many contrasting political changes; the Mexican–American War led to a territorial cession of the extant northern territories to the United States. The Pastry War, the Franco-Mexican War, a civil war, two empires, the Porfiriato occurred in the 19th century; the Porfiriato was ended by the start of the Mexican Revolution in 1910, which culminated with the promulgation of the 1917 Constitution and the emergence of the country's current political system as a federal, democratic republic. Mexico has the 11th largest by purchasing power parity; the Mexican economy is linked to those of its 1994 North American Free Trade Agreement partners the United States. In 1994, Mexico became the first Latin American member of the Organisation for Economic Co-operation and Development, it is classified as an upper-middle income country by the World Bank and a newly industrialized country by several analysts.
The country is considered both a regional power and a middle power, is identified as an emerging global power. Due to its rich culture and history, Mexico ranks first in the Americas and seventh in the world for number of UNESCO World Heritage Sites. Mexico is an ecologically megadiverse country, ranking fourth in the world for its biodiversity. Mexico receives a huge number of tourists every year: in 2018, it was the sixth most-visited country in the world, with 39 million international arrivals. Mexico is a member of the United Nations, the World Trade Organization, the G8+5, the G20, the Uniting for Consensus group of the UN, the Pacific Alliance trade bloc. Mēxihco is the Nahuatl term for the heartland of the Aztec Empire, namely the Valley of Mexico and surrounding territories, with its people being known as the Mexica, it is believed to be a toponym for the valley which became the primary ethnonym for the Aztec Triple Alliance as a result, although it could have been the other way around.
In the colonial era, back when Mexico was called New Spain this territory became the Intendency of Mexico and after New Spain achieved independence from the Spanish Empire it came to be known as the State of Mexico with the new country being named after its capital: the City of Mexico, which itself was founded in 1524 on top of the ancient Mexica capital of Mexico-Tenochtitlan. Traditionally, the name Tenochtitlan was thought to come from Nahuatl tetl and nōchtli and is thought to mean "Among the prickly pears rocks". However, one attestation in the late 16th-century manuscript known as "the Bancroft dialogues" suggests the second vowel was short, so that the true etymology remains uncertain; the suffix -co is the Nahuatl locative, making the word a place name. Beyond that, the etymology is uncertain, it has been suggested that it is derived from Mextli or Mēxihtli, a secret name for the god of war and patron of the Mexica, Huitzilopochtli, in which case Mēxihco means "place where Huitzilopochtli lives".
Another hypothesis suggests that Mēxihco derives from a portmanteau of the Nahuatl words for "moon" and navel. This meaning might refer to Tenochtitlan's position in the middle of Lake Texcoco; the system of interconnected lakes, of which Texcoco formed the center, had the form of a rabbit, which the Mesoamericans pareidolically associated with the moon rabbit. Still another hypothesis suggests that the word is derived from Mēctli, the name of the goddess of maguey; the name of the city-state was transliterated to Spanish as México with the phonetic value of the letter x in Medieval Spanish, which represented the voiceless postalveolar fricative. This sound, as well as the voiced postalveolar fricative, represented by a j, evolved into a voiceless velar fricative during the 16th century; this led to the use of the variant Méjico in many publications in Spanish, most notably in Spain, whereas in Mexico and most other Spanish–speaking countries, México was the preferred spelling. In recent years, the Real Academia Española, which regulates the Spanish l
Reptiles are tetrapod animals in the class Reptilia, comprising today's turtles, snakes, lizards and their extinct relatives. The study of these traditional reptile orders combined with that of modern amphibians, is called herpetology; because some reptiles are more related to birds than they are to other reptiles, the traditional groups of "reptiles" listed above do not together constitute a monophyletic grouping or clade. For this reason, many modern scientists prefer to consider the birds part of Reptilia as well, thereby making Reptilia a monophyletic class, including all living Diapsids; the earliest known proto-reptiles originated around 312 million years ago during the Carboniferous period, having evolved from advanced reptiliomorph tetrapods that became adapted to life on dry land. Some early examples include Casineria. In addition to the living reptiles, there are many diverse groups that are now extinct, in some cases due to mass extinction events. In particular, the Cretaceous–Paleogene extinction event wiped out the pterosaurs, plesiosaurs and sauropods, as well as many species of theropods, including troodontids, dromaeosaurids and abelisaurids, along with many Crocodyliformes, squamates.
Modern non-avian reptiles inhabit all the continents except Antarctica, although some birds are found on the periphery of Antarctica. Several living subgroups are recognized: Testudines, 350 species. Reptiles are tetrapod vertebrates, creatures that either have four limbs or, like snakes, are descended from four-limbed ancestors. Unlike amphibians, reptiles do not have an aquatic larval stage. Most reptiles are oviparous, although several species of squamates are viviparous, as were some extinct aquatic clades – the fetus develops within the mother, contained in a placenta rather than an eggshell; as amniotes, reptile eggs are surrounded by membranes for protection and transport, which adapt them to reproduction on dry land. Many of the viviparous species feed their fetuses through various forms of placenta analogous to those of mammals, with some providing initial care for their hatchlings. Extant reptiles range in size from a tiny gecko, Sphaerodactylus ariasae, which can grow up to 17 mm to the saltwater crocodile, Crocodylus porosus, which can reach 6 m in length and weigh over 1,000 kg.
In the 13th century the category of reptile was recognized in Europe as consisting of a miscellany of egg-laying creatures, including "snakes, various fantastic monsters, assorted amphibians, worms", as recorded by Vincent of Beauvais in his Mirror of Nature. In the 18th century, the reptiles were, from the outset of classification, grouped with the amphibians. Linnaeus, working from species-poor Sweden, where the common adder and grass snake are found hunting in water, included all reptiles and amphibians in class "III – Amphibia" in his Systema Naturæ; the terms "reptile" and "amphibian" were interchangeable, "reptile" being preferred by the French. Josephus Nicolaus Laurenti was the first to formally use the term "Reptilia" for an expanded selection of reptiles and amphibians similar to that of Linnaeus. Today, the two groups are still treated under the same heading as herptiles, it was not until the beginning of the 19th century that it became clear that reptiles and amphibians are, in fact, quite different animals, Pierre André Latreille erected the class Batracia for the latter, dividing the tetrapods into the four familiar classes of reptiles, amphibians and mammals.
The British anatomist Thomas Henry Huxley made Latreille's definition popular and, together with Richard Owen, expanded Reptilia to include the various fossil "antediluvian monsters", including dinosaurs and the mammal-like Dicynodon he helped describe. This was not the only possible classification scheme: In the Hunterian lectures delivered at the Royal College of Surgeons in 1863, Huxley grouped the vertebrates into mammals and ichthyoids, he subsequently proposed the names of Ichthyopsida for the latter two groups. In 1866, Haeckel demonstrated that vertebrates could be divided based on their reproductive strategies, that reptiles and mammals were united by the amniotic egg; the terms "Sauropsida" and "Theropsida" were used again in 1916 by E. S. Goodrich to distinguish between lizards and their relatives on the one hand and mammals and their extinct relatives on the other. Goodrich supported this division by the nature of the hearts and blood vessels in each group, other features, such as the structure of the forebrain.
According to Goodrich, both lineages evolved from an earlier stem group, Protosauria in which he included some animals today considered reptile-like amphibians, as well as early reptiles. In 1956, D. M. S. Watson observed that the first two groups diverged early in reptilian history, so he divided Goodrich's Protosauria between them, he reinterpreted Sauropsida and Theropsida to exclude birds and mammals, respectively. Thus his Sauropsida included Procolophonia, Millerosauria, Squamata, Rhynchocephalia
Binomial nomenclature called binominal nomenclature or binary nomenclature, is a formal system of naming species of living things by giving each a name composed of two parts, both of which use Latin grammatical forms, although they can be based on words from other languages. Such a name is called a binomen, binominal name or a scientific name; the first part of the name – the generic name – identifies the genus to which the species belongs, while the second part – the specific name or specific epithet – identifies the species within the genus. For example, humans belong within this genus to the species Homo sapiens. Tyrannosaurus rex is the most known binomial; the formal introduction of this system of naming species is credited to Carl Linnaeus beginning with his work Species Plantarum in 1753. But Gaspard Bauhin, in as early as 1623, had introduced in his book Pinax theatri botanici many names of genera that were adopted by Linnaeus; the application of binomial nomenclature is now governed by various internationally agreed codes of rules, of which the two most important are the International Code of Zoological Nomenclature for animals and the International Code of Nomenclature for algae and plants.
Although the general principles underlying binomial nomenclature are common to these two codes, there are some differences, both in the terminology they use and in their precise rules. In modern usage, the first letter of the first part of the name, the genus, is always capitalized in writing, while that of the second part is not when derived from a proper noun such as the name of a person or place. Both parts are italicized when a binomial name occurs in normal text, thus the binomial name of the annual phlox is now written as Phlox drummondii. In scientific works, the authority for a binomial name is given, at least when it is first mentioned, the date of publication may be specified. In zoology "Patella vulgata Linnaeus, 1758"; the name "Linnaeus" tells the reader who it was that first published a description and name for this species of limpet. "Passer domesticus". The original name given by Linnaeus was Fringilla domestica; the ICZN does not require that the name of the person who changed the genus be given, nor the date on which the change was made, although nomenclatorial catalogs include such information.
In botany "Amaranthus retroflexus L." – "L." is the standard abbreviation used in botany for "Linnaeus". "Hyacinthoides italica Rothm. – Linnaeus first named this bluebell species Scilla italica. The name is composed of two word-forming elements: "bi", a Latin prefix for two, "-nomial", relating to a term or terms; the word "binomium" was used in Medieval Latin to mean a two-term expression in mathematics. Prior to the adoption of the modern binomial system of naming species, a scientific name consisted of a generic name combined with a specific name, from one to several words long. Together they formed a system of polynomial nomenclature; these names had two separate functions. First, to designate or label the species, second, to be a diagnosis or description. In a simple genus, containing only two species, it was easy to tell them apart with a one-word genus and a one-word specific name; such "polynomial names" may sometimes look like binomials, but are different. For example, Gerard's herbal describes various kinds of spiderwort: "The first is called Phalangium ramosum, Branched Spiderwort.
The other... is aptly termed Phalangium Ephemerum Virginianum, Soon-Fading Spiderwort of Virginia". The Latin phrases are short descriptions, rather than identifying labels; the Bauhins, in particular Caspar Bauhin, took some important steps towards the binomial system, by pruning the Latin descriptions, in many cases to two words. The adoption by biologists of a system of binomial nomenclature is due to Swedish botanist and physician Carl von Linné, more known by his Latinized name Carl Linnaeus, it was in his 1753 Species Plantarum that he first began using a one-word "trivial name" together with a generic name in a system of binomial nomenclature. This trivial name is what is now known as specific name; the Bauhins' genus names were retained in many of these, but the descriptive part was reduced to a single word. Linnaeus's trivial names introduced an important new idea, namely that the function of a name could be to give a species a unique label; this meant. Thus Gerard's Phalangium ephemerum virginianum became Tradescantia virgi
A chordate is an animal constituting the phylum Chordata. During some period of their life cycle, chordates possess a notochord, a dorsal nerve cord, pharyngeal slits, an endostyle, a post-anal tail: these five anatomical features define this phylum. Chordates are bilaterally symmetric; the Chordata and Ambulacraria together form the superphylum Deuterostomia. Chordates are divided into three subphyla: Vertebrata. There are extinct taxa such as the Vetulicolia. Hemichordata has been presented as a fourth chordate subphylum, but now is treated as a separate phylum: hemichordates and Echinodermata form the Ambulacraria, the sister phylum of the Chordates. Of the more than 65,000 living species of chordates, about half are bony fish that are members of the superclass Osteichthyes. Chordate fossils have been found from as early as the Cambrian explosion, 541 million years ago. Cladistically, vertebrates - chordates with the notochord replaced by a vertebral column during development - are considered to be a subgroup of the clade Craniata, which consists of chordates with a skull.
The Craniata and Tunicata compose the clade Olfactores. Chordates form a phylum of animals that are defined by having at some stage in their lives all of the following anatomical features: A notochord, a stiff rod of cartilage that extends along the inside of the body. Among the vertebrate sub-group of chordates the notochord develops into the spine, in wholly aquatic species this helps the animal to swim by flexing its tail. A dorsal neural tube. In fish and other vertebrates, this develops into the spinal cord, the main communications trunk of the nervous system. Pharyngeal slits; the pharynx is the part of the throat behind the mouth. In fish, the slits are modified to form gills, but in some other chordates they are part of a filter-feeding system that extracts particles of food from the water in which the animals live. Post-anal tail. A muscular tail that extends backwards behind the anus. An endostyle; this is a groove in the ventral wall of the pharynx. In filter-feeding species it produces mucus to gather food particles, which helps in transporting food to the esophagus.
It stores iodine, may be a precursor of the vertebrate thyroid gland. There are soft constraints that separate chordates from certain other biological lineages, but are not part of the formal definition: All chordates are deuterostomes; this means. All chordates are based on a bilateral body plan. All chordates are coelomates, have a fluid filled body cavity called a coelom with a complete lining called peritoneum derived from mesoderm; the following schema is from the third edition of Vertebrate Palaeontology. The invertebrate chordate classes are from Fishes of the World. While it is structured so as to reflect evolutionary relationships, it retains the traditional ranks used in Linnaean taxonomy. Phylum Chordata †Vetulicolia? Subphylum Cephalochordata – Class Leptocardii Clade Olfactores Subphylum Tunicata – Class Ascidiacea Class Thaliacea Class Appendicularia Class Sorberacea Subphylum Vertebrata Infraphylum incertae sedis Cyclostomata Superclass'Agnatha' paraphyletic Class Myxini Class Petromyzontida or Hyperoartia Class †Conodonta Class †Myllokunmingiida Class †Pteraspidomorphi Class †Thelodonti Class †Anaspida Class †Cephalaspidomorphi Infraphylum Gnathostomata Class †Placodermi Class Chondrichthyes Class †Acanthodii Superclass Osteichthyes Class Actinopterygii Class Sarcopterygii Superclass Tetrapoda Class Amphibia Class Sauropsida Class Synapsida Craniates, one of the three subdivisions of chordates, all have distinct skulls.
They include the hagfish. Michael J. Benton commented that "craniates are characterized by their heads, just as chordates, or all deuterostomes, are by their tails". Most craniates are vertebrates; these consist of a series of bony or cartilaginous cylindrical vertebrae with neural arches that protect the spinal cord, with projections that link the vertebrae. However hagfish have incomplete braincases and no vertebrae, are therefore not regarded as vertebrates, but as members of the craniates, the group from which vertebrates are thought to have evolved; however the cladistic exclusion of hagfish from the vertebrates is controversial, as they ma
Herpetology is the branch of zoology concerned with the study of amphibians and reptiles. Birds, which are cladistically included within Reptilia, are traditionally excluded here. Thus, the definition of herpetology can be more stated as the study of ectothermic tetrapods. Under this definition "herps" exclude fish, but it is not uncommon for herpetological and ichthyological scientific societies to "team up", publishing joint journals and holding conferences in order to foster the exchange of ideas between the fields, as the American Society of Ichthyologists and Herpetologists does. Many herpetological societies have been formed to promote interest in reptiles and amphibians, both captive and wild. Herpetology offers benefits to humanity in the study of the role of amphibians and reptiles in global ecology because amphibians are very sensitive to environmental changes, offering a visible warning to humans that significant changes are taking place; some toxins and venoms produced by reptiles and amphibians are useful in human medicine.
Some snake venom has been used to create anti-coagulants that work to treat strokes and heart attacks. The word "herpetology" is from Greek: ἑρπετόν, herpeton, "creeping animal" and -λογία, -logia, "knowledge". People with an avid interest in herpetology and who keep different reptiles or amphibians refer to themselves as "herpers"."Herp" is a vernacular term for non-avian reptiles and amphibians. It is derived from the old term "herpetile", with roots back to Linnaeus's classification of animals, in which he grouped reptiles and amphibians together in the same class. There are over 9000 species of reptiles. In spite of its modern taxonomic irrelevance, the term has persisted in the names of herpetology, the scientific study of non-avian reptiles and amphibians, herpetoculture, the captive care and breeding of reptiles and amphibians; the field of herpetology amphibians. Batrachology, the study of amphibians in particular Ophiology, the study of snakes. Saurology, the study of lizards. Cheloniology, the study of turtles and tortoises.
Career options in the field of herpetology include, but are not limited to lab research, field studies and survey, zoological staff, museum staff and college teaching. In modern academic science, it is rare for individuals to consider themselves a herpetologist first and foremost. Most individuals focus on a particular field such as ecology, taxonomy, physiology, or molecular biology, within that field ask questions pertaining to or best answered by examining reptiles and amphibians. For example, an evolutionary biologist, a herpetologist may choose to work on an issue such as the evolution of warning coloration in coral snakes. Modern herpetological writers include Philip Purser. Modern herpetological showmen include Jeff Corwin, Steve Irwin, popularly known as the "Crocodile Hunter", the star Austin Stevens, popularly known as "AustinSnakeman" in the TV series Austin Stevens: Snakemaster. Most colleges or universities do not offer a major in herpetology at the undergraduate or the graduate level.
Instead, persons interested in herpetology select a major in the biological sciences. The knowledge learned about all aspects of the biology of animals is applied to an individual study of herpetology. Herping List of herpetologists List of herpetology academic journals Adler, Kraig. Contributions to the History of Herpetology. Society for the Study of Amphibians and Reptiles. Eatherley, Dan. Bushmaster: Raymond Ditmars and the Hunt for the World's Largest Viper. New York: Arcade. 320 pp. ISBN 978-1628725117. Goin, Coleman J.. Introduction to Herpetology, Third Edition. San Francisco: W. H. Freeman and Company. Xi + 378 pp. ISBN 0-7167-0020-4. Iranian Herpetological Studies Institute Field Herpetology Guide American Society of Ichthyologists and Herpetologists Herpetological Conservation and Biology Societas Europaea Herpetologica Distribution Maps for European Reptiles and Amphibians Center for North American Herpetology over 500 species of reptiles and amphibians European Field Herping Community New Zealand Herpetology Chicago Herpetological Society Biology of the Reptilia is an online copy of the full text of a 22-volume 13,000-page summary of the state of research of reptiles.
HerpMapper is a database of reptile and amphibian sightings Amphibian and Reptile Atlas of Peninsular California, San Diego Natural History Museum A Primer on Reptiles and Amphibians
Animals are multicellular eukaryotic organisms that form the biological kingdom Animalia. With few exceptions, animals consume organic material, breathe oxygen, are able to move, can reproduce sexually, grow from a hollow sphere of cells, the blastula, during embryonic development. Over 1.5 million living animal species have been described—of which around 1 million are insects—but it has been estimated there are over 7 million animal species in total. Animals range in length from 8.5 millionths of a metre to 33.6 metres and have complex interactions with each other and their environments, forming intricate food webs. The category includes humans, but in colloquial use the term animal refers only to non-human animals; the study of non-human animals is known as zoology. Most living animal species are in the Bilateria, a clade whose members have a bilaterally symmetric body plan; the Bilateria include the protostomes—in which many groups of invertebrates are found, such as nematodes and molluscs—and the deuterostomes, containing the echinoderms and chordates.
Life forms interpreted. Many modern animal phyla became established in the fossil record as marine species during the Cambrian explosion which began around 542 million years ago. 6,331 groups of genes common to all living animals have been identified. Aristotle divided animals into those with those without. Carl Linnaeus created the first hierarchical biological classification for animals in 1758 with his Systema Naturae, which Jean-Baptiste Lamarck expanded into 14 phyla by 1809. In 1874, Ernst Haeckel divided the animal kingdom into the multicellular Metazoa and the Protozoa, single-celled organisms no longer considered animals. In modern times, the biological classification of animals relies on advanced techniques, such as molecular phylogenetics, which are effective at demonstrating the evolutionary relationships between animal taxa. Humans make use of many other animal species for food, including meat and eggs. Dogs have been used in hunting, while many aquatic animals are hunted for sport.
Non-human animals have appeared in art from the earliest times and are featured in mythology and religion. The word "animal" comes from the Latin animalis, having soul or living being; the biological definition includes all members of the kingdom Animalia. In colloquial usage, as a consequence of anthropocentrism, the term animal is sometimes used nonscientifically to refer only to non-human animals. Animals have several characteristics. Animals are eukaryotic and multicellular, unlike bacteria, which are prokaryotic, unlike protists, which are eukaryotic but unicellular. Unlike plants and algae, which produce their own nutrients animals are heterotrophic, feeding on organic material and digesting it internally. With few exceptions, animals breathe oxygen and respire aerobically. All animals are motile during at least part of their life cycle, but some animals, such as sponges, corals and barnacles become sessile; the blastula is a stage in embryonic development, unique to most animals, allowing cells to be differentiated into specialised tissues and organs.
All animals are composed of cells, surrounded by a characteristic extracellular matrix composed of collagen and elastic glycoproteins. During development, the animal extracellular matrix forms a flexible framework upon which cells can move about and be reorganised, making the formation of complex structures possible; this may be calcified, forming structures such as shells and spicules. In contrast, the cells of other multicellular organisms are held in place by cell walls, so develop by progressive growth. Animal cells uniquely possess the cell junctions called tight junctions, gap junctions, desmosomes. With few exceptions—in particular, the sponges and placozoans—animal bodies are differentiated into tissues; these include muscles, which enable locomotion, nerve tissues, which transmit signals and coordinate the body. There is an internal digestive chamber with either one opening or two openings. Nearly all animals make use of some form of sexual reproduction, they produce haploid gametes by meiosis.
These fuse to form zygotes, which develop via mitosis into a hollow sphere, called a blastula. In sponges, blastula larvae swim to a new location, attach to the seabed, develop into a new sponge. In most other groups, the blastula undergoes more complicated rearrangement, it first invaginates to form a gastrula with a digestive chamber and two separate germ layers, an external ectoderm and an internal endoderm. In most cases, a third germ layer, the mesoderm develops between them; these germ layers differentiate to form tissues and organs. Repeated instances of mating with a close relative during sexual reproduction leads to inbreeding depression within a population due to the increased prevalence of harmful recessive traits. Animals have evolved numerous mechanisms for avoiding close inbreeding. In some species, such as the splendid fairywren, females benefit by mating with multiple males, thus producing more offspring of higher genetic quality; some animals are capable of asexual reproduction, which results