A planthopper is any insect in the infraorder Fulgoromorpha: in the suborder Auchenorrhyncha and exceeding 12,500 described species worldwide. The name comes from their remarkable resemblance to leaves and other plants of their environment and from the fact that they "hop" for quick transportation in a similar way to that of grasshoppers. However, planthoppers walk slowly so as not to attract attention. Distributed worldwide, all members of this group are plant-feeders, though few are considered pests; the infraorder contains Fulgoroidea. Fulgoroids are most reliably distinguished from the other Auchenorrhyncha by two features. Nymphs of many fulgoroids produce wax from special glands on the abdominal terga and other parts of the body; these help conceal the insects. Adult females of many families produce wax which may be used to protect eggs. Fulgoroid nymphs possess a biological gear mechanism at the base of the hind legs, which keeps the legs in synchrony when the insects jump; the gears, not present in the adults, were known for decades before the recent description of their function.
Planthoppers are vectors for plant diseases phytoplasmas which live in the phloem of plants and can be transmitted by planthoppers when feeding. A number of extinct member of Fulgoroidea are known from the fossil record, such as the Lutetian age Emiliana from the Green River Formation in Colorado, USA; as mentioned under Auchenorrhyncha, some authors use the name Archaeorrhyncha as a replacement for the Fulgoromorpha. The extant families of Fulgoroidea are: Acanaloniidae Achilidae Achilixiidae Caliscelidae Cixiidae Delphacidae Derbidae Dictyopharidae Eurybrachidae Flatidae Fulgoridae Gengidae Hypochthonellidae Issidae Kinnaridae Lophopidae Meenoplidae Nogodinidae Ricaniidae Tettigometridae Tropiduchidae Stephen W. Wilson Keys To The Families Of Fulgoromorpha with emphasis on planthoppers of potential economic importance in the southeastern United States. Florida Entomologist 88 PDF Bourgoin T. 1996-2015. FLOW: a world knowledge base dedicated to Fulgoromorpha. Http://www.hemiptera-databases.org/flow/ Larivière, M.-C..
J.. Fauna of New Zealand, Swzedo J.. In: Fossil Planthoppers of the world. An annotated catalogue with notes on Hemiptera classification. Swzedo, J. Th. Bourgoin & F. Lefèbvre. J. Swzedo edt. Warsaw 2004, 199 pp + 8 pl. Metcalfa pruinosa, citrus flatid planthopper on the University of Florida/IFAS Featured Creatures website Ormenaria rufifascia, a flatid planthopper on the University of Florida/Institute of Food and Agricultural Sciences Featured Creatures website www.planthopper.com
Animals are multicellular eukaryotic organisms that form the biological kingdom Animalia. With few exceptions, animals consume organic material, breathe oxygen, are able to move, can reproduce sexually, grow from a hollow sphere of cells, the blastula, during embryonic development. Over 1.5 million living animal species have been described—of which around 1 million are insects—but it has been estimated there are over 7 million animal species in total. Animals range in length from 8.5 millionths of a metre to 33.6 metres and have complex interactions with each other and their environments, forming intricate food webs. The category includes humans, but in colloquial use the term animal refers only to non-human animals; the study of non-human animals is known as zoology. Most living animal species are in the Bilateria, a clade whose members have a bilaterally symmetric body plan; the Bilateria include the protostomes—in which many groups of invertebrates are found, such as nematodes and molluscs—and the deuterostomes, containing the echinoderms and chordates.
Life forms interpreted. Many modern animal phyla became established in the fossil record as marine species during the Cambrian explosion which began around 542 million years ago. 6,331 groups of genes common to all living animals have been identified. Aristotle divided animals into those with those without. Carl Linnaeus created the first hierarchical biological classification for animals in 1758 with his Systema Naturae, which Jean-Baptiste Lamarck expanded into 14 phyla by 1809. In 1874, Ernst Haeckel divided the animal kingdom into the multicellular Metazoa and the Protozoa, single-celled organisms no longer considered animals. In modern times, the biological classification of animals relies on advanced techniques, such as molecular phylogenetics, which are effective at demonstrating the evolutionary relationships between animal taxa. Humans make use of many other animal species for food, including meat and eggs. Dogs have been used in hunting, while many aquatic animals are hunted for sport.
Non-human animals have appeared in art from the earliest times and are featured in mythology and religion. The word "animal" comes from the Latin animalis, having soul or living being; the biological definition includes all members of the kingdom Animalia. In colloquial usage, as a consequence of anthropocentrism, the term animal is sometimes used nonscientifically to refer only to non-human animals. Animals have several characteristics. Animals are eukaryotic and multicellular, unlike bacteria, which are prokaryotic, unlike protists, which are eukaryotic but unicellular. Unlike plants and algae, which produce their own nutrients animals are heterotrophic, feeding on organic material and digesting it internally. With few exceptions, animals breathe oxygen and respire aerobically. All animals are motile during at least part of their life cycle, but some animals, such as sponges, corals and barnacles become sessile; the blastula is a stage in embryonic development, unique to most animals, allowing cells to be differentiated into specialised tissues and organs.
All animals are composed of cells, surrounded by a characteristic extracellular matrix composed of collagen and elastic glycoproteins. During development, the animal extracellular matrix forms a flexible framework upon which cells can move about and be reorganised, making the formation of complex structures possible; this may be calcified, forming structures such as shells and spicules. In contrast, the cells of other multicellular organisms are held in place by cell walls, so develop by progressive growth. Animal cells uniquely possess the cell junctions called tight junctions, gap junctions, desmosomes. With few exceptions—in particular, the sponges and placozoans—animal bodies are differentiated into tissues; these include muscles, which enable locomotion, nerve tissues, which transmit signals and coordinate the body. There is an internal digestive chamber with either one opening or two openings. Nearly all animals make use of some form of sexual reproduction, they produce haploid gametes by meiosis.
These fuse to form zygotes, which develop via mitosis into a hollow sphere, called a blastula. In sponges, blastula larvae swim to a new location, attach to the seabed, develop into a new sponge. In most other groups, the blastula undergoes more complicated rearrangement, it first invaginates to form a gastrula with a digestive chamber and two separate germ layers, an external ectoderm and an internal endoderm. In most cases, a third germ layer, the mesoderm develops between them; these germ layers differentiate to form tissues and organs. Repeated instances of mating with a close relative during sexual reproduction leads to inbreeding depression within a population due to the increased prevalence of harmful recessive traits. Animals have evolved numerous mechanisms for avoiding close inbreeding. In some species, such as the splendid fairywren, females benefit by mating with multiple males, thus producing more offspring of higher genetic quality; some animals are capable of asexual reproduction, which results
In zoological nomenclature, a type species is the species name with which the name of a genus or subgenus is considered to be permanently taxonomically associated, i.e. the species that contains the biological type specimen. A similar concept is used for suprageneric groups called a type genus. In botanical nomenclature, these terms have no formal standing under the code of nomenclature, but are sometimes borrowed from zoological nomenclature. In botany, the type of a genus name is a specimen, the type of a species name; the species name that has that type can be referred to as the type of the genus name. Names of genus and family ranks, the various subdivisions of those ranks, some higher-rank names based on genus names, have such types. In bacteriology, a type species is assigned for each genus; every named genus or subgenus in zoology, whether or not recognized as valid, is theoretically associated with a type species. In practice, there is a backlog of untypified names defined in older publications when it was not required to specify a type.
A type species is both a concept and a practical system, used in the classification and nomenclature of animals. The "type species" represents the reference species and thus "definition" for a particular genus name. Whenever a taxon containing multiple species must be divided into more than one genus, the type species automatically assigns the name of the original taxon to one of the resulting new taxa, the one that includes the type species; the term "type species" is regulated in zoological nomenclature by article 42.3 of the International Code of Zoological Nomenclature, which defines a type species as the name-bearing type of the name of a genus or subgenus. In the Glossary, type species is defined as The nominal species, the name-bearing type of a nominal genus or subgenus; the type species permanently attaches a formal name to a genus by providing just one species within that genus to which the genus name is permanently linked. The species name in turn is fixed, to a type specimen. For example, the type species for the land snail genus Monacha is Helix cartusiana, the name under which the species was first described, known as Monacha cartusiana when placed in the genus Monacha.
That genus is placed within the family Hygromiidae. The type genus for that family is the genus Hygromia; the concept of the type species in zoology was introduced by Pierre André Latreille. The International Code of Zoological Nomenclature states that the original name of the type species should always be cited, it gives an example in Article 67.1. Astacus marinus Fabricius, 1775 was designated as the type species of the genus Homarus, thus giving it the name Homarus marinus. However, the type species of Homarus should always be cited using its original name, i.e. Astacus marinus Fabricius, 1775. Although the International Code of Nomenclature for algae and plants does not contain the same explicit statement, examples make it clear that the original name is used, so that the "type species" of a genus name need not have a name within that genus, thus in Article 10, Ex. 3, the type of the genus name Elodes is quoted as the type of the species name Hypericum aegypticum, not as the type of the species name Elodes aegyptica.
Glossary of scientific naming Genetypes – genetic sequence data from type specimens. Holotype Paratype Principle of Typification Type Type genus
An arthropod is an invertebrate animal having an exoskeleton, a segmented body, paired jointed appendages. Arthropods form the phylum Euarthropoda, which includes insects, arachnids and crustaceans; the term Arthropoda as proposed refers to a proposed grouping of Euarthropods and the phylum Onychophora. Arthropods are characterized by their jointed limbs and cuticle made of chitin mineralised with calcium carbonate; the arthropod body plan consists of each with a pair of appendages. The rigid cuticle inhibits growth, so arthropods replace it periodically by moulting. Arthopods are bilaterally symmetrical and their body possesses an external skeleton; some species have wings. Their versatility has enabled them to become the most species-rich members of all ecological guilds in most environments, they have over a million described species, making up more than 80 per cent of all described living animal species, some of which, unlike most other animals, are successful in dry environments. Arthropods range in size from the microscopic crustacean Stygotantulus up to the Japanese spider crab.
Arthropods' primary internal cavity is a haemocoel, which accommodates their internal organs, through which their haemolymph – analogue of blood – circulates. Like their exteriors, the internal organs of arthropods are built of repeated segments, their nervous system is "ladder-like", with paired ventral nerve cords running through all segments and forming paired ganglia in each segment. Their heads are formed by fusion of varying numbers of segments, their brains are formed by fusion of the ganglia of these segments and encircle the esophagus; the respiratory and excretory systems of arthropods vary, depending as much on their environment as on the subphylum to which they belong. Their vision relies on various combinations of compound eyes and pigment-pit ocelli: in most species the ocelli can only detect the direction from which light is coming, the compound eyes are the main source of information, but the main eyes of spiders are ocelli that can form images and, in a few cases, can swivel to track prey.
Arthropods have a wide range of chemical and mechanical sensors based on modifications of the many setae that project through their cuticles. Arthropods' methods of reproduction and development are diverse; the evolutionary ancestry of arthropods dates back to the Cambrian period. The group is regarded as monophyletic, many analyses support the placement of arthropods with cycloneuralians in a superphylum Ecdysozoa. Overall, the basal relationships of Metazoa are not yet well resolved; the relationships between various arthropod groups are still debated. Aquatic species use either external fertilization. All arthropods lay eggs, but scorpions give birth to live young after the eggs have hatched inside the mother. Arthropod hatchlings vary from miniature adults to grubs and caterpillars that lack jointed limbs and undergo a total metamorphosis to produce the adult form; the level of maternal care for hatchlings varies from nonexistent to the prolonged care provided by scorpions. Arthropods contribute to the human food supply both directly as food, more indirectly as pollinators of crops.
Some species are known to spread severe disease to humans and crops. The word arthropod comes from the Greek ἄρθρον árthron, "joint", πούς pous, i.e. "foot" or "leg", which together mean "jointed leg". Arthropods are invertebrates with jointed limbs; the exoskeleton or cuticles consists of a polymer of glucosamine. The cuticle of many crustaceans, beetle mites, millipedes is biomineralized with calcium carbonate. Calcification of the endosternite, an internal structure used for muscle attachments occur in some opiliones. Estimates of the number of arthropod species vary between 1,170,000 and 5 to 10 million and account for over 80 per cent of all known living animal species; the number of species remains difficult to determine. This is due to the census modeling assumptions projected onto other regions in order to scale up from counts at specific locations applied to the whole world. A study in 1992 estimated that there were 500,000 species of animals and plants in Costa Rica alone, of which 365,000 were arthropods.
They are important members of marine, freshwater and air ecosystems, are one of only two major animal groups that have adapted to life in dry environments. One arthropod sub-group, insects, is the most species-rich member of all ecological guilds in land and freshwater environments; the lightest insects weigh less than 25 micrograms. Some living crustaceans are much larger; the embryos of all arthropods are segmented, built from a series of repeated modules. The last common ancestor of living arthropods consisted of a series of undifferentiated segments, each with a pair of appendages that functioned as limbs. However, all known living and fossil arthropods have grouped segments into tagmata in which segments and their limbs are specialized in various ways; the three-
The Fulgorinae are a sub-family of insects in the Auchenorrhyncha: which include the spectacular "lantern-bugs" and allied insects. Nine genera are listed by the NCBI, but molecular studies question the placement of genera in the Zannini questioning whether they belong to the family Fulgoridae; the Hemiptera database suggests that there may be 15 genera in this subfamily in three tribes: Auth.: Latreille, 1807 Aphrodisias Kirkaldy, 1906 Cathedra Kirkaldy, 1903 monotypic: Cathedra serrata Diareusa Walker, 1858 Fulgora Linné, 1767 Odontoptera Carreno, 1841 Phrictus Spinola, 1839 Auth.: Distant, 1906 Datua Schmidt, 1911 Hariola Stål, 1863 Saiva Distant, 1906 Auth.: Metcalf, 1938 Pseudodictya Schmidt, 1924 Radamana Distant, 1906 Rhicnophloea Gerstaecker, 1895 Ruehlella Schmidt, 1924 Samsama Distant, 1906 Zanna Kirkaldy, 1902 Placed here by NCBI Pyrops Spinola, 1839 Sinuala O'Brien, 1991 Data related to Fulgoridae at Wikispecies Media related to Fulgoridae at Wikimedia Commons
Shaivism is one of the major traditions within Hinduism that reveres Shiva as the Supreme Being. The followers of Shaivism are called "Shaivites" or "Saivites", it is one of the largest sects that believe Shiva — worshipped as a creator and destroyer of worlds — is the supreme god over all. The Shaiva have many sub-traditions, ranging from devotional dualistic theism such as Shaiva Siddhanta to yoga-oriented monistic non-theism such as Kashmiri Shaivism, it considers the Agama texts as important sources of theology. The origin of Shaivism may be traced to the conception of Rudra in the Rig Veda. Shaivism has ancient roots, traceable in the Vedic literature of 2nd millennium BCE, but this is in the form of the Vedic deity Rudra; the ancient text Shvetashvatara Upanishad dated to late 1st millennium BCE mentions terms such as Rudra and Maheshwaram, but its interpretation as a theistic or monistic text of Shaivism is disputed. In the early centuries of the common era is the first clear evidence of Pāśupata Shaivism.
Both devotional and monistic Shaivism became popular in the 1st millennium CE becoming the dominant religious tradition of many Hindu kingdoms. It arrived in Southeast Asia shortly thereafter, leading to thousands of Shaiva temples on the islands of Indonesia as well as Cambodia and Vietnam, co-evolving with Buddhism in these regions. In the contemporary era, Shaivism is one of the major aspects of Hinduism. Shaivism theology ranges from Shiva being the creator, destroyer to being the same as the Atman within oneself and every living being, it is related to Shaktism, some Shaiva worship in Shiva and Shakti temples. It is the Hindu tradition that most accepts ascetic life and emphasizes yoga, like other Hindu traditions encourages an individual to discover and be one with Shiva within. Shaivism is one of the largest traditions within Hinduism. Shiva means kind, gracious, or auspicious; as a proper name, it means "The Auspicious One". The word Shiva is used as an adjective in the Rig Veda, as an epithet for several Rigvedic deities, including Rudra.
The term Shiva connotes "liberation, final emancipation" and "the auspicious one", this adjective sense of usage is addressed to many deities in Vedic layers of literature. The term evolved from the Vedic Rudra-Shiva to the noun Shiva in the Epics and the Puranas, as an auspicious deity, the "creator and dissolver"; the Sanskrit word śaiva or Shaiva means "relating to the god Shiva", while the related beliefs, history and sub-traditions constitute Shaivism. The reverence for Shiva is one of the pan-Hindu traditions, found across India, Sri Lanka and Nepal. While Shiva is revered broadly, Hinduism itself is a complex religion and a way of life, with a diversity of ideas on spirituality and traditions, it has no ecclesiastical order, no unquestionable religious authorities, no governing body, no prophet nor any binding holy book. Shaivism is a major tradition within Hinduism, with a theology, predominantly related to the Hindu god Shiva. Shaivism has many different sub-traditions with regional differences in philosophy.
Shaivism has a vast literature with different philosophical schools, ranging from nondualism and mixed schools. The origins of Shaivism a matter of debate among scholars; some trace the origins to the Indus Valley civilization, which reached its peak around 2500–2000 BCE. Archeological discoveries show seals. Of these is the Pashupati seal, which early scholars interpreted as someone seated in a meditating yoga pose surrounded by animals, with horns; this "Pashupati" seal has been interpreted by these scholars as a prototype of Shiva. Gavin Flood characterizes these views as "speculative", saying that it is not clear from the seal if the figure has three faces, or is seated in a yoga posture, or that the shape is intended to represent a human figure. Other scholars state that the Indus Valley script remains undeciphered, the interpretation of the Pashupati seal is uncertain. According to Srinivasan, the proposal that it is proto-Shiva may be a case of projecting "later practices into archeological findings".
Asko Parpola states that other archaeological finds such as the early Elamite seals dated to 3000–2750 BCE show similar figures and these have been interpreted as "seated bull" and not a yogi, the bull interpretation is more accurate. The Rigveda has the earliest clear mention of Rudra in its hymns such as 2.33, 1.43 and 1.114. The text includes a Satarudriya, an influential hymn with embedded hundred epithets for Rudra, cited in many medieval era Shaiva texts as well as recited in major Shiva temples of Hindus in contemporary times. Yet, the Vedic literature only present scriptural theology, but does not attest to the existence of Shaivism; the Shvetashvatara Upanishad composed before the Bhagavad Gita about 4th century BCE contains the theistic foundations of Shaivism wrapped in a monistic structure. It contains the key terms and ideas of Shaivism, such as Shiva, Maheswara, Bhakti, Atman and self-knowledge. According to Gavin Flood, "the formation of Śaiva traditions as we understand them begins to occur during the period from 200 BC to 100 AD."
According to Chakravarti, Shiva rose to prominence as he was identified to be the
Insects or Insecta are hexapod invertebrates and the largest group within the arthropod phylum. Definitions and circumscriptions vary; as used here, the term Insecta is synonymous with Ectognatha. Insects have a chitinous exoskeleton, a three-part body, three pairs of jointed legs, compound eyes and one pair of antennae. Insects are the most diverse group of animals; the total number of extant species is estimated at between ten million. Insects may be found in nearly all environments, although only a small number of species reside in the oceans, which are dominated by another arthropod group, crustaceans. Nearly all insects hatch from eggs. Insect growth is constrained by the inelastic exoskeleton and development involves a series of molts; the immature stages differ from the adults in structure and habitat, can include a passive pupal stage in those groups that undergo four-stage metamorphosis. Insects that undergo three-stage metamorphosis lack a pupal stage and adults develop through a series of nymphal stages.
The higher level relationship of the insects is unclear. Fossilized insects of enormous size have been found from the Paleozoic Era, including giant dragonflies with wingspans of 55 to 70 cm; the most diverse insect groups appear to have coevolved with flowering plants. Adult insects move about by walking, flying, or sometimes swimming; as it allows for rapid yet stable movement, many insects adopt a tripedal gait in which they walk with their legs touching the ground in alternating triangles, composed of the front & rear on one side with the middle on the other side. Insects are the only invertebrates to have evolved flight, all flying insects derive from one common ancestor. Many insects spend at least part of their lives under water, with larval adaptations that include gills, some adult insects are aquatic and have adaptations for swimming; some species, such as water striders, are capable of walking on the surface of water. Insects are solitary, but some, such as certain bees and termites, are social and live in large, well-organized colonies.
Some insects, such as earwigs, show maternal care, guarding their eggs and young. Insects can communicate with each other in a variety of ways. Male moths can sense the pheromones of female moths over great distances. Other species communicate with sounds: crickets stridulate, or rub their wings together, to attract a mate and repel other males. Lampyrid beetles communicate with light. Humans regard certain insects as pests, attempt to control them using insecticides, a host of other techniques; some insects damage crops by feeding on sap, fruits, or wood. Some species are parasitic, may vector diseases; some insects perform complex ecological roles. Insect pollinators are essential to the life cycle of many flowering plant species on which most organisms, including humans, are at least dependent. Many insects are considered ecologically beneficial as predators and a few provide direct economic benefit. Silkworms produce silk and honey bees produce honey and both have been domesticated by humans.
Insects are consumed as food in 80% of the world's nations, by people in 3000 ethnic groups. Human activities have effects on insect biodiversity; the word "insect" comes from the Latin word insectum, meaning "with a notched or divided body", or "cut into", from the neuter singular perfect passive participle of insectare, "to cut into, to cut up", from in- "into" and secare "to cut". A calque of Greek ἔντομον, "cut into sections", Pliny the Elder introduced the Latin designation as a loan-translation of the Greek word ἔντομος or "insect", Aristotle's term for this class of life in reference to their "notched" bodies. "Insect" first appears documented in English in 1601 in Holland's translation of Pliny. Translations of Aristotle's term form the usual word for "insect" in Welsh, Serbo-Croatian, etc; the precise definition of the taxon Insecta and the equivalent English name "insect" varies. In the broadest circumscription, Insecta sensu lato consists of all hexapods. Traditionally, insects defined in this way were divided into "Apterygota" —the wingless insects—and Pterygota—the winged insects.
However, modern phylogenetic studies have shown that "Apterygota" is not monophyletic, so does not form a good taxon. A narrower circumscription restricts insects to those hexapods with external mouthparts, comprises only the last three groups in the table. In this sense, Insecta sensu stricto is equivalent to Ectognatha. In the narrowest circumscription, insects are restricted to hexapods that are either winged or descended from winged ancestors. Insecta sensu strictissimo is equivalent to Pterygota. For the purposes of this article, the middle definition is used; the evolutionary relationship of insects to other animal groups remains unclear. Although traditionally grouped with millipedes and centiped