A leaf is an organ of a vascular plant and is the principal lateral appendage of the stem. The leaves and stem together form the shoot. Leaves are collectively referred to as foliage, as in "autumn foliage". A leaf is a thin, dorsiventrally flattened organ borne above ground and specialized for photosynthesis. In most leaves, the primary photosynthetic tissue, the palisade mesophyll, is located on the upper side of the blade or lamina of the leaf but in some species, including the mature foliage of Eucalyptus, palisade mesophyll is present on both sides and the leaves are said to be isobilateral. Most leaves have distinct upper surface and lower surface that differ in colour, the number of stomata, the amount and structure of epicuticular wax and other features. Leaves can have many different shapes and textures; the broad, flat leaves with complex venation of flowering plants are known as megaphylls and the species that bear them, the majority, as broad-leaved or megaphyllous plants. In the clubmosses, with different evolutionary origins, the leaves are simple and are known as microphylls.
Some leaves, such as bulb scales, are not above ground. In many aquatic species the leaves are submerged in water. Succulent plants have thick juicy leaves, but some leaves are without major photosynthetic function and may be dead at maturity, as in some cataphylls and spines. Furthermore, several kinds of leaf-like structures found in vascular plants are not homologous with them. Examples include flattened plant stems called phylloclades and cladodes, flattened leaf stems called phyllodes which differ from leaves both in their structure and origin; some structures of non-vascular plants function much like leaves. Examples include the phyllids of liverworts. Leaves are the most important organs of most vascular plants. Green plants are autotrophic, meaning that they do not obtain food from other living things but instead create their own food by photosynthesis, they capture the energy in sunlight and use it to make simple sugars, such as glucose and sucrose, from carbon dioxide and water. The sugars are stored as starch, further processed by chemical synthesis into more complex organic molecules such as proteins or cellulose, the basic structural material in plant cell walls, or metabolised by cellular respiration to provide chemical energy to run cellular processes.
The leaves draw water from the ground in the transpiration stream through a vascular conducting system known as xylem and obtain carbon dioxide from the atmosphere by diffusion through openings called stomata in the outer covering layer of the leaf, while leaves are orientated to maximise their exposure to sunlight. Once sugar has been synthesized, it needs to be transported to areas of active growth such as the plant shoots and roots. Vascular plants transport sucrose in a special tissue called the phloem; the phloem and xylem are parallel to each other but the transport of materials is in opposite directions. Within the leaf these vascular systems branch to form veins which supply as much of the leaf as possible, ensuring that cells carrying out photosynthesis are close to the transportation system. Leaves are broad and thin, thereby maximising the surface area directly exposed to light and enabling the light to penetrate the tissues and reach the chloroplasts, thus promoting photosynthesis.
They are arranged on the plant so as to expose their surfaces to light as efficiently as possible without shading each other, but there are many exceptions and complications. For instance plants adapted to windy conditions may have pendent leaves, such as in many willows and eucalyptss; the flat, or laminar, shape maximises thermal contact with the surrounding air, promoting cooling. Functionally, in addition to carrying out photosynthesis, the leaf is the principal site of transpiration, providing the energy required to draw the transpiration stream up from the roots, guttation. Many gymnosperms have thin needle-like or scale-like leaves that can be advantageous in cold climates with frequent snow and frost; these are interpreted as reduced from megaphyllous leaves of their Devonian ancestors. Some leaf forms are adapted to modulate the amount of light they absorb to avoid or mitigate excessive heat, ultraviolet damage, or desiccation, or to sacrifice light-absorption efficiency in favour of protection from herbivory.
For xerophytes the major constraint drought. Some window plants such as Fenestraria species and some Haworthia species such as Haworthia tesselata and Haworthia truncata are examples of xerophytes. and Bulbine mesembryanthemoides. Leaves function to store chemical energy and water and may become specialised organs serving other functions, such as tendrils of peas and other legumes, the protective spines of cacti and the insect traps in carnivorous plants such as Nepenthes and Sarracenia. Leaves are the fundamental structural units from which cones are constructed in gymnosperms and from which flowers are constructed in flowering plants; the internal organisation of most kinds of leaves has evolved to maximise exposure of the photosynthetic organelles, the chloroplasts, to light and to increase the absorption of carbon dioxide while at the same time controlling water loss. Their surfaces are waterproofed by the plant cuticle and gas exchange between the mesophyll cells and the atmosphere is controlled by minute openings called stomata which open or close to regulate the rate exchange of carbon dioxide and water vapour into
The term incubous describes the way in which the leaves of a liverwort are attached to the stem. If one were to look down from above on a plant where the leaf attachment is incubous, the upper edge of each leaf would overlap the next higher leaf along the stem; because of this, the upper edge of each leaf is visible from above, but the lower edge of each leaf is obscured by its neighboring leaf. The opposite of incubous is succubous; the dictionary definition of incubous at Wiktionary
The Marchantiophyta are a division of non-vascular land plants referred to as hepatics or liverworts. Like mosses and hornworts, they have a gametophyte-dominant life cycle, in which cells of the plant carry only a single set of genetic information, it is estimated. Some of the more familiar species grow as a flattened leafless thallus, but most species are leafy with a form much like a flattened moss. Leafy species can be distinguished from the similar mosses on the basis of a number of features, including their single-celled rhizoids. Leafy liverworts differ from most mosses in that their leaves never have a costa and may bear marginal cilia. Other differences are not universal for all mosses and liverworts, but the occurrence of leaves arranged in three ranks, the presence of deep lobes or segmented leaves, or a lack of differentiated stem and leaves all point to the plant being a liverwort. Liverworts are small from 2–20 mm wide with individual plants less than 10 cm long, are therefore overlooked.
However, certain species may cover large patches of ground, trees or any other reasonably firm substrate on which they occur. They are distributed globally in every available habitat, most in humid locations although there are desert and Arctic species as well; some species can be a weed in gardens. Most liverworts are small, measuring from 2–20 millimetres wide with individual plants less than 10 centimetres long, so they are overlooked; the most familiar liverworts consist of a prostrate, ribbon-like or branching structure called a thallus. However, most liverworts produce flattened stems with overlapping scales or leaves in two or more ranks, the middle rank is conspicuously different from the outer ranks. Liverworts can most reliably be distinguished from the similar mosses by their single-celled rhizoids. Other differences are not universal for all mosses and all liverworts. Unlike any other embryophytes, most liverworts contain unique membrane-bound oil bodies containing isoprenoids in at least some of their cells, lipid droplets in the cytoplasm of all other plants being unenclosed.
The overall physical similarity of some mosses and leafy liverworts means that confirmation of the identification of some groups can be performed with certainty only with the aid of microscopy or an experienced bryologist. Liverworts have a gametophyte-dominant life cycle, with the sporophyte dependent on the gametophyte. Cells in a typical liverwort plant each contain only a single set of genetic information, so the plant's cells are haploid for the majority of its life cycle; this contrasts with the pattern exhibited by nearly all animals and by most other plants. In the more familiar seed plants, the haploid generation is represented only by the tiny pollen and the ovule, while the diploid generation is the familiar tree or other plant. Another unusual feature of the liverwort life cycle is that sporophytes are short-lived, withering away not long after releasing spores. In other bryophytes, the sporophyte is persistent and disperses spores over an extended period; the life of a liverwort starts from the germination of a haploid spore to produce a protonema, either a mass of thread-like filaments or else a flattened thallus.
The protonema is a transitory stage in the life of a liverwort, from which will grow the mature gametophore plant that produces the sex organs. The male organs produce the sperm cells. Clusters of antheridia are enclosed by a protective layer of cells called the perigonium; as in other land plants, the female organs are known as archegonia and are protected by the thin surrounding perichaetum. Each archegonium has a slender hollow tube, the "neck", down which the sperm swim to reach the egg cell. Liverwort species may be either monoicous. In dioicous liverworts and male sex organs are borne on different and separate gametophyte plants. In monoicous liverworts, the two kinds of reproductive structures are borne on different branches of the same plant. In either case, the sperm must move from the antheridia where they are produced to the archegonium where the eggs are held; the sperm of liverworts is biflagellate, i.e. they have two tail-like flagellae that enable them to swim short distances, provided that at least a thin film of water is present.
Their journey may be assisted by the splashing of raindrops. In 2008, Japanese researchers discovered that some liverworts are able to fire sperm-containing water up to 15 cm in the air, enabling them to fertilize female plants growing more than a metre from the nearest male; when sperm reach the archegonia, fertilisation occurs, leading to the production of a diploid sporophyte. After fertilisation, the immature sporophyte within the archegonium develops three distinct regions: a foot, which both anchors the sporophyte in place and receives nutrients from its "mother" plant, a spherical or ellipsoidal capsule, inside which the spores will be produced for dispersing to new locations, a seta which lies between the other two
A stem is one of two main structural axes of a vascular plant, the other being the root. The stem is divided into nodes and internodes: The nodes hold one or more leaves, as well as buds which can grow into branches. Adventitious roots may be produced from the nodes; the internodes distance one node from another. The term "shoots" is confused with "stems". In most plants stems are located above the soil surface but some plants have underground stems. Stems have four main functions which are: Support for and the elevation of leaves and fruits; the stems keep the leaves in the light and provide a place for the plant to keep its flowers and fruits. Transport of fluids between the roots and the shoots in the xylem and phloem Storage of nutrients Production of new living tissue; the normal lifespan of plant cells is one to three years. Stems have cells called meristems. Stems are specialized for storage, asexual reproduction, protection or photosynthesis, including the following: Acaulescent – used to describe stems in plants that appear to be stemless.
These stems are just short, the leaves appearing to rise directly out of the ground, e.g. some Viola species. Arborescent – tree like with woody stems with a single trunk. Axillary bud – a bud which grows at the point of attachment of an older leaf with the stem, it gives rise to a shoot. Branched – aerial stems are described as being branched or unbranched Bud – an embryonic shoot with immature stem tip. Bulb – a short vertical underground stem with fleshy storage leaves attached, e.g. onion, tulip. Bulbs function in reproduction by splitting to form new bulbs or producing small new bulbs termed bulblets. Bulbs are a combination of stem and leaves so may better be considered as leaves because the leaves make up the greater part. Caespitose – when stems grow in a tangled mass or clump or in low growing mats. Cladode – a flattened stem that appears more-or-less leaf like and is specialized for photosynthesis, e.g. cactus pads. Climbing -- stems that wrap around other plants or structures. Corm – a short enlarged underground, storage stem, e.g. taro, gladiolus.
Decumbent -- stems that lie flat on the turn upwards at the ends. Fruticose -- stems. Herbaceous – non woody, they die at the end of the growing season. Internode – an interval between two successive nodes, it possesses the ability to elongate, either from its base or from its extremity depending on the species. Node – a point of attachment of a leaf or a twig on the stem in seed plants. A node is a small growth zone. Pedicel – stems that serve as the stalk of an individual flower in an inflorescence or infrutescence. Peduncle – a stem that supports an inflorescence Prickle – a sharpened extension of the stem's outer layers, e.g. roses. Pseudostem – a false stem made of the rolled bases of leaves, which may be 2 or 3 m tall as in banana Rhizome – a horizontal underground stem that functions in reproduction but in storage, e.g. most ferns, iris Runner – a type of stolon, horizontally growing on top of the ground and rooting at the nodes, aids in reproduction. E.g. garden strawberry, Chlorophytum comosum.
Scape – a stem that holds flowers that comes out of the ground and has no normal leaves. Hosta, Iris, Garlic. Stolon – a horizontal stem that produces rooted plantlets at its nodes and ends, forming near the surface of the ground. Thorn – a modified stem with a sharpened point. Tuber – a swollen, underground storage stem adapted for storage and reproduction, e.g. potato. Woody – hard textured stems with secondary xylem. Stem consist of three tissues, dermal tissue, ground tissue and vascular tissue; the dermal tissue covers the outer surface of the stem and functions to waterproof and control gas exchange. The ground tissue consists of parenchyma cells and fills in around the vascular tissue, it sometimes functions in photosynthesis. Vascular tissue provides structural support. Most or all ground tissue may be lost in woody stems; the dermal tissue of aquatic plants stems. The arrangement of the vascular tissues varies among plant species. Dicot stems with primary growth have pith in the center, with vascular bundles forming a distinct ring visible when the stem is viewed in cross section.
The outside of the stem is covered with an epidermis, covered by a waterproof cuticle. The epidermis may contain stomata for gas exchange and multicellular stem hairs called trichomes. A cortex consisting of hypodermis and endodermis is present above the pericycle and vascular bundles. Woody dicots and many nonwoody dicots have secondary growth originating from their lateral or secondary meristems: the vascular cambium and the cork cambium or phellogen; the vascular cambium forms between the xylem and phloem in the vascular bundles and connects to form a continuous cylinder. The vascular cambium cells divide to produce secondary xylem to the inside and secondary phloem to the outside; as the stem increases in diameter due to production of secondary xylem and secondary phloem, the cortex and epidermis are destroyed. Before the cortex is destroyed, a cork cambium develops there; the cork cambium divides to produce waterproof cork cells externally and sometimes phelloderm cells internally. Those three tissues form the periderm.
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Anatomical terms of location
Standard anatomical terms of location deal unambiguously with the anatomy of animals, including humans. All vertebrates have the same basic body plan – they are bilaterally symmetrical in early embryonic stages and bilaterally symmetrical in adulthood; that is, they have mirror-image left and right halves if divided down the middle. For these reasons, the basic directional terms can be considered to be those used in vertebrates. By extension, the same terms are used for many other organisms as well. While these terms are standardized within specific fields of biology, there are unavoidable, sometimes dramatic, differences between some disciplines. For example, differences in terminology remain a problem that, to some extent, still separates the terminology of human anatomy from that used in the study of various other zoological categories. Standardized anatomical and zoological terms of location have been developed based on Latin and Greek words, to enable all biological and medical scientists to delineate and communicate information about animal bodies and their component organs though the meaning of some of the terms is context-sensitive.
The vertebrates and Craniata share a substantial heritage and common structure, so many of the same terms are used for location. To avoid ambiguities this terminology is based on the anatomy of each animal in a standard way. For humans, one type of vertebrate, anatomical terms may differ from other forms of vertebrates. For one reason, this is because humans have a different neuraxis and, unlike animals that rest on four limbs, humans are considered when describing anatomy as being in the standard anatomical position, thus what is on "top" of a human is the head, whereas the "top" of a dog may be its back, the "top" of a flounder could refer to either its left or its right side. For invertebrates, standard application of locational terminology becomes difficult or debatable at best when the differences in morphology are so radical that common concepts are not homologous and do not refer to common concepts. For example, many species are not bilaterally symmetrical. In these species, terminology depends on their type of symmetry.
Because animals can change orientation with respect to their environment, because appendages like limbs and tentacles can change position with respect to the main body, positional descriptive terms need to refer to the animal as in its standard anatomical position. All descriptions are with respect to the organism in its standard anatomical position when the organism in question has appendages in another position; this helps avoid confusion in terminology. In humans, this refers to the body in a standing position with arms at the side and palms facing forward. While the universal vertebrate terminology used in veterinary medicine would work in human medicine, the human terms are thought to be too well established to be worth changing. Many anatomical terms can be combined, either to indicate a position in two axes or to indicate the direction of a movement relative to the body. For example, "anterolateral" indicates a position, both anterior and lateral to the body axis. In radiology, an X-ray image may be said to be "anteroposterior", indicating that the beam of X-rays pass from their source to patient's anterior body wall through the body to exit through posterior body wall.
There is no definite limit to the contexts in which terms may be modified to qualify each other in such combinations. The modifier term is truncated and an "o" or an "i" is added in prefixing it to the qualified term. For example, a view of an animal from an aspect at once dorsal and lateral might be called a "dorsolateral" view. Again, in describing the morphology of an organ or habitus of an animal such as many of the Platyhelminthes, one might speak of it as "dorsiventrally" flattened as opposed to bilaterally flattened animals such as ocean sunfish. Where desirable three or more terms may be agglutinated or concatenated, as in "anteriodorsolateral"; such terms sometimes used to be hyphenated. There is however little basis for any strict rule to interfere with choice of convenience in such usage. Three basic reference planes are used to describe location; the sagittal plane is a plane parallel to the sagittal suture. All other sagittal planes are parallel to it, it is known as a "longitudinal plane".
The plane is perpendicular to the ground. The median plane or midsagittal plane is in the midline of the body, divides the body into left and right portions; this passes through the head, spinal cord, and, in many animals, the tail. The term "median plane" can refer to the midsagittal plane of other structures, such as a digit; the frontal plane or coronal plane divides the body into ventral portions. For post-embryonic humans a coronal plane is vertical and a transverse plane is horizontal, but for embryos and quadrupeds a coronal plane is horizontal and a transverse plane is vertical. A longitudinal plane is any plane perpendicular to the transverse plane; the coronal plane and the sagittal plane are examples of longitudinal planes. A transverse plane known as a cross-section, divides the body into cranial and caudal portions. In human anatomy: A transverse plane is an X-Z plane, parallel to the ground, which s
A succubus is a demon in female form, or supernatural entity in folklore, that appears in dreams and takes the form of a woman in order to seduce men through sexual activity. The male counterpart is the incubus. Religious traditions hold that repeated sexual activity with a succubus may result in the deterioration of health or mental state, or death. In modern representations, a succubus may or may not appear in dreams and is depicted as a attractive seductress or enchantress; the word is derived from Late Latin succuba "paramour". The word "succubus" originates from the late 14th century; as depicted in the Jewish mystical work Zohar and the medieval rabbinical text Alphabet of Ben Sira, Lilith was Adam's first wife, who became a succubus. She refused to return to the Garden of Eden after she mated with the archangel Samael. In Zoharistic Kabbalah, there were four succubi. There were four original queens of the demons: Lilith, Agrat bat Mahlat, Naamah. A succubus may take a form of a beautiful young girl but closer inspection may reveal deformities of her body, such as bird-like claws or serpentine tails.
Folklore describes the act of sexually penetrating a succubus as akin to entering a cavern of ice, there are reports of succubi forcing men to perform cunnilingus on their vulvas, which drip with urine and other fluids. In folklore, a succubus took the form of a siren. Throughout history and rabbis, including Hanina Ben Dosa and Abaye, tried to curb the power of succubi over humans. However, not all succubi were malevolent. According to Walter Map in the satire De Nugis Curialium, Pope Sylvester II was involved with a succubus named Meridiana, who helped him achieve his high rank in the Catholic Church. Before his death, he died repentant. According to the Kabbalah and the school of Rashba, the original three queens of the demons, Agrat Bat Mahlat, Eisheth Zenunim, all their cohorts give birth to children, except Lilith. According to other legends, the children of Lilith are called Lilin. According to the Malleus Maleficarum, or "Witches' Hammer", written by Heinrich Kramer in 1486, succubi collect semen from men they seduce.
Incubi, or male demons use the semen to impregnate human females, thus explaining how demons could sire children despite the traditional belief that they were incapable of reproduction. Children so begotten – cambions – were supposed to be those that were born deformed, or more susceptible to supernatural influences. While the book does not address why a human female impregnated with the semen of a human male would not produce regular human offspring, an explanation could be that the semen is altered before being transferred to the female host; however in some lore, the child is born deformed. King James in his dissertation titled Dæmonologie refutes the possibility for angelic entities to reproduce and instead offered a suggestion that a devil would carry out two methods of impregnating women: the first, to steal the sperm out of a dead man and deliver it into a woman. If a demon could extract the semen the transportation of the substance could not be transported to a female host, causing it to go cold.
This explains his view that succubae and incubi were the same demonic entity only to be described differently based on the tormented sexes being conversed with. The second method was the idea that a dead body could be possessed by a devil, causing it to rise and have sexual relations with others. However, there is no mention of a female corpse being possessed to elicit sex from men. In Arabian mythology, the qarînah is a spirit similar to the succubus, with origins in ancient Egyptian religion or in the animistic beliefs of pre-Islamic Arabia. A qarînah "sleeps with the person and has relations during sleep as is known by the dreams." They are said to be invisible, but a person with "second sight" can see them in the form of a cat, dog, or other household pet. "In Omdurman it is a spirit which possesses.... Only certain people are possessed and such people cannot marry or the qarina will harm them." To date, many African myths claim that men who have similar experience with such principality in dreams find themselves exhausted as soon as they awaken.
Local rituals/divination are invoked in order to appeal the god for divine protection and intervention. In the field of medicine, there is some belief that the stories relating to encounters with succubi bear resemblance to the contemporary phenomenon of people reporting alien abductions, ascribed to the condition known as sleep paralysis, it is therefore suggested that historical accounts of people experiencing encounters with succubi may rather have been symptoms of sleep paralysis, with the hallucination of the said creatures coming from their contemporary culture. Throughout history, succubi have been popular characters in music, film and more in pornography and manga/anime characters. Similar creatures in folklore "The Wiki of the Succubi - SuccuWiki". Www.succubus.net. Retrieved 6 November 2016