Squirrels are members of the family Sciuridae, a family that includes small or medium-size rodents. The squirrel family includes tree squirrels, ground squirrels, marmots, flying squirrels, prairie dogs amongst other rodents. Squirrels are indigenous to the Americas and Africa, were introduced by humans to Australia; the earliest known squirrels date from the Eocene period and are most related to the mountain beaver and to the dormouse among other living rodent families. The word "squirrel", first attested in 1327, comes from the Anglo-Norman esquirel, from the Old French escurel, the reflex of a Latin word sciurus; this Latin word was borrowed from the Ancient Greek word σκίουρος, which means shadow-tailed, referring to the bushy appendage possessed by many of its members. The native Old English word for the squirrel, ācweorna, survived only into Middle English before being replaced; the Old English word is of Common Germanic origin, cognates of which are still used in other Germanic languages, including the German Eichhörnchen, the Norwegian ikorn/ekorn, the Dutch eekhoorn, the Swedish ekorre and the Danish egern.
Squirrels are small animals, ranging in size from the African pygmy squirrel at 7–10 cm in length and just 10 g in weight, to the Laotian giant flying squirrel at 1.08 m in length and the Alpine marmot, which weighs from 5 to 8 kg. Squirrels have slender bodies with bushy tails and large eyes. In general, their fur is silky, though much thicker in some species than others; the coat color of squirrels is variable between—and even within—species. In most squirrel species, the hind limbs are longer than the fore limbs, while all species have either four or five toes on each paw; the paws, which include an poorly developed thumb, have soft pads on the undersides and versatile, sturdy claws for grasping and climbing. Tree squirrels, unlike most mammals, can descend a tree head-first, they do so by rotating their ankles 180 degrees, enabling the hind paws to point backward and thus grip the tree bark from the opposite direction. Squirrels live in every habitat, from tropical rainforest to semiarid desert, avoiding only the high polar regions and the driest of deserts.
They are predominantly herbivorous, subsisting on seeds and nuts, but many will eat insects and small vertebrates. As their large eyes indicate, squirrels have an excellent sense of vision, important for the tree-dwelling species. Many have a good sense of touch, with vibrissae on their limbs as well as their heads; the teeth of sciurids follow the typical rodent pattern, with large incisors that grow throughout life, cheek teeth that are set back behind a wide gap, or diastema. The typical dental formula for sciurids is 188.8.131.52.0.1.3. Many juvenile squirrels die in the first year of life. Adult squirrels can have a lifespan of 5 to 10 years in the wild; some can survive 10 to 20 years in captivity. Premature death may be caused when a nest falls from the tree, in which case the mother may abandon her young if their body temperature is not correct. Many such baby squirrels have been rescued and fostered by a professional wildlife rehabilitator until they could be safely returned to the wild, although the density of squirrel populations in many places and the constant care required by premature squirrels means that few rehabilitators are willing to spend their time doing this and such animals are euthanized instead.
Squirrels mate either once or twice a year and, following a gestation period of three to six weeks, give birth to a number of offspring that varies by species. The young are altricial, being born naked and blind. In most species of squirrel, the female alone looks after the young, which are weaned at six to ten weeks and become sexually mature by the end of their first year. In general, the ground-dwelling squirrel species are social living in well-developed colonies, while the tree-dwelling species are more solitary. Ground squirrels and tree squirrels are either diurnal or crepuscular, while the flying squirrels tend to be nocturnal—except for lactating flying squirrels and their young, which have a period of diurnality during the summer; because squirrels cannot digest cellulose, they must rely on foods rich in protein and fats. In temperate regions, early spring is the hardest time of year for squirrels because the nuts they buried are beginning to sprout, while many of the usual food sources have not yet become available.
During these times, squirrels rely on the buds of trees. Squirrels, being herbivores, eat a wide variety of plants, as well as nuts, conifer cones, fruits and green vegetation; some squirrels, however consume meat when faced with hunger. Squirrels have been known to eat small birds, young snakes, smaller rodents, as well as bird eggs and insects. Indeed, some tropical squirrel species have shifted entirely to a diet of insects. Predatory behavior has been observed in various species of ground squirrels, in particular the thirteen-lined ground squirrel. For example, Bailey, a scientist in the 1920s, observed a thirteen-lined ground squirrel preying upon a young chicken. Wistrand reported seeing this same species eating a freshly killed snake. Whitaker examined the stomachs of 139 thirteen-lined ground squirrels and found bird flesh in four of the specimens and the remains of a short-tailed shrew in one.
A leaf is an organ of a vascular plant and is the principal lateral appendage of the stem. The leaves and stem together form the shoot. Leaves are collectively referred to as foliage, as in "autumn foliage". A leaf is a thin, dorsiventrally flattened organ borne above ground and specialized for photosynthesis. In most leaves, the primary photosynthetic tissue, the palisade mesophyll, is located on the upper side of the blade or lamina of the leaf but in some species, including the mature foliage of Eucalyptus, palisade mesophyll is present on both sides and the leaves are said to be isobilateral. Most leaves have distinct upper surface and lower surface that differ in colour, the number of stomata, the amount and structure of epicuticular wax and other features. Leaves can have many different shapes and textures; the broad, flat leaves with complex venation of flowering plants are known as megaphylls and the species that bear them, the majority, as broad-leaved or megaphyllous plants. In the clubmosses, with different evolutionary origins, the leaves are simple and are known as microphylls.
Some leaves, such as bulb scales, are not above ground. In many aquatic species the leaves are submerged in water. Succulent plants have thick juicy leaves, but some leaves are without major photosynthetic function and may be dead at maturity, as in some cataphylls and spines. Furthermore, several kinds of leaf-like structures found in vascular plants are not homologous with them. Examples include flattened plant stems called phylloclades and cladodes, flattened leaf stems called phyllodes which differ from leaves both in their structure and origin; some structures of non-vascular plants function much like leaves. Examples include the phyllids of liverworts. Leaves are the most important organs of most vascular plants. Green plants are autotrophic, meaning that they do not obtain food from other living things but instead create their own food by photosynthesis, they capture the energy in sunlight and use it to make simple sugars, such as glucose and sucrose, from carbon dioxide and water. The sugars are stored as starch, further processed by chemical synthesis into more complex organic molecules such as proteins or cellulose, the basic structural material in plant cell walls, or metabolised by cellular respiration to provide chemical energy to run cellular processes.
The leaves draw water from the ground in the transpiration stream through a vascular conducting system known as xylem and obtain carbon dioxide from the atmosphere by diffusion through openings called stomata in the outer covering layer of the leaf, while leaves are orientated to maximise their exposure to sunlight. Once sugar has been synthesized, it needs to be transported to areas of active growth such as the plant shoots and roots. Vascular plants transport sucrose in a special tissue called the phloem; the phloem and xylem are parallel to each other but the transport of materials is in opposite directions. Within the leaf these vascular systems branch to form veins which supply as much of the leaf as possible, ensuring that cells carrying out photosynthesis are close to the transportation system. Leaves are broad and thin, thereby maximising the surface area directly exposed to light and enabling the light to penetrate the tissues and reach the chloroplasts, thus promoting photosynthesis.
They are arranged on the plant so as to expose their surfaces to light as efficiently as possible without shading each other, but there are many exceptions and complications. For instance plants adapted to windy conditions may have pendent leaves, such as in many willows and eucalyptss; the flat, or laminar, shape maximises thermal contact with the surrounding air, promoting cooling. Functionally, in addition to carrying out photosynthesis, the leaf is the principal site of transpiration, providing the energy required to draw the transpiration stream up from the roots, guttation. Many gymnosperms have thin needle-like or scale-like leaves that can be advantageous in cold climates with frequent snow and frost; these are interpreted as reduced from megaphyllous leaves of their Devonian ancestors. Some leaf forms are adapted to modulate the amount of light they absorb to avoid or mitigate excessive heat, ultraviolet damage, or desiccation, or to sacrifice light-absorption efficiency in favour of protection from herbivory.
For xerophytes the major constraint drought. Some window plants such as Fenestraria species and some Haworthia species such as Haworthia tesselata and Haworthia truncata are examples of xerophytes. and Bulbine mesembryanthemoides. Leaves function to store chemical energy and water and may become specialised organs serving other functions, such as tendrils of peas and other legumes, the protective spines of cacti and the insect traps in carnivorous plants such as Nepenthes and Sarracenia. Leaves are the fundamental structural units from which cones are constructed in gymnosperms and from which flowers are constructed in flowering plants; the internal organisation of most kinds of leaves has evolved to maximise exposure of the photosynthetic organelles, the chloroplasts, to light and to increase the absorption of carbon dioxide while at the same time controlling water loss. Their surfaces are waterproofed by the plant cuticle and gas exchange between the mesophyll cells and the atmosphere is controlled by minute openings called stomata which open or close to regulate the rate exchange of carbon dioxide and water vapour into
A birch is a thin-leaved deciduous hardwood tree of the genus Betula, in the family Betulaceae, which includes alders and hornbeams. It is related to the beech-oak family Fagaceae; the genus Betula contains 30 to 60 known taxa of which 11 are on the IUCN 2011 Red List of Threatened Species. They are a rather short-lived pioneer species widespread in the Northern Hemisphere in northern areas of temperate climates and in boreal climates. Birch species are small to medium-sized trees or shrubs of northern temperate and boreal climates; the simple leaves are alternate, singly or doubly serrate, feather-veined and stipulate. They appear in pairs, but these pairs are borne on spur-like, two-leaved, lateral branchlets; the fruit is a small samara. They differ from the alders in that the female catkins are not woody and disintegrate at maturity, falling apart to release the seeds, unlike the woody, cone-like female alder catkins; the bark of all birches is characteristically marked with long, horizontal lenticels, separates into thin, papery plates upon the paper birch.
Distinctive colors give the common names gray, black and yellow birch to different species. The buds form early and are full grown by midsummer, all are lateral, no terminal bud is formed; the wood of all the species is close-grained with a satiny texture and capable of taking a fine polish. The flowers are monoecious, opening with or before the leaves and borne once grown these leaves are 3–6 millimetres long on three-flowered clusters in the axils of the scales of drooping or erect catkins or aments. Staminate aments are pendulous, clustered or solitary in the axils of the last leaves of the branch of the year or near the ends of the short lateral branchlets of the year, they remain rigid during the winter. The scales of the staminate aments when mature are broadly ovate, yellow or orange color below the middle, dark chestnut brown at apex; each scale bears two bractlets and three sterile flowers, each flower consisting of a sessile, membranaceous two-lobed, calyx. Each calyx bears four short filaments with one-celled anthers or two filaments divided into two branches, each bearing a half-anther.
Anther cells open longitudinally. The pistillate aments are pendulous, solitary; the pistillate scales are oblong-ovate, three-lobed, pale yellow-green tinged with red, becoming brown at maturity. These scales bear each flower consisting of a naked ovary; the ovary is compressed, two-celled, crowned with two slender styles. Each scale bears a single small, winged nut, oval, with two persistent stigmas at the apex. Betula species are organised into five subgenera. Birches native to Europe and Asia include Betula albosinensis – Chinese red birch Betula alnoides – alder-leaf birch Betula ashburneri – Betula baschkirica – Betula bomiensis – Betula browicziana – Betula calcicola – Betula celtiberica – Betula chichibuensis – Betula chinensis – Chinese dwarf birch Betula coriaceifolia – Betula corylifolia – Betula costata – Betula cylindrostachya – Betula dahurica – Betula delavayi – Betula ermanii – Erman's birch Betula falcata – Betula fargesii – Betula fruticosa – Betula globispica – Betula gmelinii – Betula grossa – Japanese cherry birch Betula gynoterminalis – Betula honanensis – Betula humilis or Betula kamtschatica – Kamchatka birch platyphylla Betula insignis – Betula karagandensis – Betula klokovii – Betula kotulae – Betula litvinovii – Betula luminifera – Betula maximowiczii – monarch birch Betula medwediewii – Caucasian birch Betula megrelica – Betula microphylla – Betula nana – dwarf birch ) Betula pendula – silver birch Betula platyphylla – —Siberian silver birch Betula potamophila – Betula potaninii – Betula psammophila – Betula pubescens – downy birch known as white, European white or hairy birch Betula raddeana – Betula saksaren
The Niobrara River is a tributary of the Missouri River 568 miles long, running through the U. S. states of Nebraska. The river drains one of the most arid sections of the Great Plains, has a low flow for a river of its length; the Niobrara's watershed includes the northern tier of Nebraska Sandhills, a small south-central section of South Dakota, as well as a small area of eastern Wyoming. The river rises in southern Niobrara County; the Niobrara flows east as an intermittent stream past Lusk and southeast into northwestern Nebraska. It flows southeast across the Pine Ridge country of Sioux County east through Agate Fossil Beds National Monument, past Marsland, through Box Butte Reservoir; the stream flows east across northern Nebraska, near the northern edge of the Sandhills and past Valentine. It is joined by the Snake River about 13 miles southwest of Valentine. In north-central Nebraska it is joined by the Keya Paha River 6 miles west of Butte; the river joins the Missouri northwest of Niobrara in northern Knox County, just upstream of Lewis and Clark Lake.
Its total drainage basin is about 11,580 square miles. Although the annual runoff is low relative to the size of its drainage basin, the Niobrara has a stronger and more consistent flow than many other streams in the region. An estimated 70 percent of the river's water results from seepage from the Ogallala Aquifer that underlies the area, with the remaining 30 percent from precipitation; the river is lowest in early fall. Low flows in late summer and fall are the result of large irrigation diversions; the Niobrara's average discharge between 1958 and 2013, measured at the U. S. Geological Survey stream gage at Verdel, Nebraska, 14.8 miles above the mouth, is 1,757 cubic feet per second. The highest flow recorded was 39,100 cubic feet per second on March 27, 1960; the lowest daily mean was 102 cubic feet per second on November 13, 1960. The lower Niobrara valley is the traditional home of the Ponca tribe of Native Americans. Between 1861 and 1882, the stretch of the Niobrara River from the mouth of the Keya Paha to its confluence with the Missouri marked the boundary between Nebraska and the Dakota Territory.
A 76-mile stretch of the Niobrara River in central Nebraska, from the town of Valentine east to Nebraska State Highway 137, has been designated as the Niobrara National Scenic River since 1991. It is managed by the Department of the Interior to protect the water quality, paleontologic, fish & wildlife and recreation values. Most of the lands within the boundary of the National Scenic River are, will remain, in private ownership. Management is based upon working with private, county and federal landowners and stakeholders to coordinate protection of the river while ensuring a quality experience for river visitors; the U. S. Fish and Wildlife Service manages the 9 miles of river that flow through the Fort Niobrara Refuge for wilderness and wildlife habitat, but allows recreation downstream from Cornell Dam; the National Park Service manages the remaining 67 miles, acting as a facilitator for resource protection by landowners and river users, providing law enforcement and visitor education services, coordinating resource management activities.
The Box Butte Dam, completed in 1946 by the U. S. Bureau of Reclamation, is the only major dam on the Niobrara River proper. Located in Dawes County in western Nebraska, the dam is part of the Mirage Flats Project, which irrigates 11,670 acres on the north side of the Niobrara River. Dunlap Diversion Dam, 8 miles below Box Butte, diverts water through a 13-mile canal to the farmland; the Snake River tributary is impounded by the Merritt Dam and irrigates about 34,540 acres in the area of Valentine, Nebraska. The project is part of the Ainsworth Unit of the Pick-Sloan Missouri Basin Program operated by the USBR. Cornell Dam, built in 1915 at the confluence of Minnechaduza Creek near Valentine, generated power until 1985; the following year the dam was acquired by the U. S. Department of the Interior. Although decommissioned, the dam remains standing; the feasibility of removing the defunct dam has been studied, although the accumulation of sediment behind the dam, which may include high levels of chemicals from pesticides, may be harmful to the river environment if released.
Spencer Dam, about 50 miles from the mouth of the Niobrara, was the last operational hydroelectric plant on the river. The dam was operated by the Nebraska Public Power District, it includes two Westinghouse generators, with a combined capacity of 3,000 KW. In a 2015 agreement with Nebraska local and state government entities, NPPD agreed to decommission the dam in 2017; the dam was breached by flooding caused by a March 2019 storm. In the Cheyenne language, the river is Hisse Yovi Yoe, meaning "surprise river". Niobrara National Scenic River Niobrara State Park, located at the confluence of the Niobrara and Missouri Rivers Smith Falls Fort Niobrara, a U. S. Army outpost Fort Niobrara National Wildlife Refuge Samuel R. McKelvie National Forest Box Butte Reservoir Agate Fossil Beds National Monument List of longest rivers of the United States List of rivers of the United States List of Nebraska rivers List of Wyoming rivers Niobrara Na
Quebec is one of the thirteen provinces and territories of Canada. It is bordered to the west by the province of Ontario and the bodies of water James Bay and Hudson Bay. S. states of Maine, New Hampshire and New York. It shares maritime borders with Nunavut, Prince Edward Island, Nova Scotia. Quebec is Canada's largest province by its second-largest administrative division, it is and politically considered to be part of Central Canada. Quebec is the second-most populous province of Canada, after Ontario, it is the only one to have a predominantly French-speaking population, with French as the sole provincial official language. Most inhabitants live in urban areas near the Saint Lawrence River between Montreal and Quebec City, the capital. Half of Quebec residents live in the Greater Montreal Area, including the Island of Montreal. English-speaking communities and English-language institutions are concentrated in the west of the island of Montreal but are significantly present in the Outaouais, Eastern Townships, Gaspé regions.
The Nord-du-Québec region, occupying the northern half of the province, is sparsely populated and inhabited by Aboriginal peoples. The climate around the major cities is four-seasons continental with cold and snowy winters combined with warm to hot humid summers, but farther north long winter seasons dominate and as a result the northern areas of the province are marked by tundra conditions. In central Quebec, at comparatively southerly latitudes, winters are severe in inland areas. Quebec independence debates have played a large role in the politics of the province. Parti Québécois governments held referendums on sovereignty in 1980 and 1995. Although neither passed, the 1995 referendum saw the highest voter turnout in Quebec history, at over 93%, only failed by less than 1%. In 2006, the House of Commons of Canada passed a symbolic motion recognizing the "Québécois as a nation within a united Canada". While the province's substantial natural resources have long been the mainstay of its economy, sectors of the knowledge economy such as aerospace and communication technologies and the pharmaceutical industry play leading roles.
These many industries have all contributed to helping Quebec become an economically influential province within Canada, second only to Ontario in economic output. The name "Québec", which comes from the Algonquin word kébec meaning "where the river narrows" referred to the area around Quebec City where the Saint Lawrence River narrows to a cliff-lined gap. Early variations in the spelling of the name included Kébec. French explorer Samuel de Champlain chose the name Québec in 1608 for the colonial outpost he would use as the administrative seat for the French colony of New France; the province is sometimes referred to as "La belle province". The Province of Quebec was founded in the Royal Proclamation of 1763 after the Treaty of Paris formally transferred the French colony of Canada to Britain after the Seven Years' War; the proclamation restricted the province to an area along the banks of the Saint Lawrence River. The Quebec Act of 1774 expanded the territory of the province to include the Great Lakes and the Ohio River Valley and south of Rupert's Land, more or less restoring the borders existing under French rule before the Conquest of 1760.
The Treaty of Paris ceded territories south of the Great Lakes to the United States. After the Constitutional Act of 1791, the territory was divided between Lower Canada and Upper Canada, with each being granted an elected legislative assembly. In 1840, these become Canada East and Canada West after the British Parliament unified Upper and Lower Canada into the Province of Canada; this territory was redivided into the Provinces of Quebec and Ontario at Confederation in 1867. Each became one of the first four provinces. In 1870, Canada purchased Rupert's Land from the Hudson's Bay Company and over the next few decades the Parliament of Canada transferred to Quebec portions of this territory that would more than triple the size of the province. In 1898, the Canadian Parliament passed the first Quebec Boundary Extension Act that expanded the provincial boundaries northward to include the lands of the local aboriginal peoples; this was followed by the addition of the District of Ungava through the Quebec Boundaries Extension Act of 1912 that added the northernmost lands of the Inuit to create the modern Province of Quebec.
In 1927, the border between Quebec and Newfoundland and Labrador was established by the British Judicial Committee of the Privy Council. Quebec disputes this boundary. Located in the eastern part of Canada, part of Central Canada, Quebec occupies a territory nearly three times the size of France or Texas, most of, sparsely populated, its topography is different from one region to another due to the varying composition of the ground, the climate, the proximity to water. The Saint Lawrence Lowland and the Appalachians are the two main topographic regions in southern Quebec, while the Canadian Shield occupies most of central and northern Quebec. Quebec has one of the world's largest reserves of fresh water, occupying 12% of its surface, it has 3 % of the world's renewable fresh water. Mor
Carl Linnaeus known after his ennoblement as Carl von Linné, was a Swedish botanist and zoologist who formalised binomial nomenclature, the modern system of naming organisms. He is known as the "father of modern taxonomy". Many of his writings were in Latin, his name is rendered in Latin as Carolus Linnæus. Linnaeus was born in the countryside of Småland in southern Sweden, he received most of his higher education at Uppsala University and began giving lectures in botany there in 1730. He lived abroad between 1735 and 1738, where he studied and published the first edition of his Systema Naturae in the Netherlands, he returned to Sweden where he became professor of medicine and botany at Uppsala. In the 1740s, he was sent on several journeys through Sweden to find and classify plants and animals. In the 1750s and 1760s, he continued to collect and classify animals and minerals, while publishing several volumes, he was one of the most acclaimed scientists in Europe at the time of his death. Philosopher Jean-Jacques Rousseau sent him the message: "Tell him I know no greater man on earth."
Johann Wolfgang von Goethe wrote: "With the exception of Shakespeare and Spinoza, I know no one among the no longer living who has influenced me more strongly." Swedish author August Strindberg wrote: "Linnaeus was in reality a poet who happened to become a naturalist." Linnaeus has been called Princeps botanicorum and "The Pliny of the North". He is considered as one of the founders of modern ecology. In botany and zoology, the abbreviation L. is used to indicate Linnaeus as the authority for a species' name. In older publications, the abbreviation "Linn." is found. Linnaeus's remains comprise the type specimen for the species Homo sapiens following the International Code of Zoological Nomenclature, since the sole specimen that he is known to have examined was himself. Linnaeus was born in the village of Råshult in Småland, Sweden, on 23 May 1707, he was the first child of Christina Brodersonia. His siblings were Anna Maria Linnæa, Sofia Juliana Linnæa, Samuel Linnæus, Emerentia Linnæa, his father taught him Latin as a small child.
One of a long line of peasants and priests, Nils was an amateur botanist, a Lutheran minister, the curate of the small village of Stenbrohult in Småland. Christina was the daughter of the rector of Samuel Brodersonius. A year after Linnaeus's birth, his grandfather Samuel Brodersonius died, his father Nils became the rector of Stenbrohult; the family moved into the rectory from the curate's house. In his early years, Linnaeus seemed to have a liking for plants, flowers in particular. Whenever he was upset, he was given a flower, which calmed him. Nils spent much time in his garden and showed flowers to Linnaeus and told him their names. Soon Linnaeus was given his own patch of earth. Carl's father was the first in his ancestry to adopt a permanent surname. Before that, ancestors had used the patronymic naming system of Scandinavian countries: his father was named Ingemarsson after his father Ingemar Bengtsson; when Nils was admitted to the University of Lund, he had to take on a family name. He adopted the Latinate name Linnæus after a giant linden tree, lind in Swedish, that grew on the family homestead.
This name was spelled with the æ ligature. When Carl was born, he was named Carl Linnæus, with his father's family name; the son always spelled it with the æ ligature, both in handwritten documents and in publications. Carl's patronymic would have been Nilsson, as in Carl Nilsson Linnæus. Linnaeus's father began teaching him basic Latin and geography at an early age; when Linnaeus was seven, Nils decided to hire a tutor for him. The parents picked a son of a local yeoman. Linnaeus did not like him, writing in his autobiography that Telander "was better calculated to extinguish a child's talents than develop them". Two years after his tutoring had begun, he was sent to the Lower Grammar School at Växjö in 1717. Linnaeus studied going to the countryside to look for plants, he reached the last year of the Lower School when he was fifteen, taught by the headmaster, Daniel Lannerus, interested in botany. Lannerus gave him the run of his garden, he introduced him to Johan Rothman, the state doctor of Småland and a teacher at Katedralskolan in Växjö.
A botanist, Rothman broadened Linnaeus's interest in botany and helped him develop an interest in medicine. By the age of 17, Linnaeus had become well acquainted with the existing botanical literature, he remarks in his journal that he "read day and night, knowing like the back of my hand, Arvidh Månsson's Rydaholm Book of Herbs, Tillandz's Flora Åboensis, Palmberg's Serta Florea Suecana, Bromelii Chloros Gothica and Rudbeckii Hortus Upsaliensis...."Linnaeus entered the Växjö Katedralskola in 1724, where he studied Greek, Hebrew and mathematics, a curriculum designed for boys preparing for the priesthood. In the last year at the gymnasium, Linnaeus's father visited to ask the professors how his son's studies were progressing. Rothman believed otherwise; the doctor offered to have Linnaeus live with his family in Växjö and to teach him physiology and botany. Nils accepted this offer. Rothman showed Linnaeus that botany was a serious sub
In botany, stipule is a term coined by Linnaeus which refers to outgrowths borne on either side of the base of a leafstalk. A pair of stipules is considered part of the anatomy of the leaf of a typical flowering plant, although in many species the stipules are inconspicuous or absent. In some older botanical writing, the term "stipule" was used more to refer to any small leaves or leaf-parts, notably prophylls; the position of stipules on a plant varies from species to species, though they are located near the base of a leaf. Stipules are most common on dicotyledons; some monocotyledon plants only display one per leaf. A relationship exists between the anatomy of the stem node and the presence or absence of stipules: most plants with trilacunar nodes have stipules. Stipules are morphologically variable and might appear as glands, hairs, spines, or laminar structures. If a single stipule goes all the way around the stem, it is known as an ochrea; the three types of stipules according to duration are caducous and persistent.
Caducous stipules fall off before the leaf unfolds, while deciduous stipules fall off after the leaf unfolds. Persistent stipules remain attached to the plant. Stipules can be considered free lateral, interpetiolar, ochreate, bud scales, tendrillar or spiny. A stipule can be fused to the other stipule from the same node. A stipule is "adnate" if it's fused together on part of the petiole length, but the anterior is still free. A stipule is "interpetiolar" if it is located in between the petioles, as opposed to being attached to the petioles, one stipule from each leaf is fused together, so it appears that there's just one stipule between each leaf. A stipule is "intrapetiolar" if it is located in the angle that's between a petiole. In this case, the two stipules form together and appear to be one stipule. A stipule is "ochreate" if a single stipule appears to be a solid tube that goes all the way around the stem. A stipule is "foliaceous"; these are used to photosynthesize. A stipule protects leaf buds as they form.
These fall off as soon as the leaf unfolds. A stipule is considered "tendrillar" if they are long thin tendrils, are used by climbing plants. A stipule is considered "spiny" if they are pointy; these are used to deter animals. A stipule is considered to be "abaxial", "counter" or "leaf opposed" if it's located on the opposite side to where the leaf meets the stem. Stipules have various functions; some stipules may be vestigial. It is known. Sometimes stipules protect the next leaf or bud as it grows in falls off after the leaf unfolds, as with Tulip Poplars. Stipules can be used as climbing tendrils by climbing plants. Spiny stipules can be used to help protect the plant from animals. Esau, K. 1953. Plant Anatomy. Second Edition. John Wiley & Sons, Inc. New York, Sidney. 767 pp. Stipules and stipels