Arachnids are a class of joint-legged invertebrate animals, in the subphylum Chelicerata. All adult arachnids have eight legs, although the front pair of legs in some species has converted to a sensory function, while in other species, different appendages can grow large enough to take on the appearance of extra pairs of legs; the term is derived from the Greek word ἀράχνη, from the myth of the hubristic human weaver Arachne, turned into a spider. Spiders are the largest order in the class, which includes scorpions, mites and solifuges. In 2019, a molecular phylogenetic study placed horseshoe crabs in Arachnida. All extant arachnids are terrestrial, living on land. However, some inhabit freshwater environments and, with the exception of the pelagic zone, marine environments as well, they comprise over 100,000 named species. All adult arachnids have eight legs, arachnids may be distinguished from insects by this fact, since insects have six legs. However, arachnids have two further pairs of appendages that have become adapted for feeding and sensory perception.
The first pair, the chelicerae, serve in defense. The next pair of appendages, the pedipalps, have been adapted for feeding, and/or reproductive functions. In Solifugae, the palps are quite leg-like; the larvae of mites and Ricinulei have only six legs. However, mites are variable: as well as eight, there are adult mites with six or four legs. Arachnids are further distinguished from insects by the fact, their body is organized into two tagmata, called the prosoma, or cephalothorax, the opisthosoma, or abdomen. The cephalothorax is derived from the fusion of the cephalon and the thorax, is covered by a single, unsegmented carapace; the abdomen is segmented in the more primitive forms, but varying degrees of fusion between the segments occur in many groups. It is divided into a preabdomen and postabdomen, although this is only visible in scorpions, in some orders, such as the Acari, the abdominal sections are fused. A telson is present in scorpions, where it has been modified to a stinger, in the Schizomida, whip scorpions and Palpigradi.
Like all arthropods, arachnids have an exoskeleton, they have an internal structure of cartilage-like tissue, called the endosternite, to which certain muscle groups are attached. The endosternite is calcified in some Opiliones. Most arachnids lack extensor muscles in the distal joints of their appendages. Spiders and whipscorpions extend their limbs hydraulically using the pressure of their hemolymph. Solifuges and some harvestmen extend their knees by the use of elastic thickenings in the joint cuticle. Scorpions and some harvestmen have evolved muscles that extend two leg joints at once; the equivalent joints of the pedipalps of scorpions though, are extended by elastic recoil. There are characteristics that are important for the terrestrial lifestyle of arachnids, such as internal respiratory surfaces in the form of tracheae, or modification of the book gill into a book lung, an internal series of vascular lamellae used for gas exchange with the air. While the tracheae are individual systems of tubes, similar to those in insects, ricinuleids and some spiders possess sieve tracheae, in which several tubes arise in a bundle from a small chamber connected to the spiracle.
This type of tracheal system has certainly evolved from the book lungs, indicates that the tracheae of arachnids are not homologous with those of insects. Further adaptations to terrestrial life are appendages modified for more efficient locomotion on land, internal fertilisation, special sensory organs, water conservation enhanced by efficient excretory structures as well as a waxy layer covering the cuticle; the excretory glands of arachnids include up to four pairs of coxal glands along the side of the prosoma, one or two pairs of Malpighian tubules, emptying into the gut. Many arachnids have the other type of excretory gland, although several do have both; the primary nitrogenous waste product in arachnids is guanine. Arachnid blood is variable in composition, depending on the mode of respiration. Arachnids with an efficient tracheal system do not need to transport oxygen in the blood, may have a reduced circulatory system. In scorpions and some spiders, the blood contains haemocyanin, a copper-based pigment with a similar function to haemoglobin in vertebrates.
The heart is located in the forward part of the abdomen, may or may not be segmented. Some mites have no heart at all. Arachnids are carnivorous, feeding on the pre-digested bodies of insects and other small animals. Only in the harvestmen and among mites, such as the house dust mite, is there ingestion of solid food particles, thus exposure to internal parasites, although it is not unusual for spiders to eat their own silk. Several groups secrete venom from specialized glands to kill prey or enemies. Several mites and ticks are parasites. Arachnids produce digestive juices in their stomachs, use their pedipalps and chelicerae to pour them over their dead prey; the digestive juices turn the prey into a broth of nutrients, which the arachnid sucks into a pre-buccal cavity located in front of the mouth. Behind the mouth is a muscular, sclerotised pharynx, which acts as a pump, sucking the food through the mouth and on into the oesophagus and stomach. In some arachnids, the oesophagus a
Eriophyidae is a family of more than 200 genera of mites, which live as plant parasites causing galls or other damage to the plant tissues and hence known as gall mites. About 3,600 species have been described, but this is less than 10% of the actual number existing in this poorly researched family, they are yellow to pinkish white to purplish in color. The mites are worm like, have only two pairs of legs, their primary method of population spread is by wind. They affect a wide range of plants, several are major pest species causing substantial economic damage to crops; some species, are used as biological agents to control weeds and invasive plant species. Notable species in this family include: Abacarus hystrix, the cereal rust mite Abacarus sacchari, the sugarcane rust mite Acalitus essigi, the redberry mite, which affects blackberries Aceria chondrillae, the chondrilla gall mite, an agent of biological control against skeleton weed Aceria guerreronis, a mite of coconuts Aceria malherbae, the bindweed gall mite, an agent of biological control against field bindweed Eriophyes padi, a mite that causes cherry pouch galls on black cherry trees Digital museum of Nature: Classification of Eriophyoid Mites Fauna Europaea Joel Hallan's Biology Catalog: Eriophyidae Acalitus vaccinii, blueberry bud mite on the UF / IFAS Featured Creatures Web site Acathrix trymatus, a coconut mite on the UF / IFAS Featured Creatures Web site Aceria guerreronis, a coconut mite on the UF / IFAS Featured Creatures Web site
Galls or cecidia are a kind of swelling growth on the external tissues of plants, fungi, or animals. Plant galls are abnormal outgrowths of similar to benign tumors or warts in animals, they can be caused by various parasites, from viruses and bacteria, to other plants and mites. Plant galls are highly organized structures and because of this the cause of the gall can be determined without the actual agent being identified; this applies to some insect and mite plant galls. The study of plant galls is known as cecidology. In human pathology, a gall is a raised sore on the skin caused by chafing or rubbing. Insect galls are the distinctive plant structures formed by some herbivorous insects as their own microhabitats, they are plant tissue, controlled by the insect. Galls act as food source for the maker of the gall; the interior of a gall can contain other tissues. Some galls act as "physiologic sinks", concentrating resources in the gall from the surrounding plant parts. Galls may provide the insect with physical protection from predators.
Insect galls are induced by chemicals injected by the larvae of the insects into the plants, mechanical damage. After the galls are formed, the larvae develop inside until grown, when they leave. In order to form galls, the insects must take advantage of the time when plant cell division occurs quickly: the growing season spring in temperate climates, but, extended in the tropics; the meristems, where plant cell division occurs, are the usual sites of galls, though insect galls can be found on other parts of the plant, such as the leaves, branches, buds and flowers and fruits. Gall-inducing insects are species-specific and sometimes tissue-specific on the plants they gall. Gall-inducing insects include gall wasps, gall midges, gall flies, Agromyzidae aphids, scale insects, goldenrod gall fly and weevils. Galls produced by insects and mites include: Ash flower gall: this gall is caused by a small mite that causes irregular distortion of male flowers; the galls are green dry and turn brown. Ash midrib gall: 0.5 to 1 inch long, these galls are succulent and have thick walls.
A small cavity within each gall contains one or more small maggots, the larval stages of small flies called midges. Female midges lay their eggs in young leaflets during early spring. Gall formation begins. Specifics of the biology of this insect are not known; the galls do not harm tree health. Elm cockscomb gall: these distinct galls, caused by an aphid, are about one inch long and about 1/4 inch high; the irregular edge of the gall and its red color at maturity account for the common name. The galls dry and turn brown as they age. Aphids may be seen through a slit-like opening in the underside of the gall; this insect has a complex life cycle—it forms galls on elm in early summer feeds on grass roots in the summer. The galls do not cause significant harm to the tree. Hackberry leaf gall: this gall is caused by a small aphid-like insect with sucking mouthparts called a jumping plant louse; the adults can invade houses in large numbers in the fall. Females lay eggs over a long period of time beginning when leaves begin to unfold from the buds in the spring.
Feeding by the nymphs that hatch from these eggs causes abnormal plant growth. The psyllids remain inside the galIs. There is one generation each year. Heavy infestations can result in premature leaf drop which over a series of years may affect tree health. Honeylocust pod gall: this gall is caused by a small fly; the sunburst cultivar appears to be susceptible to this pest. Infested leaves have globular or pod-like distortions. Infestations begin. There are five or more generations each year. Infested leaves drop prematurely and repeated damage can kill small branches. New shoots develop at the base of dead twigs; as a result, the natural shape of the tree may be lost. Oak gall: see Oak apple Petiole and stipule galls: thick globe-like galls can develop on leaf petioles and stems. Many of these are caused by insects called phylloxerans which are similar to aphids; the hard, woody galls may remain on the tree for several years. There is one generation each year and the insects over winter on the tree in the egg stage.
Willow shoot galls: these swellings on shoots, twigs, or leaf petioles, may be caused by small flies or small wasps. The gall increases in size, they cause no significant injury. The infestation may be reduced by pruning and destroying the galled areas before the adult insect emerges in late summer. Witchhazel gall: this gall is caused by an aphid that passes the winter in eggs laid on twigs of the plant. Feeding by the aphid causes the formation of conical galls on the upper side of the leaf; each gall, produced by a single aphid becomes filled with offspring. Mature aphids with wings fly to birch. After several generations there, the insects return to witch hazel to lay the eggs that survive the winter. No galls are formed on the birch. Many rust fungi induce gall formation, including western gall rust, which infects a variety of pine trees and cedar-apple rust. G
Wikispecies is a wiki-based online project supported by the Wikimedia Foundation. Its aim is to create a comprehensive free content catalogue of all species. Jimmy Wales stated that editors are not required to fax in their degrees, but that submissions will have to pass muster with a technical audience. Wikispecies is available under the GNU Free Documentation License and CC BY-SA 3.0. Started in September 2004, with biologists across the world invited to contribute, the project had grown a framework encompassing the Linnaean taxonomy with links to Wikipedia articles on individual species by April 2005. Benedikt Mandl co-ordinated the efforts of several people who are interested in getting involved with the project and contacted potential supporters in early summer 2004. Databases were evaluated and the administrators contacted, some of them have agreed on providing their data for Wikispecies. Mandl defined two major tasks: Figure out how the contents of the data base would need to be presented—by asking experts, potential non-professional users and comparing that with existing databases Figure out how to do the software, which hardware is required and how to cover the costs—by asking experts, looking for fellow volunteers and potential sponsorsAdvantages and disadvantages were discussed by the wikimedia-I mailing list.
The board of directors of the Wikimedia Foundation voted by 4 to 0 in favor of the establishment of a Wikispecies. The project is hosted at species.wikimedia.org. It was merged to a sister project of Wikimedia Foundation on September 14, 2004. On October 10, 2006, the project exceeded 75,000 articles. On May 20, 2007, the project exceeded 100,000 articles with a total of 5,495 registered users. On September 8, 2008, the project exceeded 150,000 articles with a total of 9,224 registered users. On October 23, 2011, the project reached 300,000 articles. On June 16, 2014, the project reached 400,000 articles. On January 7, 2017, the project reached 500,000 articles. On October 30, 2018, the project reached 600,000 articles, a total of 1.12 million pages. Wikispecies comprises taxon pages, additionally pages about synonyms, taxon authorities, taxonomical publications, institutions or repositories holding type specimen. Wikispecies asks users to use images from Wikimedia Commons. Wikispecies does not allow the use of content.
All Species Foundation Catalogue of Life Encyclopedia of Life Tree of Life Web Project List of online encyclopedias The Plant List Wikispecies, The free species directory that anyone can edit Species Community Portal The Wikispecies Charter, written by Wales
Vasates quadripedes, the maple bladder-gall mite, is an eriophyid mite in the genus Vasates, which causes galls on the leaves of silver maple, red maple, sugar maple. The gall is rounded, sometimes elongate, has a short, thin neck. Galls are 2–3 millimetres in diameter, may be numerous on the upper surfaces of leaves, they have an opening in the lower surface. At first they are yellowish-green or bright red they become dark red and black. In Britain, the mite affects introduced silver maple; the species is new to Britain, being first recorded in London in 2002
An arthropod is an invertebrate animal having an exoskeleton, a segmented body, paired jointed appendages. Arthropods form the phylum Euarthropoda, which includes insects, arachnids and crustaceans; the term Arthropoda as proposed refers to a proposed grouping of Euarthropods and the phylum Onychophora. Arthropods are characterized by their jointed limbs and cuticle made of chitin mineralised with calcium carbonate; the arthropod body plan consists of each with a pair of appendages. The rigid cuticle inhibits growth, so arthropods replace it periodically by moulting. Arthopods are bilaterally symmetrical and their body possesses an external skeleton; some species have wings. Their versatility has enabled them to become the most species-rich members of all ecological guilds in most environments, they have over a million described species, making up more than 80 per cent of all described living animal species, some of which, unlike most other animals, are successful in dry environments. Arthropods range in size from the microscopic crustacean Stygotantulus up to the Japanese spider crab.
Arthropods' primary internal cavity is a haemocoel, which accommodates their internal organs, through which their haemolymph – analogue of blood – circulates. Like their exteriors, the internal organs of arthropods are built of repeated segments, their nervous system is "ladder-like", with paired ventral nerve cords running through all segments and forming paired ganglia in each segment. Their heads are formed by fusion of varying numbers of segments, their brains are formed by fusion of the ganglia of these segments and encircle the esophagus; the respiratory and excretory systems of arthropods vary, depending as much on their environment as on the subphylum to which they belong. Their vision relies on various combinations of compound eyes and pigment-pit ocelli: in most species the ocelli can only detect the direction from which light is coming, the compound eyes are the main source of information, but the main eyes of spiders are ocelli that can form images and, in a few cases, can swivel to track prey.
Arthropods have a wide range of chemical and mechanical sensors based on modifications of the many setae that project through their cuticles. Arthropods' methods of reproduction and development are diverse; the evolutionary ancestry of arthropods dates back to the Cambrian period. The group is regarded as monophyletic, many analyses support the placement of arthropods with cycloneuralians in a superphylum Ecdysozoa. Overall, the basal relationships of Metazoa are not yet well resolved; the relationships between various arthropod groups are still debated. Aquatic species use either external fertilization. All arthropods lay eggs, but scorpions give birth to live young after the eggs have hatched inside the mother. Arthropod hatchlings vary from miniature adults to grubs and caterpillars that lack jointed limbs and undergo a total metamorphosis to produce the adult form; the level of maternal care for hatchlings varies from nonexistent to the prolonged care provided by scorpions. Arthropods contribute to the human food supply both directly as food, more indirectly as pollinators of crops.
Some species are known to spread severe disease to humans and crops. The word arthropod comes from the Greek ἄρθρον árthron, "joint", πούς pous, i.e. "foot" or "leg", which together mean "jointed leg". Arthropods are invertebrates with jointed limbs; the exoskeleton or cuticles consists of a polymer of glucosamine. The cuticle of many crustaceans, beetle mites, millipedes is biomineralized with calcium carbonate. Calcification of the endosternite, an internal structure used for muscle attachments occur in some opiliones. Estimates of the number of arthropod species vary between 1,170,000 and 5 to 10 million and account for over 80 per cent of all known living animal species; the number of species remains difficult to determine. This is due to the census modeling assumptions projected onto other regions in order to scale up from counts at specific locations applied to the whole world. A study in 1992 estimated that there were 500,000 species of animals and plants in Costa Rica alone, of which 365,000 were arthropods.
They are important members of marine, freshwater and air ecosystems, are one of only two major animal groups that have adapted to life in dry environments. One arthropod sub-group, insects, is the most species-rich member of all ecological guilds in land and freshwater environments; the lightest insects weigh less than 25 micrograms. Some living crustaceans are much larger; the embryos of all arthropods are segmented, built from a series of repeated modules. The last common ancestor of living arthropods consisted of a series of undifferentiated segments, each with a pair of appendages that functioned as limbs. However, all known living and fossil arthropods have grouped segments into tagmata in which segments and their limbs are specialized in various ways; the three-
Animals are multicellular eukaryotic organisms that form the biological kingdom Animalia. With few exceptions, animals consume organic material, breathe oxygen, are able to move, can reproduce sexually, grow from a hollow sphere of cells, the blastula, during embryonic development. Over 1.5 million living animal species have been described—of which around 1 million are insects—but it has been estimated there are over 7 million animal species in total. Animals range in length from 8.5 millionths of a metre to 33.6 metres and have complex interactions with each other and their environments, forming intricate food webs. The category includes humans, but in colloquial use the term animal refers only to non-human animals; the study of non-human animals is known as zoology. Most living animal species are in the Bilateria, a clade whose members have a bilaterally symmetric body plan; the Bilateria include the protostomes—in which many groups of invertebrates are found, such as nematodes and molluscs—and the deuterostomes, containing the echinoderms and chordates.
Life forms interpreted. Many modern animal phyla became established in the fossil record as marine species during the Cambrian explosion which began around 542 million years ago. 6,331 groups of genes common to all living animals have been identified. Aristotle divided animals into those with those without. Carl Linnaeus created the first hierarchical biological classification for animals in 1758 with his Systema Naturae, which Jean-Baptiste Lamarck expanded into 14 phyla by 1809. In 1874, Ernst Haeckel divided the animal kingdom into the multicellular Metazoa and the Protozoa, single-celled organisms no longer considered animals. In modern times, the biological classification of animals relies on advanced techniques, such as molecular phylogenetics, which are effective at demonstrating the evolutionary relationships between animal taxa. Humans make use of many other animal species for food, including meat and eggs. Dogs have been used in hunting, while many aquatic animals are hunted for sport.
Non-human animals have appeared in art from the earliest times and are featured in mythology and religion. The word "animal" comes from the Latin animalis, having soul or living being; the biological definition includes all members of the kingdom Animalia. In colloquial usage, as a consequence of anthropocentrism, the term animal is sometimes used nonscientifically to refer only to non-human animals. Animals have several characteristics. Animals are eukaryotic and multicellular, unlike bacteria, which are prokaryotic, unlike protists, which are eukaryotic but unicellular. Unlike plants and algae, which produce their own nutrients animals are heterotrophic, feeding on organic material and digesting it internally. With few exceptions, animals breathe oxygen and respire aerobically. All animals are motile during at least part of their life cycle, but some animals, such as sponges, corals and barnacles become sessile; the blastula is a stage in embryonic development, unique to most animals, allowing cells to be differentiated into specialised tissues and organs.
All animals are composed of cells, surrounded by a characteristic extracellular matrix composed of collagen and elastic glycoproteins. During development, the animal extracellular matrix forms a flexible framework upon which cells can move about and be reorganised, making the formation of complex structures possible; this may be calcified, forming structures such as shells and spicules. In contrast, the cells of other multicellular organisms are held in place by cell walls, so develop by progressive growth. Animal cells uniquely possess the cell junctions called tight junctions, gap junctions, desmosomes. With few exceptions—in particular, the sponges and placozoans—animal bodies are differentiated into tissues; these include muscles, which enable locomotion, nerve tissues, which transmit signals and coordinate the body. There is an internal digestive chamber with either one opening or two openings. Nearly all animals make use of some form of sexual reproduction, they produce haploid gametes by meiosis.
These fuse to form zygotes, which develop via mitosis into a hollow sphere, called a blastula. In sponges, blastula larvae swim to a new location, attach to the seabed, develop into a new sponge. In most other groups, the blastula undergoes more complicated rearrangement, it first invaginates to form a gastrula with a digestive chamber and two separate germ layers, an external ectoderm and an internal endoderm. In most cases, a third germ layer, the mesoderm develops between them; these germ layers differentiate to form tissues and organs. Repeated instances of mating with a close relative during sexual reproduction leads to inbreeding depression within a population due to the increased prevalence of harmful recessive traits. Animals have evolved numerous mechanisms for avoiding close inbreeding. In some species, such as the splendid fairywren, females benefit by mating with multiple males, thus producing more offspring of higher genetic quality; some animals are capable of asexual reproduction, which results