The Fabaceae or Leguminosae known as the legume, pea, or bean family, are a large and economically important family of flowering plants. It includes trees and perennial or annual herbaceous plants, which are recognized by their fruit and their compound, stipulate leaves. Many legumes have characteristic fruits; the family is distributed, is the third-largest land plant family in terms of number of species, behind only the Orchidaceae and Asteraceae, with about 751 genera and about 19,000 known species. The five largest of the genera are Astragalus, Indigofera and Mimosa, which constitute about a quarter of all legume species; the ca. 19,000 known legume species amount to about 7% of flowering plant species. Fabaceae is the most common family found in tropical rainforests and in dry forests in the Americas and Africa. Recent molecular and morphological evidence supports the fact that the Fabaceae is a single monophyletic family; this conclusion has been supported not only by the degree of interrelation shown by different groups within the family compared with that found among the Leguminosae and their closest relations, but by all the recent phylogenetic studies based on DNA sequences.
These studies confirm that the Fabaceae are a monophyletic group, related to the Polygalaceae and Quillajaceae families and that they belong to the order Fabales. Along with the cereals, some fruits and tropical roots, a number of Leguminosae have been a staple human food for millennia and their use is related to human evolution; the Fabaceae family includes a number of important agricultural and food plants, including Glycine max, Pisum sativum, Cicer arietinum, Medicago sativa, Arachis hypogaea, Ceratonia siliqua, Glycyrrhiza glabra. A number of species are weedy pests in different parts of the world, including: Cytisus scoparius, Robinia pseudoacacia, Ulex europaeus, Pueraria lobata, a number of Lupinus species; the name'Fabaceae' comes from the defunct genus Faba, now included in Vicia. The term "faba" comes from Latin, appears to mean "bean". Leguminosae is an older name still considered valid, refers to the fruit of these plants, which are called legumes. Fabaceae range in habit from giant trees to small annual herbs, with the majority being herbaceous perennials.
Plants have indeterminate inflorescences. The flowers have a short hypanthium and a single carpel with a short gynophore, after fertilization produce fruits that are legumes; the Leguminosae have a wide variety of growth forms, including trees, herbaceous plants, vines or lianas. The herbaceous plants can be annuals, biennials, or perennials, without basal or terminal leaf aggregations. Many Legumes have tendrils, they are epiphytes, or vines. The latter support themselves by means of shoots that twist around a support or through cauline or foliar tendrils. Plants can be mesophytes, or xerophytes; the leaves are alternate and compound. Most they are even- or odd-pinnately compound trifoliate and palmately compound, in the Mimosoideae and the Caesalpinioideae bipinnate, they always have stipules, which can be rather inconspicuous. Leaf margins are entire or serrate. Both the leaves and the leaflets have wrinkled pulvini to permit nastic movements. In some species, leaflets have evolved into tendrils.
Many species have leaves with structures that attract ants that protect the plant from herbivore insects. Extrafloral nectaries are common among the Mimosoideae and the Caesalpinioideae, are found in some Faboideae. In some Acacia, the modified hollow stipules are known as domatia. Many Fabaceae host bacteria in their roots within structures called root nodules; these bacteria, known as rhizobia, have the ability to take nitrogen gas out of the air and convert it to a form of nitrogen, usable to the host plant. This process is called nitrogen fixation; the legume, acting as a host, rhizobia, acting as a provider of usable nitrate, form a symbiotic relationship. The flowers have five fused sepals and five free petals, they are hermaphrodite, have a short hypanthium cup shaped. There are ten stamens and one elongated superior ovary, with a curved style, they are arranged in indeterminate inflorescences. Fabaceae are entomophilous plants, the flowers are showy to attract pollinators. In the Caesalpinioideae, the flowers are zygomorphic, as in Cercis, or nearly symmetrical with five equal petals in Bauhinia.
The upper petal is the innermost one, unlike in the Faboideae. Some species, like some in the genus Senna, have asymmetric flowers, with one of the lower petals larger than the opposing one, the style bent to one side; the calyx, corolla, or stamens can be showy in this group. In the Mimosoideae, the flowers are actinomorphic and arranged in globose inflorescences; the petals are small and the stamens, which can be more than just 10, have long, coloured filaments, which are the showiest part of the flower. All of the flowers in an inflorescence open at once. In the Faboideae, the flowers are zygom
The Fabales are an order of flowering plants included in the rosid group of the eudicots in the Angiosperm Phylogeny Group II classification system. In the APG II circumscription, this order includes the families Fabaceae or legumes, Polygalaceae or milkworts, Surianaceae. Under the Cronquist system and some other plant classification systems, the order Fabales contains only the family Fabaceae. In the classification system of Dahlgren the Fabales were in the superorder Fabiflorae with three familiese corresponding to the subfamilies of Fabaceae in APG II; the other families treated in the Fabales by the APG II classification were placed in separate orders by Cronquist, the Polygalaceae within its own order, the Polygalales, the Quillajaceae and Surianaceae within the Rosales. The Fabaceae, as the third-largest plant family in the world, contain most of the diversity of the Fabales, the other families making up a comparatively small portion of the order's diversity. Research in the order is focused on the Fabaceae, due in part to its great biological diversity, to its importance as food plants.
The Polygalaceae are well researched among plant families, in part due to the large diversity of the genus Polygala, other members of the family being food plants for various Lepidoptera species. While taxonomists using molecular phylogenetic techniques find strong support for the order, questions remain about the morphological relationships of the Quillajaceae and Surianaceae to the rest of the order, due in part to limited research on these families; the Fabales are a cosmopolitan order of plants, except only the subfamily Papilionoideae of the Fabaceae are well dispersed throughout the northern part of the North Temperate Zone
United States Department of Agriculture
The United States Department of Agriculture known as the Agriculture Department, is the U. S. federal executive department responsible for developing and executing federal laws related to farming and food. It aims to meet the needs of farmers and ranchers, promote agricultural trade and production, work to assure food safety, protect natural resources, foster rural communities and end hunger in the United States and internationally. 80% of the USDA's $141 billion budget goes to the Food and Nutrition Service program. The largest component of the FNS budget is the Supplemental Nutrition Assistance Program, the cornerstone of USDA's nutrition assistance; the current Secretary of Agriculture is Sonny Perdue. Many of the programs concerned with the distribution of food and nutrition to people of America and providing nourishment as well as nutrition education to those in need are run and operated under the USDA Food and Nutrition Service. Activities in this program include the Supplemental Nutrition Assistance Program, which provides healthy food to over 40 million low-income and homeless people each month.
USDA is a member of the United States Interagency Council on Homelessness, where it is committed to working with other agencies to ensure these mainstream benefits are accessed by those experiencing homelessness. The USDA is concerned with assisting farmers and food producers with the sale of crops and food on both the domestic and world markets, it plays a role in overseas aid programs by providing surplus foods to developing countries. This aid can go through USAID, foreign governments, international bodies such as World Food Program, or approved nonprofits; the Agricultural Act of 1949, section 416 and Agricultural Trade Development and Assistance Act of 1954 known as Food for Peace, provides the legal basis of such actions. The USDA is a partner of the World Cocoa Foundation. Early in its history, the economy of the United States was agrarian. Officials in the federal government had long sought new and improved varieties of seeds and animals for import into the United States. In 1837 Henry Leavitt Ellsworth, a Yale-educated attorney interested in improving agriculture, became Commissioner of Patents, a position within the Department of State.
He began collecting and distributing new varieties of seeds and plants through members of the Congress and agricultural societies. In 1839, Congress established the Agricultural Division within the Patent Office and allotted $1,000 for "the collection of agricultural statistics and other agricultural purposes." Ellsworth's interest in aiding agriculture was evident in his annual reports that called for a public depository to preserve and distribute the new seeds and plants, a clerk to collect agricultural statistics, statewide reports about crops in different regions, the application of chemistry to agriculture. Ellsworth was called the "Father of the Department of Agriculture."In 1849, the Patent Office was transferred to the newly created Department of the Interior. In the ensuing years, agitation for a separate bureau of agriculture within the department or a separate department devoted to agriculture kept recurring. On May 15, 1862, Abraham Lincoln established the independent Department of Agriculture to be headed by a commissioner without Cabinet status, the agriculturalist Isaac Newton was appointed to be the first such commissioner.
Lincoln called it the "people's department." In 1868, the Department moved into the new Department of Agriculture Building in Washington, D. C. designed by famed DC architect Adolf Cluss. Located on Reservation No.2 on the National Mall between 12th Street and 14th SW, the Department had offices for its staff and the entire width of the Mall up to B Street NW to plant and experiment with plants. In the 1880s, varied advocacy groups were lobbying for Cabinet representation. Business interests sought a Department of Commerce and Industry, farmers tried to raise the Department of Agriculture to Cabinet rank. In 1887, the House of Representatives and Senate passed bills giving Cabinet status to the Department of Agriculture and Labor, but the bill was defeated in conference committee after farm interests objected to the addition of labor. On February 9, 1889, President Grover Cleveland signed a bill into law elevating the Department of Agriculture to Cabinet level. In 1887, the Hatch Act provided for the federal funding of agricultural experiment stations in each state.
The Smith-Lever Act of 1914 funded cooperative extension services in each state to teach agriculture, home economics, other subjects to the public. With these and similar provisions, the USDA reached out to every county of every state. During the Great Depression, farming remained a common way of life for millions of Americans; the Department of Agriculture's Bureau of Home Economics, established in 1923, published shopping advice and recipes to stretch family budgets and make food go farther. USDA helped ensure that food continued to be produced and distributed to those who needed it, assisted with loans for small landowners, contributed to the education of the rural youth, it was revealed on August 27th, 2018 that the U. S. Department of Agriculture would be providing U. S. farmers with a farm aid package, which will total $4.7 billion in direct payments to American farmers. This package is meant to offset the losses farmers are expected to incur from retaliatory tariffs placed on American exports during the Trump tariffs.
The Department of Agriculture was authorized a budget for Fiscal Year 2015 of $139.7 billion. The budget authorization is broken down as follows: Agricultural Stabilization and Conservation Service Animal Damage Control (
Agricultural Research Service
The Agricultural Research Service is the principal in-house research agency of the United States Department of Agriculture. ARS is one of four agencies in USDA's Research and Economics mission area. ARS is charged with extending the nation's scientific knowledge and solving agricultural problems through its four national program areas: nutrition, food safety and quality. ARS research focuses on solving problems affecting Americans every day; the ARS Headquarters is located in the Jamie L. Whitten Building on Independence Avenue in Washington, D. C. and the headquarters staff is located at the George Washington Carver Center in Beltsville, Maryland. For 2018, its budget was $1.2 billion. ARS conducts scientific research for the American public, their main focus is on research to develop solutions to agricultural problems and provide information access and dissemination to: ensure high quality, safe food and other agricultural products, assess the nutritional needs of Americans, sustain a competitive agricultural economy, enhance the natural resource base and the environment, provide economic opportunities to rural citizens and society as a whole.
ARS research complements the work of state colleges and universities, agricultural experiment stations, other federal and state agencies, the private sector. ARS research may focus on regional issues that have national implications, where there is a clear federal role. ARS provides information on its research results to USDA action and regulatory agencies and to several other federal regulatory agencies, including the Food and Drug Administration and the United States Environmental Protection Agency. ARS disseminates much of its research results through scientific journals, technical publications, Agricultural Research magazine, other forums. Information is distributed through ARS's National Agricultural Library. ARS has more than 150 librarians and other information specialists who work at two NAL locations—the Abraham Lincoln Building in Beltsville, Maryland. C. NAL provides reference and information services, document delivery, interlibrary loan and interlibrary borrowing services to a variety of audiences.
ARS supports more than 2,000 scientists and post docs working on 690 research projects within 15 National Programs at more than 90 research locations. The ARS is divided into 5 geographic areas: Midwest Area, Northeast Area, Pacific West Area, Plains Area, Southeast Area. ARS has five major regional research centers: the Western Regional Research Center in Albany, California; the research centers focus on innovation in agricultural practices, pest control and nutrition among other things. Work at these facilities has given life to numerous products and technologies; the ARS offers the Culture Collection, the largest public collection of microorganisms in the world, containing 93,000 strains of bacteria and fungi. The ARS Culture Collection is housed at Northern Regional Research Laboratory ARS' Henry A. Wallace Beltsville Agricultural Research Center in Beltsville, Maryland, is the world's largest agricultural research complex. ARS operates the U. S. Horticultural Research Laboratory in Fort Pierce and the U.
S. National Poultry Research Center in Athens, Georgia. ARS has six major human nutrition research centers that focus on solving a wide spectrum of human nutrition questions by providing authoritative, peer-reviewed, science-based evidence; the centers are located in Arkansas, Texas, North Dakota and California. ARS scientists at these centers study the role of food and dietary components in human health from conception to advanced age. Technology to produce lactose-free milk, ice cream and yogurt was developed by the USDA Agricultural Research Service in 1985; the grape breeding program, which dates back to 1923, developed seedless grapes. The ARS Citrus and Subtropical Products Laboratory in Winter Haven, Florida, is active in work to improve the taste of orange juice concentrate. ARS had a Toxoplasma gondii research program, which experimented on cats infected with the parasite, from 1982 until 2019. Cats were bred for the program and intentionally infected, kittens in the program were euthanized after research was completed.
Cats were fed raw cat and dog meat for the study, called "kitten cannibalism" by the White Coat Waste Project. A bipartisan bill to eliminate the practice was introduced into the House by Representatives Jimmy Panetta, Brian Mast, Elissa Slotkin, Will Hurd, with a companion bill introduced into the Senate by Jeff Merkley; the bills called the "Kittens In Traumatic Testing Ends Now Act of 2019", amend the Animal Welfare Act to limit USDA experimentation on cats. The bill has been referred to the Subcommittee on Livestock and Foreign Agriculture of the House Committee on Agriculture. While the bills have not passed, the USDA stated. Title 7 of the Code of Federal Regulations Agricultural Resource Management Survey Germplasm Resources Information Network Human Nutrition Research Center on Aging National Clonal Germplasm Repository National Agricultural Center and Hall of Fame U. S. Horticultural Research Laboratory National Interagency Confederation for Biological Research "Agricultural Research Service".
Archived from the original on October
The eudicots, Eudicotidae or eudicotyledons are a clade of flowering plants, called tricolpates or non-magnoliid dicots by previous authors. The botanical terms were introduced in 1991 by evolutionary botanist James A. Doyle and paleobotanist Carol L. Hotton to emphasize the evolutionary divergence of tricolpate dicots from earlier, less specialized, dicots; the close relationships among flowering plants with tricolpate pollen grains was seen in morphological studies of shared derived characters. These plants have a distinct trait in their pollen grains of exhibiting three colpi or grooves paralleling the polar axis. Molecular evidence confirmed the genetic basis for the evolutionary relationships among flowering plants with tricolpate pollen grains and dicotyledonous traits; the term means "true dicotyledons", as it contains the majority of plants that have been considered dicots and have characteristics of the dicots. The term "eudicots" has subsequently been adopted in botany to refer to one of the two largest clades of angiosperms, monocots being the other.
The remaining angiosperms include magnoliids and what are sometimes referred to as basal angiosperms or paleodicots, but these terms have not been or adopted, as they do not refer to a monophyletic group. The other name for the eudicots is tricolpates, a name which refers to the grooved structure of the pollen. Members of the group have tricolpate pollen; these pollens have three or more pores set in furrows called colpi. In contrast, most of the other seed plants produce monosulcate pollen, with a single pore set in a differently oriented groove called the sulcus; the name "tricolpates" is preferred by some botanists to avoid confusion with the dicots, a nonmonophyletic group. Numerous familiar plants are eudicots, including many common food plants and ornamentals; some common and familiar eudicots include members of the sunflower family such as the common dandelion, the forget-me-not and other members of its family, buttercup and macadamia. Most leafy trees of midlatitudes belong to eudicots, with notable exceptions being magnolias and tulip trees which belong to magnoliids, Ginkgo biloba, not an angiosperm.
The name "eudicots" is used in the APG system, of 1998, APG II system, of 2003, for classification of angiosperms. It is applied to a monophyletic group, which includes most of the dicots. "Tricolpate" is a synonym for the "Eudicot" monophyletic group, the "true dicotyledons". The number of pollen grain furrows or pores helps classify the flowering plants, with eudicots having three colpi, other groups having one sulcus. Pollen apertures are any modification of the wall of the pollen grain; these modifications include thinning and pores, they serve as an exit for the pollen contents and allow shrinking and swelling of the grain caused by changes in moisture content. The elongated apertures/ furrows in the pollen grain are called colpi, along with pores, are a chief criterion for identifying the pollen classes; the eudicots can be divided into two groups: the basal eudicots and the core eudicots. Basal eudicot is an informal name for a paraphyletic group; the core eudicots are a monophyletic group.
A 2010 study suggested the core eudicots can be divided into two clades, Gunnerales and a clade called "Pentapetalae", comprising all the remaining core eudicots. The Pentapetalae can be divided into three clades: Dilleniales superrosids consisting of Saxifragales and rosids superasterids consisting of Santalales, Berberidopsidales and asteridsThis division of the eudicots is shown in the following cladogram: The following is a more detailed breakdown according to APG IV, showing within each clade and orders: clade Eudicots order Ranunculales order Proteales order Trochodendrales order Buxales clade Core eudicots order Gunnerales order Dilleniales clade Superrosids order Saxifragales clade Rosids order Vitales clade Fabids order Fabales order Rosales order Fagales order Cucurbitales order Oxalidales order Malpighiales order Celastrales order Zygophyllales clade Malvids order Geraniales order Myrtales order Crossosomatales order Picramniales order Malvales order Brassicales order Huerteales order Sapindales clade Superasterids order Berberidopsidales order Santalales order Caryophyllales clade Asterids order Cornales order Ericales clade Campanulids order Aquifoliales order Asterales order Escalloniales order Bruniales order Apiales order Dipsacales order Paracryphiales clade Lamiids order Solanales order Lamiales order Vahliales order Gentianales order Boraginales order Garryales order Metteniusales order Icacinales Eudicots at the Encyclopedia of Life Eudicots, Tree of Life Web Project Dicots Plant Life Forms
The rosids are members of a large clade of flowering plants, containing about 70,000 species, more than a quarter of all angiosperms. The clade is divided into 16 to 20 orders, depending upon circumscription and classification; these orders, in turn, together comprise about 140 families. Fossil rosids are known from the Cretaceous period. Molecular clock estimates indicate that the rosids originated in the Aptian or Albian stages of the Cretaceous, between 125 and 99.6 million years ago. The name is based upon the name "Rosidae", understood to be a subclass. In 1967, Armen Takhtajan showed that the correct basis for the name "Rosidae" is a description of a group of plants published in 1830 by Friedrich Gottlieb Bartling; the clade was renamed "Rosidae" and has been variously delimited by different authors. The name "rosids" is informal and not assumed to have any particular taxonomic rank like the names authorized by the ICBN; the rosids are monophyletic based upon evidence found by molecular phylogenetic analysis.
Three different definitions of the rosids were used. Some authors included the orders Vitales in the rosids. Others excluded both of these orders; the circumscription used in this article is that of the APG IV classification, which includes Vitales, but excludes Saxifragales. The rosids and Saxifragales form the superrosids clade; this is one of three groups that compose the Pentapetalae, the others being Dilleniales and the superasterids. The rosids consist of two groups: the eurosids; the eurosids, in turn, are divided into two groups: malvids. The rosids consist of 17 orders. In addition to Vitales, there are 8 orders in malvids; some of the orders have only been recognized. These are Vitales, Crossosomatales and Huerteales; the phylogeny of Rosids shown below is adapted from the Angiosperm Phylogeny Group website. The nitrogen-fixing clade contains a high number of actinorhizal plants. Not all plants in this clade are actinorhizal, however. Media related to Rosids at Wikimedia Commons
Vicia is a genus of about 140 species of flowering plants that are part of the legume family, which are known as vetches. Member species are native to Europe, North America, South America and Africa; some other genera of their subfamily Faboideae have names containing "vetch", for example the vetchlings or the milk-vetches. The broad bean is sometimes separated in a monotypic genus Faba; the tribe Vicieae in which the vetches are placed is named after the genus' current name. Among the closest living relatives of vetches are the true peas. Bitter vetch was one of the first domesticated crops, it was grown in the Near East about 9,500 years ago, starting even one or two millennia earlier during the Pre-Pottery Neolithic A. By the time of the Central European Linear Pottery culture – about 7,000 years ago – broad bean had been domesticated. Vetch has been found at Neolithic and Eneolithic sites in Bulgaria and Slovakia, and at the same time, at the opposite end of Eurasia, the Hoabinhian people utilized the broad bean in their path towards agriculture, as shown by the seeds found in Spirit Cave, Thailand.
Bernard of Clairvaux shared a bread-of-vetch meal with his monks during the famine of 1124 to 1126, as an emblem of humility. However, Bitter Vetch was dropped from human use over time, it was only used to save as a crop of last resort in times of starvation: vetches "featured in the frugal diet of the poor until the eighteenth century, reappeared on the black market in the South of France during the Second World War", Maguelonne Toussaint-Samat, of Marseillais background, has remarked. However, broad beans remained prominent. In the Near East the seeds are mentioned in Hittite and Ancient Egyptian sources dating from more than 3,000 years ago as well as in the Christian Bible, in the large Celtic Oppidum of Manching from the La Tène culture in Europe some 2,200 years ago. Dishes resembling ful medames are attested in the Jerusalem Talmud, compiled before 400 AD. In our time, the common vetch has risen to prominence. Together with broad bean cultivars such as horse bean or field bean, the FAO includes it among the 11 most important pulses in the world.
The main usage of the common vetch is as forage for ruminant animals, both as fodder and legume, but there are other uses, as tufted vetch, V. cracca is grown as a mid-summer pollen source for honeybees. The bitter vetch, too, is grown extensively for forage and fodder, as are hairy vetch, bard vetch, French vetch and Narbon bean. V. benghalensis and Hungarian vetch are cultivated for green manure. The vetches have a broad variety of other purposes; the Hairy Vetch has well-established uses as an allelopathic cover crop. As regards the broad bean, it is known to accumulate aluminum in its tissue; the robust plants are useful as a beetle bank to provide habitat and shelter for carnivorous beetles and other arthropods to keep down pest invertebrates. When the root nodules of broad bean are inoculated with the rhodospirillacean bacterium Azospirillum brasilense and the glomeracean fungus Glomus clarum, the species can be productively grown in salty soils. In the 1980s, the auxin 4-Cl-IAA was studied in V. amurensis and the broad bean, since 1990, the antibacterial γ-thionins fabatin-1 and -2 have been isolated from the latter species.
Despite a small chromosome count of n=6, the broad bean has a high DNA content, making it easy for a micronucleus test of its root tips to recognize genotoxic compounds. A lectin from V. graminea is used to test for the medically significant N blood group. The vetches grown as forage are toxic to non-ruminants, at least if eaten in quantity. Cattle and horses have been poisoned by V. villosa and V. benghalensis, two species that contain canavanine in their seeds. Canavanine, a toxic analogue of the amino acid arginine, has been identified in Hairy Vetch as an appetite suppressant for monogastric animals, while Narbon bean contains the quicker-acting but weaker γ-glutamyl-S-ethenylcysteine. In common vetch, γ-glutamyl-β-cyanoalanine has been found; the active part of this molecule is β-cyanoalanine. It inhibits the conversion of the sulfur amino acid methionine to cysteine. Cystathionine, an intermediary product of this biochemical pathway, is secreted in urine; this process can lead to the depletion of vital protective reserves of the sulfur amino acid cysteine and thereby making Vicia sativa seed a dangerous component in mixture with other toxin sources.
The Spanish pulse mix comuña contains common vetch and bitter vetch in addition to vetchling seeds. Moreover, common vetch as well as broad bean – and other species of Vicia too – contain oxidants like convicine, isouramil and vicine in quantities sufficient to lower glutathione levels in G6PD-deficient persons to cause favism disease. At least broad beans contain the lectin phytohemagglutinin and are somewhat poisonous if eaten raw. Split common vetch seeds resemble split red lentils, has been mislabelled as such by exporters or importers to be sold for human consumption. I