Binomial nomenclature called binominal nomenclature or binary nomenclature, is a formal system of naming species of living things by giving each a name composed of two parts, both of which use Latin grammatical forms, although they can be based on words from other languages. Such a name is called a binomen, binominal name or a scientific name; the first part of the name – the generic name – identifies the genus to which the species belongs, while the second part – the specific name or specific epithet – identifies the species within the genus. For example, humans belong within this genus to the species Homo sapiens. Tyrannosaurus rex is the most known binomial; the formal introduction of this system of naming species is credited to Carl Linnaeus beginning with his work Species Plantarum in 1753. But Gaspard Bauhin, in as early as 1623, had introduced in his book Pinax theatri botanici many names of genera that were adopted by Linnaeus; the application of binomial nomenclature is now governed by various internationally agreed codes of rules, of which the two most important are the International Code of Zoological Nomenclature for animals and the International Code of Nomenclature for algae and plants.
Although the general principles underlying binomial nomenclature are common to these two codes, there are some differences, both in the terminology they use and in their precise rules. In modern usage, the first letter of the first part of the name, the genus, is always capitalized in writing, while that of the second part is not when derived from a proper noun such as the name of a person or place. Both parts are italicized when a binomial name occurs in normal text, thus the binomial name of the annual phlox is now written as Phlox drummondii. In scientific works, the authority for a binomial name is given, at least when it is first mentioned, the date of publication may be specified. In zoology "Patella vulgata Linnaeus, 1758"; the name "Linnaeus" tells the reader who it was that first published a description and name for this species of limpet. "Passer domesticus". The original name given by Linnaeus was Fringilla domestica; the ICZN does not require that the name of the person who changed the genus be given, nor the date on which the change was made, although nomenclatorial catalogs include such information.
In botany "Amaranthus retroflexus L." – "L." is the standard abbreviation used in botany for "Linnaeus". "Hyacinthoides italica Rothm. – Linnaeus first named this bluebell species Scilla italica. The name is composed of two word-forming elements: "bi", a Latin prefix for two, "-nomial", relating to a term or terms; the word "binomium" was used in Medieval Latin to mean a two-term expression in mathematics. Prior to the adoption of the modern binomial system of naming species, a scientific name consisted of a generic name combined with a specific name, from one to several words long. Together they formed a system of polynomial nomenclature; these names had two separate functions. First, to designate or label the species, second, to be a diagnosis or description. In a simple genus, containing only two species, it was easy to tell them apart with a one-word genus and a one-word specific name; such "polynomial names" may sometimes look like binomials, but are different. For example, Gerard's herbal describes various kinds of spiderwort: "The first is called Phalangium ramosum, Branched Spiderwort.
The other... is aptly termed Phalangium Ephemerum Virginianum, Soon-Fading Spiderwort of Virginia". The Latin phrases are short descriptions, rather than identifying labels; the Bauhins, in particular Caspar Bauhin, took some important steps towards the binomial system, by pruning the Latin descriptions, in many cases to two words. The adoption by biologists of a system of binomial nomenclature is due to Swedish botanist and physician Carl von Linné, more known by his Latinized name Carl Linnaeus, it was in his 1753 Species Plantarum that he first began using a one-word "trivial name" together with a generic name in a system of binomial nomenclature. This trivial name is what is now known as specific name; the Bauhins' genus names were retained in many of these, but the descriptive part was reduced to a single word. Linnaeus's trivial names introduced an important new idea, namely that the function of a name could be to give a species a unique label; this meant. Thus Gerard's Phalangium ephemerum virginianum became Tradescantia virgi
Tanzania the United Republic of Tanzania, is a country in eastern Africa within the African Great Lakes region. It borders Uganda to the north. Mount Kilimanjaro, Africa's highest mountain, is in north-eastern Tanzania; the first humans known lived in Pliocene Tanzania 6 million years ago. The genus Australopithecus ranged all over Africa 4-2 million years ago. Following the rise of Homo erectus 1.8 million years ago, mankind spread all over the Old World, in the New World and Australia under the species Homo sapiens. Homo sapiens overtook Africa and absorbed the older archaic species and subspecies of humanity. One of the oldest known ethnic groups still existing, the Hadzabe, appears to have originated in Tanzania, their oral history recalls ancestors who were tall and were the first to use fire and lived in caves, much like Homo erectus or Homo heidelbergensis who lived in the same region before them. In the Stone and Bronze Age, prehistoric migrations into Tanzania included Southern Cushitic speakers who moved south from present-day Ethiopia.
These movements took place at about the same time as the settlement of the Mashariki Bantu from West Africa in the Lake Victoria and Lake Tanganyika areas. They subsequently migrated across the rest of Tanzania between 1,700 years ago. European colonialism began in mainland Tanzania during the late 19th century when Germany formed German East Africa, which gave way to British rule following World War I; the mainland was governed as Tanganyika, with the Zanzibar Archipelago remaining a separate colonial jurisdiction. Following their respective independence in 1961 and 1963, the two entities merged in April 1964 to form the United Republic of Tanzania; the United Nations estimated Tanzania's 2016 population at 55.57 million. The population is composed of several ethnic and religious groups; the sovereign state of Tanzania is a presidential constitutional republic and since 1996 its official capital city has been Dodoma where the president's office, the National Assembly, some government ministries are located.
Dar es Salaam, the former capital, retains most government offices and is the country's largest city, principal port, leading commercial centre. Tanzania is a de facto one-party state with the democratic socialist Chama Cha Mapinduzi party in power. Tanzania is densely forested in the north-east, where Mount Kilimanjaro is located. Three of Africa's Great Lakes are within Tanzania. To the north and west lie Lake Victoria, Africa's largest lake, Lake Tanganyika, the continent's deepest lake, known for its unique species of fish. To the south lies Lake Malawi; the eastern shore is humid, with the Zanzibar Archipelago just offshore. The Menai Bay Conservation Area is Zanzibar's largest marine protected area; the Kalambo Falls, located on the Kalambo River at the Zambian border, is the second highest uninterrupted waterfall in Africa. Over 100 different languages are spoken in Tanzania, making it the most linguistically diverse country in East Africa; the country does not have a de jure official language.
Swahili is used in parliamentary debate, in the lower courts, as a medium of instruction in primary school. English is used in foreign trade, in diplomacy, in higher courts, as a medium of instruction in secondary and higher education, although the Tanzanian government is planning to discontinue English as a language of instruction altogether. 10 percent of Tanzanians speak Swahili as a first language, up to 90 percent speak it as a second language. The name "Tanzania" was created as a clipped compound of the names of the two states that unified to create the country: Tanganyika and Zanzibar, it comprises the first three letters of the two states, "Tan" and "Zan" as well as the only two vowels in the names of two states, "I" and "a" to form Tanzania. The name "Tanganyika" is derived from the Swahili words tanga and nyika, creating the phrase "sail in the wilderness", it is sometimes understood as a reference to Lake Tanganyika. The name of Zanzibar comes from "zenji", the name for a local people, the Arabic word "barr", which means coast or shore.
The indigenous populations of eastern Africa are thought to be the linguistically isolated Hadza and Sandawe hunter-gatherers of Tanzania. The first wave of migration was by Southern Cushitic speakers who moved south from Ethiopia and Somalia into Tanzania, they are ancestral to the Iraqw and Burunge. Based on linguistic evidence, there may have been two movements into Tanzania of Eastern Cushitic people at about 4,000 and 2,000 years ago, originating from north of Lake Turkana. Archaeological evidence supports the conclusion that Southern Nilotes, including the Datoog, moved south from the present-day South Sudan / Ethiopia border region into central northern Tanzania between 2,900 and 2,400 years ago; these movements took place at the same time as the settlement of the iron-making Mashariki Bantu from West Africa in the Lake Victoria and Lake Tanganyika areas. They brought with them the west African planting tradition and the p
Titanosaurs were a diverse group of sauropod dinosaurs which included Saltasaurus and Isisaurus. The titanosaurs were the last surviving group of long-necked sauropods, with taxa still thriving at the time of the extinction event at the end of the Cretaceous; the group includes the largest land animals known to have existed, such as Patagotitan—estimated at 37 m long with a weight of 69 tonnes —and the comparably sized Argentinosaurus and Puertasaurus from the same region. The group's name alludes to the mythological Titans of Ancient Greece, via the type genus Titanosaurus. Together with the brachiosaurids and relatives, titanosaurs make up the larger clade Titanosauriformes. Titanosaurs had small heads when compared with other sauropods; the head was wide, similar to the heads of Camarasaurus and Brachiosaurus but more elongated. Their nostrils were large and they all had crests formed by these nasal bones, their teeth were either somewhat spatulate or like pegs or pencils, but were always small.
Their necks were of average length, for sauropods, their tails were whip-like, but not as long as in the diplodocids. While the pelvis was slimmer than some sauropods, the pectoral was much wider, giving them a uniquely'wide-gauged' stance; as a result, the fossilised trackways of titanosaurs are distinctly broader than other sauropods. Their forelimbs were stocky, longer than their hind limbs, their vertebrae were solid. Their spinal column was more flexible, so they were more agile than their cousins and better at rearing up. Unlike other sauropods, some titanosaurs had no digits or digit bones, walked only on horseshoe-shaped "stumps" made up of the columnar metacarpal bones. From skin impressions found with the fossils, it has been determined that the skin of many titanosaur species was armored with a small mosaic of small, bead-like scales around a larger scale. One species, has been discovered with bony plates, like the ankylosaurs. Studies published in 2011 indicate that titanosaurs such as Rapetosaurus, may have used the osteoderms common in the various genera for storing minerals during harsh changes in climate, such as drought.
While they were all huge, many were average in size compared with the other giant dinosaurs. There were some island-dwelling dwarf species such as Magyarosaurus the result of allopatric speciation and insular dwarfism; the family Titanosauridae was once used for derived titanosaurs, but Wilson and Upchurch found the type genus Titanosaurus dubious based on the figures and original description. Weishampel et al. in the second edition of The Dinosauria did not use the family Titanosauridae, instead used several smaller titanosaur families such as Saltasauridae and Nemegtosauridae, coining Lithostrotia for derived titanosaurs. A handful of Argentine sauropod workers, continue to use Titanosauridae for titanosaurs now placed in Lithostrotia. In the second edition of The Dinosauria, the clade Titanosauria was defined as all sauropods closer to Saltasaurus than to Brachiosaurus. Subsequent cladistic analyses have defined Titanosauria as including Saltasaurus but not Euhelopus or Brachiosaurus. Relationships within the Titanosauria have been variable from study to study, complicated by the fact that clade and rank names have been applied inconsistently by various scientists.
One possible cladogram is presented here, follows a 2007 analysis by Calvo and colleagues. The authors notably used the family Titanosauridae in a broader fashion than other recent studies, coined the new clade name Lognkosauria. In the description of Mansourasaurus, Sallam et al. published a phylogenetic analysis of Titanosauria including the most taxa of any analysis of the clade. The relationships within Titanosauria can be seen below. Fossilized dung associated with late Cretaceous titanosaurids has revealed phytoliths, silicified plant fragments, that offer clues to a broad, unselective plant diet. Besides the plant remains that might have been expected, such as cycads and conifers, discoveries published in 2005 revealed an unexpectedly wide range of monocotyledons, including palms and grasses, including ancestors of rice and bamboo, which has given rise to speculation that herbivorous dinosaurs and grasses co-evolved. A large titanosaurid nesting ground was discovered in Auca Mahuevo, in Patagonia and another colony has been discovered in Spain.
Several hundred female saltasaurs dug holes with their back feet, laid eggs in clutches averaging around 25 eggs each, buried the nests under dirt and vegetation. The small eggs, about 11–12 centimetres in diameter, contained fossilised embryos, complete with skin impressions; the impressions showed. The huge number of individuals gives evidence of herd behavior, along with their armor, could have helped provide protection against large predators such as Abelisaurus; the titanosaurs were the last great group of sauropods, which existed from about 136 to 66 million years ago, before the Cretaceous–Paleogene extinction event, were the dominant herbivores of their time. The fossil evidence suggests they replaced the other sauropods, like the diplodocids and the brachiosaurids, which died out between the late Jurassic and the mid-Cretaceous Periods. Titanosaurs were widespread. In December 2011, Argentine scientists announced titanosaur fossils had been found on Antarctica—meaning that titanosaur fossils
Lusotitan is a genus of herbivorous brachiosaurid sauropod dinosaur from the Late Jurassic Period of Portugal. In 1947 Manuel de Matos, a member of the Geological Survey of Portugal, discovered large sauropod fossils in the Portuguese Lourinhã Formation that date back to the Tithonian stage of the Late Jurassic period. In 1957 Albert-Félix de Lapparent and Georges Zbyszewski named the remains as a new species of Brachiosaurus: Brachiosaurus atalaiensis; the specific name referred to the site Atalaia. In 2003 Octávio Mateus and Miguel Telles Antunes named it as a separate genus: Lusotitan; the type species is Lusotitan atalaiensis. The generic name is derived from Luso, the Latin name for an inhabitant of Lusitania, from the Greek word "Titan", a mythological giant; the finds consisted of a partial skeleton lacking the skull and individual vertebrae uncovered in several locations. De Lapparent did not assign a holotype. In 2003 Mateus chose the skeleton as the lectotype, its bones have the inventory numbers MIGM 4798, 4801–10, 4938, 4944, 4950, 4952, 4958, 4964–6, 4981–2, 4985, 8807, 8793-5.
These remains include elements of the appendicular skeleton. It has been estimated, it had long one of the reasons Mateus assigned it to the Brachiosauridae. The lectotype was re-described by Mannion and colleagues in 2013; the Lourinhã Formation of western Portugal was to be formed during the Kimmeridgian or Tithonian ages of the Late Jurassic period. The area is a coastal region with a strong marine influence, its flora and fauna are similar to the Morrison Formation in the United States, the Tendaguru Formation in Tanzania. Lusotitan is the largest dinosaur, discovered in the area. Lusotitan lived alongside species of the predatory theropods Allosaurus, Ceratosaurus and Torvosaurus, the ankylosaurian Dracopelta, the sauropods Supersaurus and Zby, the stegosaurs Dacentrurus and Miragaia
Saurischia is one of the two basic divisions of dinosaurs. ‘Saurischia’ translates to lizard-hipped. In 1888, Harry Seeley classified dinosaurs into two orders, based on their hip structure, though today most paleontologists classify Saurischia as an unranked clade rather than an order. All carnivorous dinosaurs are traditionally classified as saurischians, as are all of the birds and one of the two primary lineages of herbivorous dinosaurs, the sauropodomorphs. At the end of the Cretaceous Period, all saurischians except the birds became extinct in the course of the Cretaceous–Paleogene extinction event. Birds, as direct descendants of one group of theropod dinosaurs, are a sub-clade of saurischian dinosaurs in phylogenetic classification. Saurischian dinosaurs are traditionally distinguished from ornithischian dinosaurs by their three-pronged pelvic structure, with the pubis pointed forward; the ornithischians' pelvis is arranged with the pubis rotated backward, parallel with the ischium also with a forward-pointing process, giving a four-pronged structure.
The saurischian hip structure led Seeley to name them "lizard-hipped" dinosaurs, because they retained the ancestral hip anatomy found in modern lizards and other reptiles. He named ornithischians "bird-hipped" dinosaurs because their hip arrangement was superficially similar to that of birds, though he did not propose any specific relationship between ornithischians and birds. However, in the view which has long been held, this "bird-hipped" arrangement evolved several times independently in dinosaurs, first in the ornithischians in the lineage of saurischians including birds, lastly in the therizinosaurians; this would be an example of convergent evolution, therizinosaurians, ornithischian dinosaurs all developed a similar hip anatomy independently of each other as an adaptation to their herbivorous or omnivorous diets. In his paper naming the two groups, Seeley reviewed previous classification schemes put forth by other paleontologists to divide up the traditional order Dinosauria, he preferred one, put forward by Othniel Charles Marsh in 1878, which divided dinosaurs into four orders: Sauropoda, Theropoda and Stegosauria.
Seeley, wanted to formulate a classification that would take into account a single primary difference between major dinosaurian groups based on a characteristic that differentiated them from other reptiles. He found this in the configuration of the hip bones, found that all four of Marsh's orders could be divided neatly into two major groups based on this feature, he placed the Stegosauria and Ornithopoda in the Ornithischia, the Theropoda and Sauropoda in the Saurischia. Furthermore, Seeley used this major difference in the hip bones, along with many other noted differences between the two groups, to argue that "dinosaurs" were not a natural grouping at all, but rather two distinct orders that had arisen independently from more primitive archosaurs; this concept that "dinosaur" was an outdated term for two distinct orders lasted many decades in the scientific and popular literature, it was not until the 1960s that scientists began to again consider the possibility that saurischians and ornithischians were more related to each other than they were to other archosaurs.
Although his concept of a polyphyletic Dinosauria is no longer accepted by most paleontologists, Seeley's basic division of the two dinosaurian groups has stood the test of time, has been supported by modern cladistic analysis of relationships among dinosaurs. One alternative hypothesis challenging Seeley's classification was proposed by Robert T. Bakker in his 1986 book The Dinosaur Heresies. Bakker's classification separated the theropods into their own group and placed the two groups of herbivorous dinosaurs together in a separate group he named the Phytodinosauria; the Phytodinosauria hypothesis was based on the supposed link between ornithischians and prosauropods, the idea that the former had evolved directly from the latter by way of an enigmatic family that seemed to possess characters of both groups, the segnosaurs. However, it was found that segnosaurs were an unusual type of herbivorous theropod saurischian related to birds, the Phytodinosauria hypothesis fell out of favor. A 2017 study by Dr Matthew Grant Baron, Dr David B. Norman and Prof. Paul M. Barrett did not find support for a monophyletic Saurischia, according to its traditional definition.
Instead, the group was found to be paraphyletic, with Theropoda removed from the group and placed as the sister group to the Ornithischia in the newly defined clade Ornithoscelida. As a result, the authors redefined Saurischia as "the most inclusive clade that contains D. carnegii, but not T. horridus", resulting in a clade containing only the Sauropodomorpha and Herrerasauridae
The Triassic is a geologic period and system which spans 50.6 million years from the end of the Permian Period 251.9 million years ago, to the beginning of the Jurassic Period 201.3 Mya. The Triassic is the shortest period of the Mesozoic Era. Both the start and end of the period are marked by major extinction events. Triassic began in the wake of the Permian–Triassic extinction event, which left the Earth's biosphere impoverished. Therapsids and archosaurs were the chief terrestrial vertebrates during this time. A specialized subgroup of archosaurs, called dinosaurs, first appeared in the Late Triassic but did not become dominant until the succeeding Jurassic Period; the first true mammals, themselves a specialized subgroup of therapsids evolved during this period, as well as the first flying vertebrates, the pterosaurs, like the dinosaurs, were a specialized subgroup of archosaurs. The vast supercontinent of Pangaea existed until the mid-Triassic, after which it began to rift into two separate landmasses, Laurasia to the north and Gondwana to the south.
The global climate during the Triassic was hot and dry, with deserts spanning much of Pangaea's interior. However, the climate became more humid as Pangaea began to drift apart; the end of the period was marked by yet another major mass extinction, the Triassic–Jurassic extinction event, that wiped out many groups and allowed dinosaurs to assume dominance in the Jurassic. The Triassic was named in 1834 by Friedrich von Alberti, after the three distinct rock layers that are found throughout Germany and northwestern Europe—red beds, capped by marine limestone, followed by a series of terrestrial mud- and sandstones—called the "Trias"; the Triassic is separated into Early and Late Triassic Epochs, the corresponding rocks are referred to as Lower, Middle, or Upper Triassic. The faunal stages from the youngest to oldest are: During the Triassic all the Earth's land mass was concentrated into a single supercontinent centered more or less on the equator and spanning from pole to pole, called Pangaea.
From the east, along the equator, the Tethys sea penetrated Pangaea, causing the Paleo-Tethys Ocean to be closed. In the mid-Triassic a similar sea penetrated along the equator from the west; the remaining shores were surrounded by the world-ocean known as Panthalassa. All the deep-ocean sediments laid down during the Triassic have disappeared through subduction of oceanic plates; the supercontinent Pangaea was rifting during the Triassic—especially late in that period—but had not yet separated. The first nonmarine sediments in the rift that marks the initial break-up of Pangaea, which separated New Jersey from Morocco, are of Late Triassic age. S. these thick sediments comprise the Newark Group. Because a super-continental mass has less shoreline compared to one broken up, Triassic marine deposits are globally rare, despite their prominence in Western Europe, where the Triassic was first studied. In North America, for example, marine deposits are limited to a few exposures in the west, thus Triassic stratigraphy is based on organisms that lived in lagoons and hypersaline environments, such as Estheria crustaceans.
At the beginning of the Mesozoic Era, Africa was joined with Earth's other continents in Pangaea. Africa shared the supercontinent's uniform fauna, dominated by theropods and primitive ornithischians by the close of the Triassic period. Late Triassic fossils are more common in the south than north; the time boundary separating the Permian and Triassic marks the advent of an extinction event with global impact, although African strata from this time period have not been studied. During the Triassic peneplains are thought to have formed in what is now southern Sweden. Remnants of this peneplain can be traced as a tilted summit accordance in the Swedish West Coast. In northern Norway Triassic peneplains may have been buried in sediments to be re-exposed as coastal plains called strandflats. Dating of illite clay from a strandflat of Bømlo, southern Norway, have shown that landscape there became weathered in Late Triassic times with the landscape also being shaped during that time. At Paleorrota geopark, located in Rio Grande do Sul, the Santa Maria Formation and Caturrita Formations are exposed.
In these formations, one of the earliest dinosaurs, Staurikosaurus, as well as the mammal ancestors Brasilitherium and Brasilodon have been discovered. The Triassic continental interior climate was hot and dry, so that typical deposits are red bed sandstones and evaporites. There is no evidence of glaciation near either pole. Pangaea's large size limited the moderating effect of the global ocean; the strong contrast between the Pangea supercontinent and the global ocean triggered intense cross-equatorial monsoons. The Triassic may have been a dry period, but evidence exists that it was punctuated by several episodes of increased rainfall in tropical and subtropical latitudes of the Tethys Sea and its surrounding land. Sediments and fossils suggestive of a more humid climate are known from the Anisian to Ladinian of the Tethysian domain, from the Carnian and Rhaetian of a larger area that includes the Boreal domain, the North