Sharks are a group of elasmobranch fish characterized by a cartilaginous skeleton, five to seven gill slits on the sides of the head, and pectoral fins that are not fused to the head. Modern sharks are classified within the clade Selachimorpha and are the sister group to the Batoidea. Some sources extend the term "shark" as an informal category including extinct members of Chondrichthyes with a shark-like morphology, such as hybodonts. Shark-like chondrichthyans such as Cladoselache and Doliodus first appeared in the Devonian Period, though some fossilized chondrichthyan-like scales are as old as the Late Ordovician. The oldest confirmed modern sharks (selachimorphs) are known from the Early Jurassic, about 200 million years ago, though records of true sharks may extend back as far as the Permian.

Elasmobranchii is a subclass of Chondrichthyes or cartilaginous fish, including modern sharks, rays, skates, and sawfish. Members of this subclass are characterised by having five to seven pairs of gill clefts opening individually to the exterior, rigid dorsal fins and small placoid scales on the skin. The teeth are in several series; the upper jaw is not fused to the cranium, and the lower jaw is articulated with the upper. The details of this jaw anatomy vary between species, and help distinguish the different elasmobranch clades. The pelvic fins in males are modified to create claspers for the transfer of sperm. There is no swim bladder; instead, these fish maintain buoyancy with large livers rich in oil.