1.
Precambrian
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The Precambrian is the earliest period of Earths history, set before the current Phanerozoic Eon. The Precambrian is a supereon that is subdivided into three eons of the time scale. It spans from the formation of Earth about 4.6 billion years ago to the beginning of the Cambrian Period, about 541 million years ago, the Precambrian accounts for 89% of geologic time. Relatively little is known about the Precambrian, despite it making up roughly seven-eighths of the Earths history, the Precambrian fossil record is poorer than that of the succeeding Phanerozoic, and fossils from that time are of limited biostratigraphic use. This is because many Precambrian rocks have been metamorphosed, obscuring their origins, while others have been destroyed by erosion. A stable crust was apparently in place by 4,412 Ma, the term Precambrian is recognized by the International Commission on Stratigraphy as a general term including the Archean and Proterozoic eons. It is still used by geologists and paleontologists for general discussions not requiring the more specific eon names and it was briefly called the Cryptozoic eon. A specific date for the origin of life has not been determined, carbon found in 3.8 billion year old rocks from islands off western Greenland may be of organic origin. Well-preserved microscopic fossils of bacteria older than 3.46 billion years have found in Western Australia. Probable fossils 100 million years older have been found in the same area, there is a fairly solid record of bacterial life throughout the remainder of the Precambrian. The oldest fossil evidence from that era of such complex life comes from the Lantian formation of the Ediacarian period, a very diverse collection of soft-bodied forms is found in a variety of locations worldwide and date to between 635 and 542 Ma. These are referred to as Ediacaran or Vendian biota, hard-shelled creatures appeared toward the end of that time span, marking the beginning of the Phanerozoic era. By the middle of the following Cambrian period, a diverse fauna is recorded in the Burgess Shale. The explosion in diversity of lifeforms during the early Cambrian is called the Cambrian explosion of life, while land seems to have been devoid of plants and animals, cyanobacteria and other microbes formed prokaryotic mats that covered terrestrial areas. Evidence of the details of plate motions and other activity in the Precambrian has been poorly preserved. It is generally believed that small proto-continents existed prior to 4280 Ma, the supercontinent, known as Rodinia, broke up around 750 Ma. A number of glacial periods have been identified going as far back as the Huronian epoch, one of the best studied is the Sturtian-Varangian glaciation, around 850–635 Ma, which may have brought glacial conditions all the way to the equator, resulting in a Snowball Earth. The atmosphere of the early Earth is not well understood, most geologists believe it was composed primarily of nitrogen, carbon dioxide, and other relatively inert gases, and was lacking in free oxygen
2.
Cambrian
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The Cambrian Period was the first geological period of the Paleozoic Era, of the Phanerozoic Eon. The Cambrian lasted 55.6 million years from the end of the preceding Ediacaran Period 541 million years ago to the beginning of the Ordovician Period 485.4 mya and its subdivisions, and its base, are somewhat in flux. The period was established by Adam Sedgwick, who named it after Cambria, the Latinised form of Cymru, the Welsh name for Wales, as a result, our understanding of the Cambrian biology surpasses that of some later periods. The rapid diversification of lifeforms in the Cambrian, known as the Cambrian explosion, most of the continents were probably dry and rocky due to a lack of vegetation. Shallow seas flanked the margins of several continents created during the breakup of the supercontinent Pannotia, the seas were relatively warm, and polar ice was absent for much of the period. The United States Federal Geographic Data Committee uses a barred capital C ⟨Є⟩ character similar to the capital letter Ukrainian Ye ⟨Є⟩ to represent the Cambrian Period, the proper Unicode character is U+A792 Ꞓ LATIN CAPITAL LETTER C WITH BAR. Despite the long recognition of its distinction from younger Ordovician Period rocks and older Supereon Precambrian rocks, the base of the Cambrian lies atop a complex assemblage of trace fossils known as the Treptichnus pedum assemblage. Pedum in Namibia, Spain and Newfoundland, and possibly, in the western USA, the stratigraphic range of T. pedum overlaps the range of the Ediacaran fossils in Namibia, and probably in Spain. The Cambrian Period followed the Ediacaran Period and was followed by the Ordovician Period, the Cambrian is divided into four epochs and ten ages. Currently only two series and five stages are named and have a GSSP, because the international stratigraphic subdivision is not yet complete, many local subdivisions are still widely used. In some of these subdivisions the Cambrian is divided into three epochs with locally differing names – the Early Cambrian, Middle Cambrian and Furongian, rocks of these epochs are referred to as belonging to the Lower, Middle, or Upper Cambrian. Trilobite zones allow biostratigraphic correlation in the Cambrian, each of the local epochs is divided into several stages. The International Commission on Stratigraphy list the Cambrian period as beginning at 541 million years ago, the lower boundary of the Cambrian was originally held to represent the first appearance of complex life, represented by trilobites. The recognition of small shelly fossils before the first trilobites, and Ediacara biota substantially earlier and this formal designation allowed radiometric dates to be obtained from samples across the globe that corresponded to the base of the Cambrian. Early dates of 570 million years ago quickly gained favour, though the used to obtain this number are now considered to be unsuitable. A more precise date using modern radiometric dating yield a date of 541 ±0.3 million years ago, most continental land was clustered in the Southern Hemisphere at this time, but was drifting north. Large, high-velocity rotational movement of Gondwana appears to have occurred in the Early Cambrian, the sea levels fluctuated somewhat, suggesting there were ice ages, associated with pulses of expansion and contraction of a south polar ice cap. In Baltoscandia a Lower Cambrian transgression transformed large swathes of the Sub-Cambrian peneplain into a epicontinental sea, the Earth was generally cold during the early Cambrian, probably due to the ancient continent of Gondwana covering the South Pole and cutting off polar ocean currents
3.
Ordovician
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The Ordovician is a geologic period and system, the second of six periods of the Paleozoic Era. The Ordovician spans 41.2 million years from the end of the Cambrian Period 485.4 million years ago to the start of the Silurian Period 443.8 Mya. Lapworth recognized that the fauna in the disputed strata were different from those of either the Cambrian or the Silurian periods. It received international sanction in 1960, when it was adopted as a period of the Paleozoic Era by the International Geological Congress. Life continued to flourish during the Ordovician as it did in the earlier Cambrian period, invertebrates, namely molluscs and arthropods, dominated the oceans. The Great Ordovician Biodiversification Event considerably increased the diversity of life, fish, the worlds first true vertebrates, continued to evolve, and those with jaws may have first appeared late in the period. Life had yet to diversify on land, about 100 times as many meteorites struck the Earth during the Ordovician compared with today. The Ordovician Period began with a major extinction called the Cambrian–Ordovician extinction event and it lasted for about 42 million years and ended with the Ordovician–Silurian extinction event, about 443.8 Mya which wiped out 60% of marine genera. The dates given are recent radiometric dates and vary slightly from those found in other sources and this second period of the Paleozoic era created abundant fossils that became major petroleum and gas reservoirs. The boundary chosen for the beginning of both the Ordovician Period and the Tremadocian stage is highly significant and it correlates well with the occurrence of widespread graptolite, conodont, and trilobite species. The base of the Tremadocian allows scientists to relate these species not only to each other and this makes it easier to place many more species in time relative to the beginning of the Ordovician Period. A number of terms have been used to subdivide the Ordovician Period. In 2008, the ICS erected an international system of subdivisions. There exist Baltoscandic, British, Siberian, North American, Australian, the Ordovician Period in Britain was traditionally broken into Early, Middle and Late epochs. The corresponding rocks of the Ordovician System are referred to as coming from the Lower, Middle, the Floian corresponds to the lower Arenig, the Arenig continues until the early Darriwilian, subsuming the Dapingian. The Llanvirn occupies the rest of the Darriwilian, and terminates with it at the base of the Late Ordovician. The Sandbian represents the first half of the Caradoc, the Caradoc ends in the mid-Katian, during the Ordovician, the southern continents were collected into Gondwana. Gondwana started the period in equatorial latitudes and, as the period progressed, drifted toward the South Pole, the small continent Avalonia separated from Gondwana and began to move north towards Baltica and Laurentia, opening the Rheic Ocean between Gondwana and Avalonia
4.
Silurian
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The Silurian is a geologic period and system spanning 24.6 million years from the end of the Ordovician Period, at 443.8 million years ago, to the beginning of the Devonian Period,419.2 Mya. As with other periods, the rock beds that define the periods start and end are well identified. The base of the Silurian is set at a major Ordovician-Silurian extinction event when 60% of marine species were wiped out, a significant evolutionary milestone during the Silurian was the diversification of jawed and bony fish. However, terrestrial life would not greatly diversify and affect the landscape until the Devonian, the Silurian system was first identified by British geologist Sir Roderick Impey Murchison, who was examining fossil-bearing sedimentary rock strata in south Wales in the early 1830s. He named the sequences for a Celtic tribe of Wales, the Silures, inspired by his friend Adam Sedgwick and this naming does not indicate any correlation between the occurrence of the Silurian rocks and the land inhabited by the Silures. As it was first identified, the Silurian series when traced farther afield quickly came to overlap Sedgwicks Cambrian sequence, however, charles Lapworth resolved the conflict by defining a new Ordovician system including the contested beds. An early alternative name for the Silurian was Gotlandian after the strata of the Baltic island of Gotland, the French geologist Joachim Barrande, building on Murchisons work, used the term Silurian in a more comprehensive sense than was justified by subsequent knowledge. He divided the Silurian rocks of Bohemia into eight stages and his interpretation was questioned in 1854 by Edward Forbes, and the later stages of Barrande, F, G and H, have since been shown to be Devonian. Despite these modifications in the groupings of the strata, it is recognized that Barrande established Bohemia as a classic ground for the study of the earliest fossils. The epoch is named for the town of Llandovery in Carmarthenshire, the Wenlock, which lasted from 433.4 ±1.5 to 427.4 ±2.8 mya, is subdivided into the Sheinwoodian and Homerian ages. It is named after Wenlock Edge in Shropshire, England, during the Wenlock, the oldest known tracheophytes of the genus Cooksonia, appear. The first terrestrial animals also appear in the Wenlock, represented by air-breathing millipedes from Scotland. The Ludlow, lasting from 427.4 ±1.5 to 423 ±2.8 mya, comprises the Gorstian stage, lasting until 425.6 million years ago, and it is named for the town of Ludlow in Shropshire, England. The Pridoli, lasting from 423 ±1.5 to 419.2 ±2.8 mya, is the final and it is named after one locality at the Homolka a Přídolí nature reserve near the Prague suburb Slivenec in the Czech Republic. Přídolí is the old name of a field area. The high sea levels of the Silurian and the flat land resulted in a number of island chains. The southern continents remained united during this period, the melting of icecaps and glaciers contributed to a rise in sea level, recognizable from the fact that Silurian sediments overlie eroded Ordovician sediments, forming an unconformity. The continents of Avalonia, Baltica, and Laurentia drifted together near the equator and this event is the Caledonian orogeny, a spate of mountain building that stretched from New York State through conjoined Europe and Greenland to Norway
5.
Devonian
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The Devonian is a geologic period and system of the Paleozoic, spanning 60 million years from the end of the Silurian,419.2 million years ago, to the beginning of the Carboniferous,358.9 Mya. It is named after Devon, England, where rocks from this period were first studied, the first significant adaptive radiation of life on dry land occurred during the Devonian. Free-sporing vascular plants began to spread across dry land, forming extensive forests which covered the continents, by the middle of the Devonian, several groups of plants had evolved leaves and true roots, and by the end of the period the first seed-bearing plants appeared. Various terrestrial arthropods also became well-established, Fish reached substantial diversity during this time, leading the Devonian to often be dubbed the Age of Fish. The first ray-finned and lobe-finned bony fish appeared, while the placodermi began dominating almost every aquatic environment. The ancestors of all four-limbed vertebrates began adapting to walking on land, as their strong pectoral, in the oceans, primitive sharks became more numerous than in the Silurian and Late Ordovician. The first ammonites, species of molluscs, appeared, trilobites, the mollusk-like brachiopods and the great coral reefs, were still common. The Late Devonian extinction which started about 375 million years ago severely affected marine life, killing off all placodermi, and all trilobites, save for a few species of the order Proetida. The palaeogeography was dominated by the supercontinent of Gondwana to the south, the continent of Siberia to the north, while the rock beds that define the start and end of the Devonian period are well identified, the exact dates are uncertain. According to the International Commission on Stratigraphy, the Devonian extends from the end of the Silurian 419.2 Mya, another common term is Age of the Fishes, referring to the evolution of several major groups of fish that took place during the period. Older literature on the Anglo-Welsh basin divides it into the Downtonian, Dittonian, Breconian and Farlovian stages, in the Late Devonian, by contrast, arid conditions were less prevalent across the world and temperate climates were more common. The Devonian Period is formally broken into Early, Middle and Late subdivisions, the rocks corresponding to those epochs are referred to as belonging to the Lower, Middle and Upper parts of the Devonian System. Early Devonian The Early Devonian lasted from 419.2 ±2.8 to 393.3 ±2.5 and began with the Lochkovian stage, which lasted until the Pragian. It spanned from 410.8 ±2.8 to 407.6 ±2.5, and was followed by the Emsian, which lasted until the Middle Devonian began,393. 3±2.7 million years ago. Middle Devonian The Middle Devonian comprised two subdivisions, first the Eifelian, which gave way to the Givetian 387. 7±2.7 million years ago. Late Devonian Finally, the Late Devonian started with the Frasnian,382.7 ±2.8 to 372.2 ±2.5, during which the first forests took shape on land. The first tetrapods appeared in the record in the ensuing Famennian subdivision. This lasted until the end of the Devonian,358. 9±2.5 million years ago, the Devonian was a relatively warm period, and probably lacked any glaciers
6.
Carboniferous
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The Carboniferous is a geologic period and system that spans 60 million years from the end of the Devonian Period 358.9 million years ago, to the beginning of the Permian Period,298.9 Mya. The name Carboniferous means coal-bearing and derives from the Latin words carbō and ferō, and was coined by geologists William Conybeare and William Phillips in 1822. Based on a study of the British rock succession, it was the first of the system names to be employed. The Carboniferous is often treated in North America as two periods, the earlier Mississippian and the later Pennsylvanian. Terrestrial life was established by the Carboniferous period. Amphibians were the dominant land vertebrates, of one branch would eventually evolve into amniotes. Arthropods were also common, and many were much larger than those of today. Vast swaths of forest covered the land, which would eventually be laid down, the atmospheric content of oxygen also reached their highest levels in geological history during the period, 35% compared with 21% today, allowing terrestrial invertebrates to evolve to great size. A major marine and terrestrial extinction event, the Carboniferous rainforest collapse, occurred in the middle of the period, the later half of the period experienced glaciations, low sea level, and mountain building as the continents collided to form Pangaea. In the United States the Carboniferous is usually broken into Mississippian and Pennsylvanian subperiods, the Silesian is roughly contemporaneous with the late Mississippian Serpukhovian plus the Pennsylvanian. In Britain the Dinantian is traditionally known as the Carboniferous Limestone, the Namurian as the Millstone Grit, and the Westphalian as the Coal Measures and Pennant Sandstone. There was also a drop in south polar temperatures, southern Gondwanaland was glaciated throughout the period and these conditions apparently had little effect in the deep tropics, where lush swamps, later to become coal, flourished to within 30 degrees of the northernmost glaciers. Mid-Carboniferous, a drop in sea level precipitated a major extinction, one that hit crinoids. This sea level drop and the unconformity in North America separate the Mississippian subperiod from the Pennsylvanian subperiod. This happened about 323 million years ago, at the onset of the Permo-Carboniferous Glaciation, the Carboniferous was a time of active mountain-building, as the supercontinent Pangaea came together. The southern continents remained tied together in the supercontinent Gondwana, which collided with North America–Europe along the present line of eastern North America, in the same time frame, much of present eastern Eurasian plate welded itself to Europe along the line of the Ural mountains. Most of the Mesozoic supercontinent of Pangea was now assembled, although North China, the Late Carboniferous Pangaea was shaped like an O. There were two major oceans in the Carboniferous—Panthalassa and Paleo-Tethys, which was inside the O in the Carboniferous Pangaea, other minor oceans were shrinking and eventually closed - Rheic Ocean, the small, shallow Ural Ocean and Proto-Tethys Ocean
7.
Permian
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The Permian is a geologic period and system which spans 46.7 million years from the end of the Carboniferous Period 298.9 million years ago, to the beginning of the Triassic Period 252.2 Mya. It is the last period of the Paleozoic Era, the following Triassic Period belongs to the Mesozoic Era, the concept of the Permian was introduced in 1841 by geologist Sir Roderick Murchison, who named it after the city of Perm. The Permian witnessed the diversification of the early amniotes into the groups of the mammals, turtles, lepidosaurs. The world at the time was dominated by two known as Pangaea and Siberia, surrounded by a global ocean called Panthalassa. The Carboniferous rainforest collapse left behind vast regions of desert within the continental interior, amniotes, who could better cope with these drier conditions, rose to dominance in place of their amphibian ancestors. The Permian ended with the Permian–Triassic extinction event, the largest mass extinction in Earths history, in which nearly 90% of marine species and it would take well into the Triassic for life to recover from this catastrophe. Recovery from the Permian-Triassic extinction event was protracted, on land, the term Permian was introduced into geology in 1841 by Sir R. I. Murchison, president of the Geological Society of London, who identified typical strata in extensive Russian explorations undertaken with Edouard de Verneuil, the region now lies in the Perm Krai of Russia. This could have in part caused the extinctions of marine species at the end of the period by severely reducing shallow coastal areas preferred by many marine organisms. During the Permian, all the Earths major landmasses were collected into a supercontinent known as Pangaea. The Cimmeria continent rifted away from Gondwana and drifted north to Laurasia, a new ocean was growing on its southern end, the Tethys Ocean, an ocean that would dominate much of the Mesozoic Era. Large continental landmass interiors experience climates with extreme variations of heat and cold, deserts seem to have been widespread on Pangaea. Such dry conditions favored gymnosperms, plants with seeds enclosed in a cover, over plants such as ferns that disperse spores in a wetter environment. The first modern trees appeared in the Permian, the climate in the Permian was quite varied. At the start of the Permian, the Earth was still in an Ice Age, glaciers receded around the mid-Permian period as the climate gradually warmed, drying the continents interiors. In the late Permian period, the drying continued although the temperature cycled between warm and cool cycles, Permian marine deposits are rich in fossil mollusks, echinoderms, and brachiopods. By the close of the Permian, trilobites and a host of other groups became extinct. Terrestrial life in the Permian included diverse plants, fungi, arthropods, the period saw a massive desert covering the interior of Pangaea
8.
Triassic
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The Triassic is a geologic period and system which spans 50.9 million years from the end of the Permian Period 252.17 million years ago, to the beginning of the Jurassic Period 201.3 Mya. The Triassic is the first period of the Mesozoic Era, both the start and end of the period are marked by major extinction events. Therapsids and archosaurs were the terrestrial vertebrates during this time. A specialized subgroup of archosaurs, called dinosaurs, first appeared in the Late Triassic, the vast supercontinent of Pangaea existed until the mid-Triassic, after which it began to gradually rift into two separate landmasses, Laurasia to the north and Gondwana to the south. The global climate during the Triassic was mostly hot and dry, however, the climate shifted and became more humid as Pangaea began to drift apart. The end of the period was marked by yet another mass extinction, the Triassic-Jurassic extinction event. The Triassic is usually separated into Early, Middle, and Late Triassic Epochs, from the east, along the equator, the Tethys sea penetrated Pangaea, causing the Paleo-Tethys Ocean to be closed. Later in the mid-Triassic a similar sea penetrated along the equator from the west, the remaining shores were surrounded by the world-ocean known as Panthalassa. All the deep-ocean sediments laid down during the Triassic have disappeared through subduction of oceanic plates, thus, the supercontinent Pangaea was rifting during the Triassic—especially late in that period—but had not yet separated. In North America, for example, marine deposits are limited to a few exposures in the west, thus Triassic stratigraphy is mostly based on organisms that lived in lagoons and hypersaline environments, such as Estheria crustaceans. At the beginning of the Mesozoic Era, Africa was joined with Earths other continents in Pangaea, Africa shared the supercontinents relatively uniform fauna which was dominated by theropods, prosauropods and primitive ornithischians by the close of the Triassic period. Late Triassic fossils are found throughout Africa, but are common in the south than north. The time boundary separating the Permian and Triassic marks the advent of an event with global impact. At Paleorrota geopark, located in Rio Grande do Sul, Brazil, in these formations, one of the earliest dinosaurs, Staurikosaurus, as well as the mammal ancestors Brasilitherium and Brasilodon have been discovered. The Triassic continental interior climate was hot and dry, so that typical deposits are red bed sandstones and evaporites. Pangaeas large size limited the effect of the global ocean, its continental climate was highly seasonal, with very hot summers. The strong contrast between the Pangea supercontinent and the global ocean triggered intense cross-equatorial monsoons, the best studied of such episodes of humid climate, and probably the most intense and widespread, was the Carnian Pluvial Event. On land, the vascular plants included the lycophytes, the dominant cycadophytes, ginkgophyta, ferns, horsetails
9.
Jurassic
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The Jurassic is a geologic period and system that spans 56.3 million years from the end of the Triassic Period 201.3 million years ago to the beginning of the Cretaceous Period 145 Mya. The Jurassic constitutes the middle period of the Mesozoic Era, also known as the Age of Reptiles, the start of the period is marked by the major Triassic–Jurassic extinction event. The Jurassic is named after the Jura Mountains within the European Alps, by the beginning of the Jurassic, the supercontinent Pangaea had begun rifting into two landmasses, Laurasia to the north and Gondwana to the south. This created more coastlines and shifted the continental climate from dry to humid, on land, the fauna transitioned from the Triassic fauna, dominated by both dinosauromorph and crocodylomorph archosaurs, to one dominated by dinosaurs alone. The first birds also appeared during the Jurassic, having evolved from a branch of theropod dinosaurs, other major events include the appearance of the earliest lizards, and the evolution of therian mammals, including primitive placentals. Crocodilians made the transition from a terrestrial to a mode of life. The oceans were inhabited by marine reptiles such as ichthyosaurs and plesiosaurs, the chronostratigraphic term Jurassic is directly linked to the Jura Mountains. The name Jura is derived from the Celtic root jor, which was Latinised into juria, the Jurassic period is divided into the Early Jurassic, Middle, and Late Jurassic epochs. The Jurassic System, in stratigraphy, is divided into the Lower Jurassic, Middle, the separation of the term Jurassic into three sections goes back to Leopold von Buch. The Jurassic North Atlantic Ocean was relatively narrow, while the South Atlantic did not open until the following Cretaceous period, the Tethys Sea closed, and the Neotethys basin appeared. Climates were warm, with no evidence of glaciation, as in the Triassic, there was apparently no land over either pole, and no extensive ice caps existed. In contrast, the North American Jurassic record is the poorest of the Mesozoic, the Jurassic was a time of calcite sea geochemistry in which low-magnesium calcite was the primary inorganic marine precipitate of calcium carbonate. Carbonate hardgrounds were thus very common, along with calcitic ooids, calcitic cements, the first of several massive batholiths were emplaced in the northern American cordillera beginning in the mid-Jurassic, marking the Nevadan orogeny. Important Jurassic exposures are found in Russia, India, South America, Japan, Australasia. As the Jurassic proceeded, larger and more groups of dinosaurs like sauropods and ornithopods proliferated in Africa. Middle Jurassic strata are well represented nor well studied in Africa. Late Jurassic strata are also poorly represented apart from the spectacular Tendaguru fauna in Tanzania, the Late Jurassic life of Tendaguru is very similar to that found in western North Americas Morrison Formation. During the Jurassic period, the primary living in the sea were fish
10.
Cretaceous
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The Cretaceous is a geologic period and system that spans 79 million years from the end of the Jurassic Period 145 million years ago to the beginning of the Paleogene Period 66 Mya. It is the last period of the Mesozoic Era, the Cretaceous Period is usually abbreviated K, for its German translation Kreide. The Cretaceous was a period with a warm climate, resulting in high eustatic sea levels that created numerous shallow inland seas. These oceans and seas were populated with now-extinct marine reptiles, ammonites and rudists, during this time, new groups of mammals and birds, as well as flowering plants, appeared. The Cretaceous ended with a mass extinction, the Cretaceous–Paleogene extinction event, in which many groups, including non-avian dinosaurs, pterosaurs. The end of the Cretaceous is defined by the abrupt Cretaceous–Paleogene boundary, the name Cretaceous was derived from Latin creta, meaning chalk. The Cretaceous is divided into Early and Late Cretaceous epochs, or Lower and Upper Cretaceous series, in older literature the Cretaceous is sometimes divided into three series, Neocomian, Gallic and Senonian. A subdivision in eleven stages, all originating from European stratigraphy, is now used worldwide, in many parts of the world, alternative local subdivisions are still in use. As with other geologic periods, the rock beds of the Cretaceous are well identified. No great extinction or burst of diversity separates the Cretaceous from the Jurassic and this layer has been dated at 66.043 Ma. A140 Ma age for the Jurassic-Cretaceous boundary instead of the usually accepted 145 Ma was proposed in 2014 based on a study of Vaca Muerta Formation in Neuquén Basin. Víctor Ramos, one of the authors of the study proposing the 140 Ma boundary age sees the study as a first step toward formally changing the age in the International Union of Geological Sciences, due to the high sea level there was extensive space for such sedimentation. Because of the young age and great thickness of the system. Chalk is a type characteristic for the Cretaceous. It consists of coccoliths, microscopically small calcite skeletons of coccolithophores, the group is found in England, northern France, the low countries, northern Germany, Denmark and in the subsurface of the southern part of the North Sea. Chalk is not easily consolidated and the Chalk Group still consists of sediments in many places. The group also has other limestones and arenites, among the fossils it contains are sea urchins, belemnites, ammonites and sea reptiles such as Mosasaurus. In southern Europe, the Cretaceous is usually a marine system consisting of competent limestone beds or incompetent marls
11.
Paleogene
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The Paleogene is a geologic period and system that spans 43 million years from the end of the Cretaceous Period 66 million years ago to the beginning of the Neogene Period 23.03 Mya. It is the beginning of the Cenozoic Era of the present Phanerozoic Eon and this period consists of the Paleocene, Eocene and Oligocene epochs. The terms Paleogene System and lower Tertiary System are applied to the rocks deposited during the Paleogene Period. By dividing the Tertiary Period into two periods instead of directly into five epochs, the periods are more comparable to the duration of periods of the preceding Mesozoic and Paleozoic Eras. The trend was caused by the formation of the Antarctic Circumpolar Current. During the Paleogene, the continued to drift closer to their current positions. India was in the process of colliding with Asia, subsequently forming the Himalayas, the Atlantic Ocean continued to widen by a few centimeters each year. Africa was moving north to meet with Europe and form the Mediterranean, inland seas retreated from North America early in the period. Australia had also separated from Antarctica and was drifting towards Southeast Asia, mammals began a rapid diversification during this period. Some of these mammals would evolve into forms that would dominate the land, while others would become capable of living in marine, specialized terrestrial. Those that took to the oceans became modern cetaceans, while those that took to the trees became primates, the group to which humans belong. Birds, which were well established by the end of the Cretaceous. In comparison to birds and mammals, most other branches of life remained unchanged during this period. As the Earth cooled, tropical plants became less numerous and were now restricted to equatorial regions, deciduous plants, which could survive through the seasonal climates the world was now experiencing, became more common. The Paleogene is notable in the context of offshore oil drilling, and especially in Gulf of Mexico oil exploration and these rock formations represent the current cutting edge of deep-water oil discovery. Lower Tertiary explorations to date include, Kaskida Oil Field Tiber Oil Field Jack 2 Paleogene Microfossils, 180+ images of Foraminifera
12.
Neogene
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The Neogene is a geologic period and system that spans 20.45 million years from the end of the Paleogene Period 23.03 million years ago to the beginning of the present Quaternary Period 2.58 Mya. The Neogene is sub-divided into two epochs, the earlier Miocene and the later Pliocene, some geologists assert that the Neogene cannot be clearly delineated from the modern geological period, the Quaternary. During this period, mammals and birds continued to evolve into modern forms. Early hominids, the ancestors of humans, appeared in Africa near the end of the period, some continental movement took place, the most significant event being the connection of North and South America at the Isthmus of Panama, late in the Pliocene. This cut off the ocean currents from the Pacific to the Atlantic ocean. The global climate cooled considerably over the course of the Neogene, the terms Neogene System and upper Tertiary System describe the rocks deposited during the Neogene Period. The continents in the Neogene were very close to their current positions, the Isthmus of Panama formed, connecting North and South America. The Indian subcontinent continued to collide with Asia, forming the Himalayas, sea levels fell, creating land bridges between Africa and Eurasia and between Eurasia and North America. The global climate became seasonal and continued an overall drying and cooling trend which began at the start of the Paleogene. The ice caps on both poles began to grow and thicken, and by the end of the period the first of a series of glaciations of the current Ice Age began, marine and continental flora and fauna have a modern appearance. The reptile group Choristodera became extinct in the part of the period. Mammals and birds continued to be the dominant terrestrial vertebrates, the first hominids, the ancestors of humans, appeared in Africa and spread into Eurasia. In response to the cooler, seasonal climate, tropical plant species gave way to deciduous ones, grasses therefore greatly diversified, and herbivorous mammals evolved alongside it, creating the many grazing animals of today such as horses, antelope, and bison. The Neogene traditionally ended at the end of the Pliocene Epoch, just before the definition of the beginning of the Quaternary Period. However, there was a movement amongst geologists to also include ongoing geological time in the Neogene, by dividing the Cenozoic Era into three periods instead of seven epochs, the periods are more closely comparable to the duration of periods in the Mesozoic and Paleozoic eras. The International Commission on Stratigraphy once proposed that the Quaternary be considered a sub-era of the Neogene, with a date of 2.58 Ma. In the 2004 proposal of the ICS, the Neogene would have consisted of the Miocene and Pliocene epochs, thus the Neogene Period ends bounding the succeeding Quaternary Period at 2.58 Mya. Digital Atlas of Neogene Life for the Southeastern United States — by San Jose State University via Web Archive
13.
Phyllosoma
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The phyllosoma is the larval stage of spiny, slipper and coral lobsters, and represents one of the most significant characteristics that unify them into the taxon Achelata. Its body is thin, flat, and transparent, with long legs. The phyllosoma stage may last several years, during this time the larva may grow quite large, the phyllosoma larva of spiny lobsters has a long planktonic life before metamorphosing into the puerulus stage, which is the transitional stage from planktonic to a benthic existence. Despite the importance of survival to predict recruitment, not much is known about the biology of phyllosoma larvae. In many cases even natural diet is still unclear, recent efforts to identify phyllosoma larvae using molecular techniques have provided further insight into the ecology and distribution
14.
Crustacean larva
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Crustaceans may pass through a number of larval and immature stages between hatching from their eggs and reaching their adult form. Each of the stages is separated by a moult, in which the hard exoskeleton is shed to allow the animal to grow. The larvae of crustaceans often bear little resemblance to the adult and this is especially true of crustaceans which live as benthic adults, more so than where the larvae are planktonic and therefore more easily caught. Many crustacean larvae were not immediately recognised as larvae when they were discovered, the names of these genera have become generalised to cover specific larval stages across wide groups of crustaceans, such as zoea and nauplius. Other terms described forms which are found in particular groups, such as the glaucothoe of hermit crabs, or the phyllosoma of slipper lobsters. At its most complete, a life cycle begins with an egg, which is usually fertilised. This egg hatches into a pre-larva or pre-zoea, through a series of moults, the young animal then passes through various zoea stages, followed by a megalopa or post-larva. This is followed by metamorphosis into a form, which broadly resembles the adult, and after further moults. Some crustaceans continue to moult as adults, while for others, antonie van Leeuwenhoek was the first person to observe the difference between larval crustaceans and the adults when he watched the eggs of Cyclops hatching in 1699. This controversy persisted until the 1840s, and the first descriptions of a series of larval forms were not published until the 1870s. The genus name Nauplius was published posthumously by Otto Friedrich Müller in 1785 for animals now known to be the larvae of copepods, the nauplius stage is characterised by the use of the appendages of the head for swimming. The nauplius is also the stage at which a simple, unpaired eye first appears. The eye is known for that reason as the naupliar eye, the genus Zoea was initially described by Louis Augustin Guillaume Bosc in 1802 for an animal now known to be the larva of a crab. The zoea stage is characterised by the use of the thoracic appendages for swimming, the post-larva is characterised by the use of abdominal appendages for propulsion. The post-larva is usually similar to the form, and so many names have been erected for the stage in different groups. William Elford Leach erected the genus Megalopa in 1813 for a post-larval crab, every other crustacean group with free larvae shows a metamorphosis, and this difference in the larvae is thought to reflect a fundamental cleavage of the crustaceans. The larvae of remipedes are lecithotrophic, consuming egg yolk rather than using external food sources and this characteristic, which is shared with malacostracan groups such as the Decapoda and Euphausiacea has been used to suggest a link between Remipedia and Malacostraca. Young isopod crustaceans hatch directly into a stage, which is similar in appearance to the adult
15.
Taxonomy (biology)
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Taxonomy is the science of defining groups of biological organisms on the basis of shared characteristics and giving names to those groups. The exact definition of taxonomy varies from source to source, but the core of the remains, the conception, naming. There is some disagreement as to whether biological nomenclature is considered a part of taxonomy, the broadest meaning of taxonomy is used here. The word taxonomy was introduced in 1813 by Candolle, in his Théorie élémentaire de la botanique, the term alpha taxonomy is primarily used today to refer to the discipline of finding, describing, and naming taxa, particularly species. In earlier literature, the term had a different meaning, referring to morphological taxonomy, ideals can, it may be said, never be completely realized. They have, however, a value of acting as permanent stimulants. Some of us please ourselves by thinking we are now groping in a beta taxonomy, turrill thus explicitly excludes from alpha taxonomy various areas of study that he includes within taxonomy as a whole, such as ecology, physiology, genetics, and cytology. He further excludes phylogenetic reconstruction from alpha taxonomy, thus, Ernst Mayr in 1968 defined beta taxonomy as the classification of ranks higher than species. This activity is what the term denotes, it is also referred to as beta taxonomy. How species should be defined in a group of organisms gives rise to practical and theoretical problems that are referred to as the species problem. The scientific work of deciding how to define species has been called microtaxonomy, by extension, macrotaxonomy is the study of groups at higher taxonomic ranks, from subgenus and above only, than species. While some descriptions of taxonomic history attempt to date taxonomy to ancient civilizations, earlier works were primarily descriptive, and focused on plants that were useful in agriculture or medicine. There are a number of stages in scientific thinking. Early taxonomy was based on criteria, the so-called artificial systems. Later came systems based on a complete consideration of the characteristics of taxa, referred to as natural systems, such as those of de Jussieu, de Candolle and Bentham. The publication of Charles Darwins Origin of Species led to new ways of thinking about classification based on evolutionary relationships and this was the concept of phyletic systems, from 1883 onwards. This approach was typified by those of Eichler and Engler, the advent of molecular genetics and statistical methodology allowed the creation of the modern era of phylogenetic systems based on cladistics, rather than morphology alone. Taxonomy has been called the worlds oldest profession, and naming and classifying our surroundings has likely been taking place as long as mankind has been able to communicate
16.
Animal
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Animals are multicellular, eukaryotic organisms of the kingdom Animalia. The animal kingdom emerged as a clade within Apoikozoa as the group to the choanoflagellates. Animals are motile, meaning they can move spontaneously and independently at some point in their lives and their body plan eventually becomes fixed as they develop, although some undergo a process of metamorphosis later in their lives. All animals are heterotrophs, they must ingest other organisms or their products for sustenance, most known animal phyla appeared in the fossil record as marine species during the Cambrian explosion, about 542 million years ago. Animals can be divided broadly into vertebrates and invertebrates, vertebrates have a backbone or spine, and amount to less than five percent of all described animal species. They include fish, amphibians, reptiles, birds and mammals, the remaining animals are the invertebrates, which lack a backbone. These include molluscs, arthropods, annelids, nematodes, flatworms, cnidarians, ctenophores, the study of animals is called zoology. The word animal comes from the Latin animalis, meaning having breath, the biological definition of the word refers to all members of the kingdom Animalia, encompassing creatures as diverse as sponges, jellyfish, insects, and humans. Aristotle divided the world between animals and plants, and this was followed by Carl Linnaeus, in the first hierarchical classification. In Linnaeuss original scheme, the animals were one of three kingdoms, divided into the classes of Vermes, Insecta, Pisces, Amphibia, Aves, and Mammalia. Since then the last four have all been subsumed into a single phylum, in 1874, Ernst Haeckel divided the animal kingdom into two subkingdoms, Metazoa and Protozoa. The protozoa were later moved to the kingdom Protista, leaving only the metazoa, thus Metazoa is now considered a synonym of Animalia. Animals have several characteristics that set apart from other living things. Animals are eukaryotic and multicellular, which separates them from bacteria and they are heterotrophic, generally digesting food in an internal chamber, which separates them from plants and algae. They are also distinguished from plants, algae, and fungi by lacking cell walls. All animals are motile, if only at life stages. In most animals, embryos pass through a stage, which is a characteristic exclusive to animals. With a few exceptions, most notably the sponges and Placozoa and these include muscles, which are able to contract and control locomotion, and nerve tissues, which send and process signals
17.
Arthropod
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An arthropod is an invertebrate animal having an exoskeleton, a segmented body, and jointed appendages. Arthropods form the phylum Arthropoda, which includes the insects, arachnids, myriapods, arthropods are characterized by their jointed limbs and cuticle made of chitin, often mineralised with calcium carbonate. The arthropod body plan consists of segments, each with a pair of appendages, the rigid cuticle inhibits growth, so arthropods replace it periodically by moulting. Their versatility has enabled them to become the most species-rich members of all guilds in most environments. They have over a million described species, making up more than 80% of all described living species, some of which. Arthropods range in size from the microscopic crustacean Stygotantulus up to the Japanese spider crab, arthropods primary internal cavity is a hemocoel, which accommodates their internal organs, and through which their haemolymph – analogue of blood – circulates, they have open circulatory systems. Like their exteriors, the organs of arthropods are generally built of repeated segments. Their nervous system is ladder-like, with paired ventral nerve cords running through all segments and their heads are formed by fusion of varying numbers of segments, and their brains are formed by fusion of the ganglia of these segments and encircle the esophagus. The respiratory and excretory systems of arthropods vary, depending as much on their environment as on the subphylum to which they belong, arthropods also have a wide range of chemical and mechanical sensors, mostly based on modifications of the many setae that project through their cuticles. Aquatic species use internal or external fertilization. Almost all arthropods lay eggs, but scorpions give birth to live young after the eggs have hatched inside the mother, arthropod hatchlings vary from miniature adults to grubs and caterpillars that lack jointed limbs and eventually undergo a total metamorphosis to produce the adult form. The level of care for hatchlings varies from nonexistent to the prolonged care provided by scorpions. The evolutionary ancestry of arthropods dates back to the Cambrian period, the group is generally regarded as monophyletic, and many analyses support the placement of arthropods with cycloneuralians in a superphylum Ecdysozoa. Overall however, the relationships of Metazoa are not yet well resolved. Likewise, the relationships between various groups are still actively debated. Arthropods contribute to the food supply both directly as food, and more importantly indirectly as pollinators of crops. Some species are known to spread disease to humans, livestock. The word arthropod comes from the Greek ἄρθρον árthron, joint, and πούς pous, i. e. foot or leg, arthropods are invertebrates with segmented bodies and jointed limbs
18.
Crustacean
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Crustaceans form a large, diverse arthropod taxon which includes such familiar animals as crabs, lobsters, crayfish, shrimp, krill, woodlice and barnacles. In other words, some crustaceans are more related to insects. The 67,000 described species range in size from Stygotantulus stocki at 0.1 mm, to the Japanese spider crab with a leg span of up to 3.8 m, like other arthropods, crustaceans have an exoskeleton, which they moult to grow. Most crustaceans are free-living aquatic animals, but some are terrestrial, some are parasitic, the group has an extensive fossil record, reaching back to the Cambrian, and includes living fossils such as Triops cancriformis, which has existed apparently unchanged since the Triassic period. More than 10 million tons of crustaceans are produced by fishery or farming for human consumption, krill and copepods are not as widely fished, but may be the animals with the greatest biomass on the planet, and form a vital part of the food chain. The scientific study of crustaceans is known as carcinology, and a scientist who works in carcinology is a carcinologist, the body of a crustacean is composed of segments, which are grouped into three regions, the cephalon or head, the thorax, and the pleon or abdomen. The head and thorax may be fused together to form a cephalothorax, the crustacean body is protected by the hard exoskeleton, which must be moulted for the animal to grow. The shell around each somite can be divided into a dorsal tergum, ventral sternum, various parts of the exoskeleton may be fused together. The abdomen bears pleopods, and ends in a telson, which bears the anus, the number and variety of appendages in different crustaceans may be partly responsible for the groups success. It is unclear whether the condition is a derived state which evolved in crustaceans. Trilobites, for instance, also possessed biramous appendages, the main body cavity is an open circulatory system, where blood is pumped into the haemocoel by a heart located near the dorsum. Malacostraca have haemocyanin as the pigment, while copepods, ostracods, barnacles. The alimentary canal consists of a tube that often has a gizzard-like gastric mill for grinding food and a pair of digestive glands that absorb food. Structures that function as kidneys are located near the antennae, a brain exists in the form of ganglia close to the antennae, and a collection of major ganglia is found below the gut. In many decapods, the first pair of pleopods are specialised in the male for sperm transfer, many terrestrial crustaceans mate seasonally and return to the sea to release the eggs. Others, such as woodlice, lay their eggs on land, in most decapods, the females retain the eggs until they hatch into free-swimming larvae. Marine crustaceans are as ubiquitous in the oceans as insects are on land, some branchiurans are able to withstand rapid changes of salinity and will also switch hosts from marine to non-marine species. Krill are the layer and the most important part of the food chain in Antarctic animal communities
19.
Malacostraca
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Malacostraca is the largest of the six classes of crustaceans, containing about 40,000 living species, divided among 16 orders. They are abundant in all environments and have colonised freshwater. They are segmented animals, united by a body plan comprising 20 body segments, and divided into a head, thorax. The name Malacostraca was coined by a French zoologist Pierre André Latreille in 1802 and he was curator of the arthropod collection at the National Museum of Natural History in Paris. The name comes from the Greek roots μαλακός and ὄστρακον, the name is misleading, since the shell is only soft immediately after moulting, and is usually hard. Malacostracans are sometimes contrasted with entomostracans, a name applied to all crustaceans outside the Malacostraca, the class Malacostraca includes about 40,000 species, and arguably. Contains a greater diversity of forms than any other class in the animal kingdom. Malacostracans have abdominal appendages, a fact that differentiates them from all other major crustacean taxa except Remipedia, each body segment bears a pair of jointed appendages, although these may be lost secondarily. The head bears two pairs of antennae, the first of which is often biramous and the second pair bear exopods which are flattened into antennal scales known as scaphocerites. The mouthparts consist of each of mandibles, maxillules and maxillae. Usually a pair of stalked compound eyes is present, although in some taxa the eyes are unstalked, reduced or lost, up to three thoracic segments may be fused with the head to form a cephalothorax, the associated appendages turn forward and are modified as maxillipeds. A carapace may be absent, present or secondarily lost, and may cover the head, part or all of the thorax and it is variable in form and may be fused dorsally with some of the thoracic segments or occasionally be in two parts, hinged dorsally. Typically, each of the thoracic appendages is biramous and the endopods are the developed of the branches. Each endopod consist of seven articulating segments, the coxa, basis, ischium, in decapods, the claw is formed by the articulation of the dactylus against an outgrowth of the propodus. In some taxa, the exopods are lost and the appendages are uniramous, there is a clear demarcation between the thorax and the six or seven-segmented abdomen. In most taxa, each abdominal segment except the last carries a pair of biramous pleopods used for swimming, burrowing, gas exchange, the first and second abdominal pleopods may be modified in the male to form gonopods. The appendages of the last segment are typically flattened into uropods and it is the sudden flexion of this tail fan that provides the thrust for the rapid escape response of these crustaceans and the tail fan is also used in steering. In Leptostraca, the appendages on the telson instead form caudal rami, the digestive tract is straight and the foregut consists of a short oesophagus and a two-chambered stomach, the first part of which contains a gizzard-like gastric mill for grinding food
20.
Decapoda
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The Decapoda or decapods are an order of crustaceans within the class Malacostraca, including many familiar groups, such as crayfish, crabs, lobsters, prawns, and shrimp. The order is estimated to contain nearly 15,000 species in around 2,700 genera, nearly half of these species are crabs, with the shrimp and Anomura (including hermit crabs, porcelain crabs, squat lobsters making up the bulk of the remainder. The earliest fossil decapod is the Devonian Palaeopalaemon, as the name Decapoda implies, all decapods have what are considered ten legs, in the form of five pairs of thoracic appendages on the last five thoracic segments. Decapods can have as many as twenty apendages, arranged in one pair per body segment, the front three pairs function as mouthparts and are generally referred to as maxillipeds, the remainder are pereiopods. In many decapods, however, one pair of legs has enlarged pincers, the remaining appendages are found on the abdomen, with each segment capable of carrying a pair of biramous pleopods, the last of which form part of the tail fan and are called uropods. Classification within the order Decapoda depends on the structure of the gills and legs, the Dendrobranchiata consist of prawns, including many species colloquially referred to as shrimp, such as the white shrimp, Litopenaeus setiferus. The Pleocyemata include the groups, including true shrimp. Those groups which usually walk rather than form a clade called Reptantia. This classification to the level of superfamilies follows De Grave et al
21.
Pleocyemata
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Pleocyemata is a sub-order of decapod crustaceans, erected by Martin Burkenroad in 1963. Burkenroads classification replaced the earlier sub-orders of Natantia and Reptantia with the monophyletic groups Dendrobranchiata and Pleocyemata, Pleocyemata contains all the members of the Reptantia, as well as the Stenopodidea, and Caridea, which contains the true shrimp. It is this characteristic that gives the group its name, Pleocyemata also possess a lamellar gill structure as opposed to the branched found in Dendrobranchiata. The earliest fossil representative is the Devonian Palaeopalaemon, data related to Pleocyemata at Wikispecies
22.
Spiny lobster
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Spiny lobsters, also known as langouste or rock lobsters, are a family of about 60 species of achelate crustaceans, in the Decapoda Reptantia. The furry lobsters were previously separated into a family of their own, the Synaxidae, the slipper lobsters are their next-closest relatives, and these two or three families make up the Achelata. Genera of spiny lobsters include Palinurus and a number of anagrams thereof, Panulirus, Linuparus, true lobsters have much smaller antennae and claws on the first three pairs of legs, with the first being particularly enlarged. Spiny lobsters have typically a slightly compressed carapace, lacking any lateral ridges and their antennae lack a scaphocerite, the flattened exopod of the antenna. This is fused to the epistome, the flagellum, at the top of the antenna, is stout, tapering, and very long. The ambulatory legs end in claws, the fossil record of spiny lobsters has been extended by the discovery in 1995 of a 110-million-year-old fossil near El Espiñal in Chiapas, Mexico. Spiny lobsters tend to live in crevices of rocks and coral reefs, only venturing out at night to seek snails, clams, sea-hares, crabs. Sometimes, they migrate in large groups in long files of lobsters across the sea floor. These lines may be more than 50 lobsters long, spiny lobsters navigate using the smell and taste of natural substances in the water that change in different parts of the ocean. It was recently discovered that spiny lobsters can also navigate by detecting the Earths magnetic field and they keep together by contact, using their long antennae. Potential predators may be deterred from eating spiny lobsters by a loud screech made by the antennae of the spiny lobsters rubbing against a part of the exoskeleton. Spiny lobsters usually exhibit the social habit of being together, however recent studies indicate that healthy lobsters move away from infected ones, leaving the diseased lobsters to fend for themselves. Like true lobsters, spiny lobsters are edible and are a significant food source, they are the biggest food export of the Bahamas. Many spiny lobsters produce rasping sounds to repel predators by rubbing the plectrum at the base of the spiny lobsters antennae against a file, the noise is produced by frictional vibrations - sticking and slipping, similar to rubber materials sliding against hard surfaces. While a number of insects use frictional vibration mechanisms to generate sound, the stridulating organ is present in all but three genera in the family, and its form can distinguish different species. An audio recording of the rasp of a spiny lobster Media related to Palinuridae at Wikimedia Commons Data related to Palinuridae at Wikispecies
23.
Slipper lobster
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Slipper lobsters are a family of about 90 species of achelate crustaceans, in the Decapoda Reptantia, found in all warm oceans and seas. They are not true lobsters, but are closely related to spiny lobsters. Slipper lobsters are instantly recognisable by their antennae, which project forward from the head as wide plates. All the species are edible, and some, such as the Moreton Bay bug, slipper lobsters have six segments in their heads and eight segments in the thorax, which are collectively covered in a thick carapace. The six segments of the abdomen each bear a pair of pleopods, the head segments bear various mouthparts and two pairs of antennae. The first antennae, or antennules, are held on a flexible stalk. The second antennae are the slipper lobsters most conspicuous feature, as they are expanded and flattened into large plates that extend forward from the animals head. There is considerable variation in size among species of slipper lobsters, the Mediterranean species Scyllarus pygmaeus is the smallest, growing to a maximum total length of 55 millimetres, and rarely more than 40 mm. The largest species, Scyllarides haanii, may reach 50 centimetres long, slipper lobsters are typically bottom dwellers of the continental shelves, found at depths of up to 500 metres. Slipper lobsters eat a variety of molluscs, including limpets, mussels and oysters, as well as crustaceans, polychaetes and they grow slowly and live to a considerable age. The most significant predators of slipper lobsters are bony fish, with the grey triggerfish being the most significant predator of Scyllarides latus in the Mediterranean Sea, after hatching out of their eggs, young slipper lobsters pass through around ten instars as phyllosoma larvae — leaf-like, planktonic zoeae. These ten or so stages last the part of a year. Almost nothing is known about the transition from stage to the adults. Although they are fished for wherever they are found, slipper lobsters have not been the subject of intense fishery as spiny lobsters or true lobsters. The methods used for catching slipper lobsters varies depending on the species ecology and those that prefer soft substrates, such as Thenus and Ibacus, are often caught by trawling, while those that prefer crevices, caves and reefs are usually caught by SCUBA divers. The global catch of slipper lobsters was reported in 1991 to be 2,100 tonnes, more recently, annual production has been around 5,000 tonnes, the majority of which is production of Thenus orientalis in Asia. A number of names have been applied to the family Scyllaridae. The most common of these is slipper lobster, followed by shovel-nosed lobster, spanish lobster is used for members of the genus Arctides, mitten lobster for Parribacus, and fan lobster for Evibacus and Ibacus
24.
Cancrinos
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Cancrinos is a genus of fossil crustacean closely allied with the slipper lobsters. One species is known, C. claviger from the Jurassic of southern Germany, fossils of Cancrinos are rare, and their state of preservation is often imperfect. Count Georg zu Münster first described Cancrinos in 1839, based on material from the Upper Jurassic Solnhofen limestones of southern Germany. He described two species, Cancrinos claviger and C. latipes, differentiated by the size of the second antennae, but the two are now considered to be synonyms. Cancrinos differs most markedly from other related animals by the form of the second antennae, unlike living slipper lobsters, however, the flattened, distal parts of the antennae retain the ancestral state of comprising many segments, rather than being reduced to a single element. Because immature specimens have been found, parts of the ontogeny of Cancrinos are known, although it is whether the smallest specimens are in the puerulus stage. Younger specimens have less flattened antennae, more like those of living spiny lobsters, thus and this ontogeny is thought to reflect the phylogeny of Cancrinos, representing a partial development from the ancestral spiny lobster-like form towards the derived slipper lobster-like form. Media related to Cancrinos at Wikimedia Commons
25.
Chela (organ)
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A chela /kˈiːlə/, also named claw, nipper, or pincer, is a pincer-like organ terminating certain limbs of some arthropods. The name comes from Greek through New Latin, legs bearing a chela are called chelipeds. Another name is claw because most chelae are curved and have a point like a claw
26.
Greek language
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Greek is an independent branch of the Indo-European family of languages, native to Greece and other parts of the Eastern Mediterranean. It has the longest documented history of any living language, spanning 34 centuries of written records and its writing system has been the Greek alphabet for the major part of its history, other systems, such as Linear B and the Cypriot syllabary, were used previously. The alphabet arose from the Phoenician script and was in turn the basis of the Latin, Cyrillic, Armenian, Coptic, Gothic and many other writing systems. Together with the Latin texts and traditions of the Roman world, during antiquity, Greek was a widely spoken lingua franca in the Mediterranean world and many places beyond. It would eventually become the official parlance of the Byzantine Empire, the language is spoken by at least 13.2 million people today in Greece, Cyprus, Italy, Albania, Turkey, and the Greek diaspora. Greek roots are used to coin new words for other languages, Greek. Greek has been spoken in the Balkan peninsula since around the 3rd millennium BC, the earliest written evidence is a Linear B clay tablet found in Messenia that dates to between 1450 and 1350 BC, making Greek the worlds oldest recorded living language. Among the Indo-European languages, its date of earliest written attestation is matched only by the now extinct Anatolian languages, the Greek language is conventionally divided into the following periods, Proto-Greek, the unrecorded but assumed last ancestor of all known varieties of Greek. The unity of Proto-Greek would have ended as Hellenic migrants entered the Greek peninsula sometime in the Neolithic era or the Bronze Age, Mycenaean Greek, the language of the Mycenaean civilisation. It is recorded in the Linear B script on tablets dating from the 15th century BC onwards, Ancient Greek, in its various dialects, the language of the Archaic and Classical periods of the ancient Greek civilisation. It was widely known throughout the Roman Empire, after the Roman conquest of Greece, an unofficial bilingualism of Greek and Latin was established in the city of Rome and Koine Greek became a first or second language in the Roman Empire. The origin of Christianity can also be traced through Koine Greek, Medieval Greek, also known as Byzantine Greek, the continuation of Koine Greek in Byzantine Greece, up to the demise of the Byzantine Empire in the 15th century. Much of the written Greek that was used as the language of the Byzantine Empire was an eclectic middle-ground variety based on the tradition of written Koine. Modern Greek, Stemming from Medieval Greek, Modern Greek usages can be traced in the Byzantine period and it is the language used by the modern Greeks, and, apart from Standard Modern Greek, there are several dialects of it. In the modern era, the Greek language entered a state of diglossia, the historical unity and continuing identity between the various stages of the Greek language is often emphasised. Greek speakers today still tend to regard literary works of ancient Greek as part of their own rather than a foreign language and it is also often stated that the historical changes have been relatively slight compared with some other languages. According to one estimation, Homeric Greek is probably closer to demotic than 12-century Middle English is to modern spoken English, Greek is spoken by about 13 million people, mainly in Greece, Albania and Cyprus, but also worldwide by the large Greek diaspora. Greek is the language of Greece, where it is spoken by almost the entire population
27.
Antenna (biology)
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Antennae, sometimes referred to as feelers, are paired appendages used for sensing in arthropods. Antennae are connected to the first one or two segments of the arthropod head and they vary widely in form, but are always made of one or more jointed segments. While they are sensory organs, the exact nature of what they sense. Functions may variously include sensing touch, air motion, heat, vibration, antennae are sometimes modified for other purposes, such as mating, brooding, swimming, and even anchoring the arthropod to a substrate. Larval arthropods have antennae that differ from those of the adult, many crustaceans, for example, have free-swimming larval forms that use their antennae for swimming. Except the chelicerates and proturans, which have none, all non-crustacean arthropods have a pair of antennae. Crustaceans bear two pairs of antennae, the pair attached to the first segment of the head are called primary antennae or antennules. This pair is generally uniramous, but is biramous in crabs and lobsters, the pair attached to the second segment are called secondary antennae or simply antennae. The second antennae are biramous, but many species later evolved uniramous pairs. The second antennae may be reduced, apparently absent, or modified to such an extent that they no longer resemble antennae. The subdivisions of crustacean antennae have many names, including flagellomeres, annuli, articles, the terminal ends of crustacean antennae have two major categorizations, segmented and flagellate. An antenna is considered segmented if each of the annuli is separate from those around it and has individual muscle attachments. Flagellate antennae, on the hand, have muscle attachments only around the base. There are several notable uses of antennae in crustaceans. Many crustaceans have a larval stage called a nauplius, which is characterized by its use of antennae for swimming. Barnacles, a highly modified crustacean, use their antennae to attach to rocks, insects evolved from prehistoric crustaceans, and they have secondary antennae like crustaceans, but not primary antennae. Antennae are the primary olfactory sensors of insects and are accordingly well-equipped with a variety of sensilla. Paired, mobile, and segmented, they are located between the eyes on the forehead, embryologically, they represent the appendages of the second head segment
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Palinurus elephas
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Palinurus elephas is a spiny lobster, which is commonly caught in the Mediterranean Sea. Its common names include European spiny lobster, crayfish or cray, common spiny lobster, Mediterranean lobster and red lobster. Palinurus elephas is a species of spiny lobster, found in the eastern Atlantic Ocean, from southern Norway to Morocco and the Azores. It lives on exposed coasts below the intertidal zone, mainly at depths of 20 to 70 metres. P. elephas may reach up to 60 cm long, although longer than 40 cm. Few achieve their weight of several kilograms. The adults are reddish-brown with yellow spots, the carapace is slightly compressed and lacks lateral ridges. It is covered with forward pointing spines, with the supraorbital spines prominent, the antennae are very heavy and spiny. Their flagellum is tapering and is longer than the body. The first walking leg is provided with subchela, the fourth segment of this leg has a characteristic row of spines. The breeding season is in September and October, with the female brooding the reddish eggs and these eggs hatch about six months later in the spring as flattened, leaf-shaped, planktonic larvae. It is nocturnal and feeds on worms, crabs or dead animals. There are also small fisheries for this species on the west coast of Scotland, Palinurus elephas, SeaLifeBase Media related to Palinurus elephas at Wikimedia Commons
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Scyllarides latus
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Scyllarides latus, the Mediterranean slipper lobster, is a species of slipper lobster found in the Mediterranean Sea and in the eastern Atlantic Ocean. It is edible and highly regarded as food, but is now rare over much of its due to overfishing. Adults may grow to 1 foot long, are camouflaged, and have no claws and they are nocturnal, emerging from caves and other shelters during the night to feed on molluscs. As well as being eaten by humans, S. latus is preyed upon by a variety of bony fish. Its closest relative is S. herklotsii, which occurs off the Atlantic coast of West Africa, other species of Scyllarides occur in the western Atlantic Ocean, the larvae and young animals are largely unknown. In Senegal, it together with a related species Scyllarides herklotsii. S. latus can grow to a body length about 45 centimetres. This is equivalent to a length of up to 12 cm. An individual may weigh as much as 1.5 kilograms, as in all slipper lobsters, the second pair of antennae are enlarged and flattened into shovels or flippers. Despite the name lobster, slipper lobsters such as Scyllarides latus have no claws, instead, the exoskeleton, and particularly the carapace, are thicker than in clawed lobsters and spiny lobsters, acting as resilient armour. Adults are cryptically coloured, and the carapace is covered in conspicuous, S. latus lives on rocky or sandy substrates at depths of 4–100 metres. They shelter during the day in dens, on the ceilings of caves, or in reefs. The diet of S. latus consists generally of molluscs, the preferred prey is, according to different sources, either limpets or bivalves. The prey, which S. latus can detect even under 3.5 cm of sediment, is opened by careful use of the strong pointed pereiopods and they will also eat oysters and squid, but not sea urchins or muricid snails. They eat more in warmer seasons, getting through 3.2 oysters per day in July, an Octopus vulgaris has been observed to eat S. latus in an artificial setting, but it is unclear whether S. latus is preyed on by octopuses in nature. Male Scyllarus latus carry spermatophores at the base of the last two pairs of pereiopods in April, fertilisation has not been observed in this species, but most reptant decapods mate with the ventral surfaces together. Between July and August, females carry around 100,000 eggs on their enlarged, the eggs develop from being a bright orange colour to a dark brown before being shed into the water after around 16 days of development. There is normally one generation per year
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Fossil
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Fossils are the preserved remains or traces of animals, plants, and other organisms from the remote past. The totality of fossils, both discovered and undiscovered, and their placement in fossiliferous rock formations and sedimentary layers is known as the fossil record. The study of fossils across geological time, how they were formed, such a preserved specimen is called a fossil if it is older than some minimum age, most often the arbitrary date of 10,000 years. The observation that fossils were associated with certain rock strata led early geologists to recognize a geological timescale in the 19th century. The development of dating techniques in the early 20th century allowed geologists to determine the numerical or absolute age of the various strata. Like extant organisms, fossils vary in size from microscopic, even single bacterial cells one micrometer in diameter, to gigantic, such as dinosaurs, Fossils may also consist of the marks left behind by the organism while it was alive, such as animal tracks or feces. These types of fossil are called trace fossils, as opposed to body fossils, finally, past life leaves some markers that cannot be seen but can be detected in the form of biochemical signals, these are known as chemofossils or biosignatures. The process of fossilization varies according to type and external conditions. Permineralization is a process of fossilization that occurs when an organism is buried, the empty spaces within an organism become filled with mineral-rich groundwater. Minerals precipitate from the groundwater, occupying the empty spaces and this process can occur in very small spaces, such as within the cell wall of a plant cell. Small scale permineralization can produce very detailed fossils, for permineralization to occur, the organism must become covered by sediment soon after death or soon after the initial decay process. The degree to which the remains are decayed when covered determines the later details of the fossil, some fossils consist only of skeletal remains or teeth, other fossils contain traces of skin, feathers or even soft tissues. This is a form of diagenesis, in some cases the original remains of the organism completely dissolve or are otherwise destroyed. The remaining organism-shaped hole in the rock is called an external mold, if this hole is later filled with other minerals, it is a cast. An endocast or internal mold is formed when sediments or minerals fill the cavity of an organism. This is a form of cast and mold formation. If the chemistry is right, the organism can act as a nucleus for the precipitation of minerals such as siderite, if this happens rapidly before significant decay to the organic tissue, very fine three-dimensional morphological detail can be preserved. Nodules from the Carboniferous Mazon Creek fossil beds of Illinois, USA, are among the best documented examples of such mineralization, replacement occurs when the shell, bone or other tissue is replaced with another mineral
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Portable Document Format
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The Portable Document Format is a file format used to present documents in a manner independent of application software, hardware, and operating systems. Each PDF file encapsulates a complete description of a fixed-layout flat document, including the text, fonts, graphics, PDF was developed in the early 1990s as a way to share computer documents, including text formatting and inline images. It was among a number of competing formats such as DjVu, Envoy, Common Ground Digital Paper, Farallon Replica, in those early years before the rise of the World Wide Web and HTML documents, PDF was popular mainly in desktop publishing workflows. Adobe Systems made the PDF specification available free of charge in 1993 and these proprietary technologies are not standardized and their specification is published only on Adobe’s website. Many of them are not supported by popular third-party implementations of PDF. So when organizations publish PDFs which use proprietary technologies, they present accessibility issues for some users. In 2014, ISO TC171 voted to deprecate XFA for ISO 32000-2, on January 9,2017, the final draft for ISO 32000-2 was published, thus reaching the approval stage. The PDF combines three technologies, A subset of the PostScript page description programming language, for generating the layout, a font-embedding/replacement system to allow fonts to travel with the documents. A structured storage system to bundle these elements and any associated content into a single file, PostScript is a page description language run in an interpreter to generate an image, a process requiring many resources. It can handle graphics and standard features of programming such as if. PDF is largely based on PostScript but simplified to remove flow control features like these, often, the PostScript-like PDF code is generated from a source PostScript file. The graphics commands that are output by the PostScript code are collected and tokenized, any files, graphics, or fonts to which the document refers also are collected. Then, everything is compressed to a single file, therefore, the entire PostScript world remains intact. PDF supports graphic transparency, PostScript does not, PostScript is an interpreted programming language with an implicit global state, so instructions accompanying the description of one page can affect the appearance of any following page. Therefore, all preceding pages in a PostScript document must be processed to determine the appearance of a given page. A PDF file is a 7-bit ASCII file, except for elements that may have binary content. A PDF file starts with a header containing the magic number, the format is a subset of a COS format. A COS tree file consists primarily of objects, of which there are eight types, Boolean values, representing true or false Numbers Strings, enclosed within parentheses, objects may be either direct or indirect
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PubMed Identifier
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PubMed is a free search engine accessing primarily the MEDLINE database of references and abstracts on life sciences and biomedical topics. The United States National Library of Medicine at the National Institutes of Health maintains the database as part of the Entrez system of information retrieval, from 1971 to 1997, MEDLINE online access to the MEDLARS Online computerized database primarily had been through institutional facilities, such as university libraries. PubMed, first released in January 1996, ushered in the era of private, free, home-, the PubMed system was offered free to the public in June 1997, when MEDLINE searches via the Web were demonstrated, in a ceremony, by Vice President Al Gore. Information about the journals indexed in MEDLINE, and available through PubMed, is found in the NLM Catalog. As of 5 January 2017, PubMed has more than 26.8 million records going back to 1966, selectively to the year 1865, and very selectively to 1809, about 500,000 new records are added each year. As of the date,13.1 million of PubMeds records are listed with their abstracts. In 2016, NLM changed the system so that publishers will be able to directly correct typos. Simple searches on PubMed can be carried out by entering key aspects of a subject into PubMeds search window, when a journal article is indexed, numerous article parameters are extracted and stored as structured information. Such parameters are, Article Type, Secondary identifiers, Language, publication type parameter enables many special features. As these clinical girish can generate small sets of robust studies with considerable precision, since July 2005, the MEDLINE article indexing process extracts important identifiers from the article abstract and puts those in a field called Secondary Identifier. The secondary identifier field is to store numbers to various databases of molecular sequence data, gene expression or chemical compounds. For clinical trials, PubMed extracts trial IDs for the two largest trial registries, ClinicalTrials. gov and the International Standard Randomized Controlled Trial Number Register, a reference which is judged particularly relevant can be marked and related articles can be identified. If relevant, several studies can be selected and related articles to all of them can be generated using the Find related data option, the related articles are then listed in order of relatedness. To create these lists of related articles, PubMed compares words from the title and abstract of each citation, as well as the MeSH headings assigned, using a powerful word-weighted algorithm. The related articles function has been judged to be so precise that some researchers suggest it can be used instead of a full search, a strong feature of PubMed is its ability to automatically link to MeSH terms and subheadings. Examples would be, bad breath links to halitosis, heart attack to myocardial infarction, where appropriate, these MeSH terms are automatically expanded, that is, include more specific terms. Terms like nursing are automatically linked to Nursing or Nursing and this important feature makes PubMed searches automatically more sensitive and avoids false-negative hits by compensating for the diversity of medical terminology. The My NCBI area can be accessed from any computer with web-access, an earlier version of My NCBI was called PubMed Cubby
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Oxford University Press
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Oxford University Press is the largest university press in the world, and the second oldest after Cambridge University Press. It is a department of the University of Oxford and is governed by a group of 15 academics appointed by the known as the delegates of the press. They are headed by the secretary to the delegates, who serves as OUPs chief executive, Oxford University has used a similar system to oversee OUP since the 17th century. The university became involved in the print trade around 1480, and grew into a printer of Bibles, prayer books. OUP took on the project became the Oxford English Dictionary in the late 19th century. Moves into international markets led to OUP opening its own offices outside the United Kingdom, by contracting out its printing and binding operations, the modern OUP publishes some 6,000 new titles around the world each year. OUP was first exempted from United States corporation tax in 1972, as a department of a charity, OUP is exempt from income tax and corporate tax in most countries, but may pay sales and other commercial taxes on its products. The OUP today transfers 30% of its surplus to the rest of the university. OUP is the largest university press in the world by the number of publications, publishing more than 6,000 new books every year, the Oxford University Press Museum is located on Great Clarendon Street, Oxford. Visits must be booked in advance and are led by a member of the archive staff, displays include a 19th-century printing press, the OUP buildings, and the printing and history of the Oxford Almanack, Alice in Wonderland and the Oxford English Dictionary. The first printer associated with Oxford University was Theoderic Rood, the first book printed in Oxford, in 1478, an edition of Rufinuss Expositio in symbolum apostolorum, was printed by another, anonymous, printer. Famously, this was mis-dated in Roman numerals as 1468, thus apparently pre-dating Caxton, roods printing included John Ankywylls Compendium totius grammaticae, which set new standards for teaching of Latin grammar. After Rood, printing connected with the university remained sporadic for over half a century, the chancellor, Robert Dudley, 1st Earl of Leicester, pleaded Oxfords case. Some royal assent was obtained, since the printer Joseph Barnes began work, Oxfords chancellor, Archbishop William Laud, consolidated the legal status of the universitys printing in the 1630s. Laud envisaged a unified press of world repute, Oxford would establish it on university property, govern its operations, employ its staff, determine its printed work, and benefit from its proceeds. To that end, he petitioned Charles I for rights that would enable Oxford to compete with the Stationers Company and the Kings Printer and these were brought together in Oxfords Great Charter in 1636, which gave the university the right to print all manner of books. Laud also obtained the privilege from the Crown of printing the King James or Authorized Version of Scripture at Oxford and this privilege created substantial returns in the next 250 years, although initially it was held in abeyance. The Stationers Company was deeply alarmed by the threat to its trade, under this, the Stationers paid an annual rent for the university not to exercise its full printing rights – money Oxford used to purchase new printing equipment for smaller purposes
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International Standard Book Number
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The International Standard Book Number is a unique numeric commercial book identifier. An ISBN is assigned to each edition and variation of a book, for example, an e-book, a paperback and a hardcover edition of the same book would each have a different ISBN. The ISBN is 13 digits long if assigned on or after 1 January 2007, the method of assigning an ISBN is nation-based and varies from country to country, often depending on how large the publishing industry is within a country. The initial ISBN configuration of recognition was generated in 1967 based upon the 9-digit Standard Book Numbering created in 1966, the 10-digit ISBN format was developed by the International Organization for Standardization and was published in 1970 as international standard ISO2108. Occasionally, a book may appear without a printed ISBN if it is printed privately or the author does not follow the usual ISBN procedure, however, this can be rectified later. Another identifier, the International Standard Serial Number, identifies periodical publications such as magazines, the ISBN configuration of recognition was generated in 1967 in the United Kingdom by David Whitaker and in 1968 in the US by Emery Koltay. The 10-digit ISBN format was developed by the International Organization for Standardization and was published in 1970 as international standard ISO2108, the United Kingdom continued to use the 9-digit SBN code until 1974. The ISO on-line facility only refers back to 1978, an SBN may be converted to an ISBN by prefixing the digit 0. For example, the edition of Mr. J. G. Reeder Returns, published by Hodder in 1965, has SBN340013818 -340 indicating the publisher,01381 their serial number. This can be converted to ISBN 0-340-01381-8, the check digit does not need to be re-calculated, since 1 January 2007, ISBNs have contained 13 digits, a format that is compatible with Bookland European Article Number EAN-13s. An ISBN is assigned to each edition and variation of a book, for example, an ebook, a paperback, and a hardcover edition of the same book would each have a different ISBN. The ISBN is 13 digits long if assigned on or after 1 January 2007, a 13-digit ISBN can be separated into its parts, and when this is done it is customary to separate the parts with hyphens or spaces. Separating the parts of a 10-digit ISBN is also done with either hyphens or spaces, figuring out how to correctly separate a given ISBN number is complicated, because most of the parts do not use a fixed number of digits. ISBN issuance is country-specific, in that ISBNs are issued by the ISBN registration agency that is responsible for country or territory regardless of the publication language. Some ISBN registration agencies are based in national libraries or within ministries of culture, in other cases, the ISBN registration service is provided by organisations such as bibliographic data providers that are not government funded. In Canada, ISBNs are issued at no cost with the purpose of encouraging Canadian culture. In the United Kingdom, United States, and some countries, where the service is provided by non-government-funded organisations. Australia, ISBNs are issued by the library services agency Thorpe-Bowker
35.
Wikidata
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Wikidata is a collaboratively edited knowledge base operated by the Wikimedia Foundation. It is intended to provide a source of data which can be used by Wikimedia projects such as Wikipedia. This is similar to the way Wikimedia Commons provides storage for files and access to those files for all Wikimedia projects. Wikidata is powered by the software Wikibase, Wikidata is a document-oriented database, focused on items. Each item represents a topic and is identified by a number, prefixed with the letter Q—for example. This enables the basic information required to identify the topic the item covers to be translated without favouring any language, information is added to items by creating statements. Statements take the form of pairs, with each statement consisting of a property. The creation of the project was funded by donations from the Allen Institute for Artificial Intelligence, the Gordon and Betty Moore Foundation, at this time, only the first phase was available. Historically, a Wikipedia article would include a list of links, being links to articles on the same topic in other editions of Wikipedia. Initially, Wikidata was a repository of interlanguage links. No Wikipedia language editions were able to access Wikidata, so they needed to continue to maintain their own lists of interlanguage links, on 14 January 2013, the Hungarian Wikipedia became the first to enable the provision of interlanguage links via Wikidata. This functionality was extended to the Hebrew and Italian Wikipedias on 30 January, to the English Wikipedia on 13 February, on 23 September 2013, phase 1 went live on Wikimedia Commons. The first aspects of the second phase were deployed on 4 February 2013, the values were initially limited to two data types, with more data types to follow later. The first new type, string, was deployed on 6 March, the ability of the various language editions of Wikipedia to access data added to Wikidata as part of phase two was rolled out progressively between 27 March and 25 April 2013. On 16 September 2015, Wikidata began allowing so-called arbitrary access, for example, in the past the article about Berlin you could not access data about Germany, but with arbitrary access it could. On 27 April 2016 arbitrary access was activated on Wikimedia Commons, phase 3 will involve database querying and the creation of lists based on data stored on Wikidata. As of October 2016 two tools for querying Wikidata were available, AutoList and PetScan, additionally to a public SPARQL endpoint, there is concern that the project is being influenced by lobbying companies, PR professionals and search engine optimizers. As of December 2015, according to Wikimedia statistics, half of the information in Wikidata is unsourced, another 30% is labeled as having come from Wikipedia, but with no indication as to which article
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Wikispecies
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Wikispecies is a wiki-based online project supported by the Wikimedia Foundation. Its aim is to create a free content catalogue of all species. Jimmy Wales stated that editors are not required to fax in their degrees, Wikispecies is available under the GNU Free Documentation License and CC BY-SA3.0. Benedikt Mandl co-ordinated the efforts of people who are interested in getting involved with the project. Databases were evaluated and the administrators contacted, some of them have agreed on providing their data for Wikispecies, the board of directors of the Wikimedia Foundation voted by 4 to 0 in favor of the establishment of a Wikispecies. The project was launched in August 2004 and is hosted at species. wikimedia. org and it was officially merged to a sister project of Wikimedia Foundation on September 14,2004. On October 10,2006, the project exceeded 75,000 articles, on May 20,2007, the project exceeded 100,000 articles with a total of 5,495 registered users. On September 8,2008, the project exceeded 150,000 articles with a total of 9,224 registered users, on October 23,2011, the project reached 300,000 articles. On June 16,2014, the project reached 400,000 articles, on January 7,2017, the project reached 500,000 articles. Wikispecies has disabled local upload and asks users to use images from Wikimedia Commons, Wikispecies does not allow the use of content that does not conform to a free license
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National Center for Biotechnology Information
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The National Center for Biotechnology Information is part of the United States National Library of Medicine, a branch of the National Institutes of Health. The NCBI is located in Bethesda, Maryland and was founded in 1988 through legislation sponsored by Senator Claude Pepper, the NCBI houses a series of databases relevant to biotechnology and biomedicine and is an important resource for bioinformatics tools and services. Major databases include GenBank for DNA sequences and PubMed, a database for the biomedical literature. Other databases include the NCBI Epigenomics database, all these databases are available online through the Entrez search engine. NCBI is directed by David Lipman, one of the authors of the BLAST sequence alignment program. He also leads a research program, including groups led by Stephen Altschul, David Landsman, Eugene Koonin, John Wilbur, Teresa Przytycka. NCBI is listed in the Registry of Research Data Repositories re3data. org, NCBI has had responsibility for making available the GenBank DNA sequence database since 1992. GenBank coordinates with individual laboratories and other databases such as those of the European Molecular Biology Laboratory. Since 1992, NCBI has grown to other databases in addition to GenBank. The NCBI assigns a unique identifier to each species of organism, the NCBI has software tools that are available by WWW browsing or by FTP. For example, BLAST is a sequence similarity searching program, BLAST can do sequence comparisons against the GenBank DNA database in less than 15 seconds. RAG2/IL2RG The NCBI Bookshelf is a collection of freely accessible, downloadable, some of the books are online versions of previously published books, while others, such as Coffee Break, are written and edited by NCBI staff. BLAST is a used for calculating sequence similarity between biological sequences such as nucleotide sequences of DNA and amino acid sequences of proteins. BLAST is a tool for finding sequences similar to the query sequence within the same organism or in different organisms. It searches the query sequence on NCBI databases and servers and post the results back to the browser in chosen format. Input sequences to the BLAST are mostly in FASTA or Genbank format while output could be delivered in variety of such as HTML, XML formatting. HTML is the output format for NCBIs web-page. Entrez is both indexing and retrieval system having data from sources for biomedical research
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World Register of Marine Species
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The World Register of Marine Species is a database that aims to provide an authoritative and comprehensive list of names of marine organisms. The content of the registry is edited and maintained by scientific specialists on each group of organism and these taxonomists control the quality of the information, which is gathered from malacological journals and several regional and taxon-specific databases. WoRMS maintains valid names of all organisms, but also provides information on synonyms. It will be a task to maintain the registry, as new species are constantly being discovered and described by scientists. In addition, the nomenclature and taxonomy of existing species is often corrected or changed as new research is constantly being published, WoRMS was founded in 2008 and grew out of the European Register of Marine Species. WoRMS has established formal agreements with other biodiversity projects, including the Global Biodiversity Information Facility. In 2008, WoRMS stated that it hoped to have a record of all marine species completed by 2010. As of November 2016, WoRMS contained listings for 242,562 valid marine species and their goal is to have a listing for each of the approximately 240,000 marine species. Census of Marine Life Ocean Biogeographic Information System Catalog of Fishes World Register of Marine Species
Media related to Achelata at Wikimedia Commons