1.
Hominidae
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Several revisions in classifying the great apes have caused the use of the term hominid to vary over time. Its original meaning referred only to humans and their closest non-extant relatives and that restrictive meaning has now been largely assumed by the term hominin, which comprises all members of the human clade after the split from the chimpanzees. The current, 21st-century meaning of hominid includes all the great apes including humans, the most recent common ancestor of all Hominidae lived roughly 14 million years ago, when the ancestors of the orangutans speciated from the ancestral line of the other three genera. Those ancestors of the family Hominidae had already speciated from the family Hylobatidae, in the early Miocene, about 22 million years ago, there were many species of arboreally adapted primitive catarrhines from East Africa, the variety suggests a long history of prior diversification. Fossils at 20 million years ago include fragments attributed to Victoriapithecus, the most recent of these far-flung Miocene apes is Oreopithecus, from the fossil-rich coal beds in northern Italy and dated to 9 million years ago. Species close to the last common ancestor of gorillas, chimpanzees and humans may be represented by Nakalipithecus fossils found in Kenya and Ouranopithecus found in Greece. Molecular evidence suggests that between 8 and 4 million years ago, first the gorillas, and then the split off from the line leading to the humans. Human DNA is approximately 98. 4% identical to that of chimpanzees when comparing single nucleotide polymorphisms, the earliest fossils argued by some to belong to the human lineage are Sahelanthropus tchadensis and Orrorin tugenensis, followed by Ardipithecus, with species Ar. kadabba and Ar. ramidus. The classification of the apes has been revised several times in the last few decades. The original meaning of the referred to only humans and their closest relatives—what is now the modern meaning of the term hominin. And the meaning of the taxon Hominidae changed gradually, leading to a different usage of hominid that today all the great apes including humans. A hominid is a member of the family Hominidae, the apes, orangutans, gorillas, chimpanzees. A hominine is a member of the subfamily Homininae, gorillas, chimpanzees, a hominin is a member of the tribe Hominini, Chimpanzees and humans. A homininan is a member of the subtribe Hominina of the tribe Hominini, a human is a member of the genus Homo, of which Homo sapiens is the only extant species, and within that Homo sapiens sapiens is the only surviving subspecies. For each clade it is indicated approximately when newer extant clades emerged, some texts will refer to Homonini as the Hominina branch. Many scientists, including paleoanthropologists, continue to use the term hominid to mean humans, as mentioned, Hominidae was originally the name given to the family of humans and their close relatives, with the other great apes all being placed in a separate family, the Pongidae. However, that eventually made Pongidae paraphyletic because at least one great ape species proved to be more closely related to humans than to other great apes. Most taxonomists today encourage monophyletic groups—this would require, in this case, thus, many biologists now assign Pongo to the family Hominidae
2.
Precambrian
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The Precambrian is the earliest period of Earths history, set before the current Phanerozoic Eon. The Precambrian is a supereon that is subdivided into three eons of the time scale. It spans from the formation of Earth about 4.6 billion years ago to the beginning of the Cambrian Period, about 541 million years ago, the Precambrian accounts for 89% of geologic time. Relatively little is known about the Precambrian, despite it making up roughly seven-eighths of the Earths history, the Precambrian fossil record is poorer than that of the succeeding Phanerozoic, and fossils from that time are of limited biostratigraphic use. This is because many Precambrian rocks have been metamorphosed, obscuring their origins, while others have been destroyed by erosion. A stable crust was apparently in place by 4,412 Ma, the term Precambrian is recognized by the International Commission on Stratigraphy as a general term including the Archean and Proterozoic eons. It is still used by geologists and paleontologists for general discussions not requiring the more specific eon names and it was briefly called the Cryptozoic eon. A specific date for the origin of life has not been determined, carbon found in 3.8 billion year old rocks from islands off western Greenland may be of organic origin. Well-preserved microscopic fossils of bacteria older than 3.46 billion years have found in Western Australia. Probable fossils 100 million years older have been found in the same area, there is a fairly solid record of bacterial life throughout the remainder of the Precambrian. The oldest fossil evidence from that era of such complex life comes from the Lantian formation of the Ediacarian period, a very diverse collection of soft-bodied forms is found in a variety of locations worldwide and date to between 635 and 542 Ma. These are referred to as Ediacaran or Vendian biota, hard-shelled creatures appeared toward the end of that time span, marking the beginning of the Phanerozoic era. By the middle of the following Cambrian period, a diverse fauna is recorded in the Burgess Shale. The explosion in diversity of lifeforms during the early Cambrian is called the Cambrian explosion of life, while land seems to have been devoid of plants and animals, cyanobacteria and other microbes formed prokaryotic mats that covered terrestrial areas. Evidence of the details of plate motions and other activity in the Precambrian has been poorly preserved. It is generally believed that small proto-continents existed prior to 4280 Ma, the supercontinent, known as Rodinia, broke up around 750 Ma. A number of glacial periods have been identified going as far back as the Huronian epoch, one of the best studied is the Sturtian-Varangian glaciation, around 850–635 Ma, which may have brought glacial conditions all the way to the equator, resulting in a Snowball Earth. The atmosphere of the early Earth is not well understood, most geologists believe it was composed primarily of nitrogen, carbon dioxide, and other relatively inert gases, and was lacking in free oxygen
3.
Cambrian
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The Cambrian Period was the first geological period of the Paleozoic Era, of the Phanerozoic Eon. The Cambrian lasted 55.6 million years from the end of the preceding Ediacaran Period 541 million years ago to the beginning of the Ordovician Period 485.4 mya and its subdivisions, and its base, are somewhat in flux. The period was established by Adam Sedgwick, who named it after Cambria, the Latinised form of Cymru, the Welsh name for Wales, as a result, our understanding of the Cambrian biology surpasses that of some later periods. The rapid diversification of lifeforms in the Cambrian, known as the Cambrian explosion, most of the continents were probably dry and rocky due to a lack of vegetation. Shallow seas flanked the margins of several continents created during the breakup of the supercontinent Pannotia, the seas were relatively warm, and polar ice was absent for much of the period. The United States Federal Geographic Data Committee uses a barred capital C ⟨Є⟩ character similar to the capital letter Ukrainian Ye ⟨Є⟩ to represent the Cambrian Period, the proper Unicode character is U+A792 Ꞓ LATIN CAPITAL LETTER C WITH BAR. Despite the long recognition of its distinction from younger Ordovician Period rocks and older Supereon Precambrian rocks, the base of the Cambrian lies atop a complex assemblage of trace fossils known as the Treptichnus pedum assemblage. Pedum in Namibia, Spain and Newfoundland, and possibly, in the western USA, the stratigraphic range of T. pedum overlaps the range of the Ediacaran fossils in Namibia, and probably in Spain. The Cambrian Period followed the Ediacaran Period and was followed by the Ordovician Period, the Cambrian is divided into four epochs and ten ages. Currently only two series and five stages are named and have a GSSP, because the international stratigraphic subdivision is not yet complete, many local subdivisions are still widely used. In some of these subdivisions the Cambrian is divided into three epochs with locally differing names – the Early Cambrian, Middle Cambrian and Furongian, rocks of these epochs are referred to as belonging to the Lower, Middle, or Upper Cambrian. Trilobite zones allow biostratigraphic correlation in the Cambrian, each of the local epochs is divided into several stages. The International Commission on Stratigraphy list the Cambrian period as beginning at 541 million years ago, the lower boundary of the Cambrian was originally held to represent the first appearance of complex life, represented by trilobites. The recognition of small shelly fossils before the first trilobites, and Ediacara biota substantially earlier and this formal designation allowed radiometric dates to be obtained from samples across the globe that corresponded to the base of the Cambrian. Early dates of 570 million years ago quickly gained favour, though the used to obtain this number are now considered to be unsuitable. A more precise date using modern radiometric dating yield a date of 541 ±0.3 million years ago, most continental land was clustered in the Southern Hemisphere at this time, but was drifting north. Large, high-velocity rotational movement of Gondwana appears to have occurred in the Early Cambrian, the sea levels fluctuated somewhat, suggesting there were ice ages, associated with pulses of expansion and contraction of a south polar ice cap. In Baltoscandia a Lower Cambrian transgression transformed large swathes of the Sub-Cambrian peneplain into a epicontinental sea, the Earth was generally cold during the early Cambrian, probably due to the ancient continent of Gondwana covering the South Pole and cutting off polar ocean currents
4.
Ordovician
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The Ordovician is a geologic period and system, the second of six periods of the Paleozoic Era. The Ordovician spans 41.2 million years from the end of the Cambrian Period 485.4 million years ago to the start of the Silurian Period 443.8 Mya. Lapworth recognized that the fauna in the disputed strata were different from those of either the Cambrian or the Silurian periods. It received international sanction in 1960, when it was adopted as a period of the Paleozoic Era by the International Geological Congress. Life continued to flourish during the Ordovician as it did in the earlier Cambrian period, invertebrates, namely molluscs and arthropods, dominated the oceans. The Great Ordovician Biodiversification Event considerably increased the diversity of life, fish, the worlds first true vertebrates, continued to evolve, and those with jaws may have first appeared late in the period. Life had yet to diversify on land, about 100 times as many meteorites struck the Earth during the Ordovician compared with today. The Ordovician Period began with a major extinction called the Cambrian–Ordovician extinction event and it lasted for about 42 million years and ended with the Ordovician–Silurian extinction event, about 443.8 Mya which wiped out 60% of marine genera. The dates given are recent radiometric dates and vary slightly from those found in other sources and this second period of the Paleozoic era created abundant fossils that became major petroleum and gas reservoirs. The boundary chosen for the beginning of both the Ordovician Period and the Tremadocian stage is highly significant and it correlates well with the occurrence of widespread graptolite, conodont, and trilobite species. The base of the Tremadocian allows scientists to relate these species not only to each other and this makes it easier to place many more species in time relative to the beginning of the Ordovician Period. A number of terms have been used to subdivide the Ordovician Period. In 2008, the ICS erected an international system of subdivisions. There exist Baltoscandic, British, Siberian, North American, Australian, the Ordovician Period in Britain was traditionally broken into Early, Middle and Late epochs. The corresponding rocks of the Ordovician System are referred to as coming from the Lower, Middle, the Floian corresponds to the lower Arenig, the Arenig continues until the early Darriwilian, subsuming the Dapingian. The Llanvirn occupies the rest of the Darriwilian, and terminates with it at the base of the Late Ordovician. The Sandbian represents the first half of the Caradoc, the Caradoc ends in the mid-Katian, during the Ordovician, the southern continents were collected into Gondwana. Gondwana started the period in equatorial latitudes and, as the period progressed, drifted toward the South Pole, the small continent Avalonia separated from Gondwana and began to move north towards Baltica and Laurentia, opening the Rheic Ocean between Gondwana and Avalonia
5.
Silurian
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The Silurian is a geologic period and system spanning 24.6 million years from the end of the Ordovician Period, at 443.8 million years ago, to the beginning of the Devonian Period,419.2 Mya. As with other periods, the rock beds that define the periods start and end are well identified. The base of the Silurian is set at a major Ordovician-Silurian extinction event when 60% of marine species were wiped out, a significant evolutionary milestone during the Silurian was the diversification of jawed and bony fish. However, terrestrial life would not greatly diversify and affect the landscape until the Devonian, the Silurian system was first identified by British geologist Sir Roderick Impey Murchison, who was examining fossil-bearing sedimentary rock strata in south Wales in the early 1830s. He named the sequences for a Celtic tribe of Wales, the Silures, inspired by his friend Adam Sedgwick and this naming does not indicate any correlation between the occurrence of the Silurian rocks and the land inhabited by the Silures. As it was first identified, the Silurian series when traced farther afield quickly came to overlap Sedgwicks Cambrian sequence, however, charles Lapworth resolved the conflict by defining a new Ordovician system including the contested beds. An early alternative name for the Silurian was Gotlandian after the strata of the Baltic island of Gotland, the French geologist Joachim Barrande, building on Murchisons work, used the term Silurian in a more comprehensive sense than was justified by subsequent knowledge. He divided the Silurian rocks of Bohemia into eight stages and his interpretation was questioned in 1854 by Edward Forbes, and the later stages of Barrande, F, G and H, have since been shown to be Devonian. Despite these modifications in the groupings of the strata, it is recognized that Barrande established Bohemia as a classic ground for the study of the earliest fossils. The epoch is named for the town of Llandovery in Carmarthenshire, the Wenlock, which lasted from 433.4 ±1.5 to 427.4 ±2.8 mya, is subdivided into the Sheinwoodian and Homerian ages. It is named after Wenlock Edge in Shropshire, England, during the Wenlock, the oldest known tracheophytes of the genus Cooksonia, appear. The first terrestrial animals also appear in the Wenlock, represented by air-breathing millipedes from Scotland. The Ludlow, lasting from 427.4 ±1.5 to 423 ±2.8 mya, comprises the Gorstian stage, lasting until 425.6 million years ago, and it is named for the town of Ludlow in Shropshire, England. The Pridoli, lasting from 423 ±1.5 to 419.2 ±2.8 mya, is the final and it is named after one locality at the Homolka a Přídolí nature reserve near the Prague suburb Slivenec in the Czech Republic. Přídolí is the old name of a field area. The high sea levels of the Silurian and the flat land resulted in a number of island chains. The southern continents remained united during this period, the melting of icecaps and glaciers contributed to a rise in sea level, recognizable from the fact that Silurian sediments overlie eroded Ordovician sediments, forming an unconformity. The continents of Avalonia, Baltica, and Laurentia drifted together near the equator and this event is the Caledonian orogeny, a spate of mountain building that stretched from New York State through conjoined Europe and Greenland to Norway
6.
Devonian
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The Devonian is a geologic period and system of the Paleozoic, spanning 60 million years from the end of the Silurian,419.2 million years ago, to the beginning of the Carboniferous,358.9 Mya. It is named after Devon, England, where rocks from this period were first studied, the first significant adaptive radiation of life on dry land occurred during the Devonian. Free-sporing vascular plants began to spread across dry land, forming extensive forests which covered the continents, by the middle of the Devonian, several groups of plants had evolved leaves and true roots, and by the end of the period the first seed-bearing plants appeared. Various terrestrial arthropods also became well-established, Fish reached substantial diversity during this time, leading the Devonian to often be dubbed the Age of Fish. The first ray-finned and lobe-finned bony fish appeared, while the placodermi began dominating almost every aquatic environment. The ancestors of all four-limbed vertebrates began adapting to walking on land, as their strong pectoral, in the oceans, primitive sharks became more numerous than in the Silurian and Late Ordovician. The first ammonites, species of molluscs, appeared, trilobites, the mollusk-like brachiopods and the great coral reefs, were still common. The Late Devonian extinction which started about 375 million years ago severely affected marine life, killing off all placodermi, and all trilobites, save for a few species of the order Proetida. The palaeogeography was dominated by the supercontinent of Gondwana to the south, the continent of Siberia to the north, while the rock beds that define the start and end of the Devonian period are well identified, the exact dates are uncertain. According to the International Commission on Stratigraphy, the Devonian extends from the end of the Silurian 419.2 Mya, another common term is Age of the Fishes, referring to the evolution of several major groups of fish that took place during the period. Older literature on the Anglo-Welsh basin divides it into the Downtonian, Dittonian, Breconian and Farlovian stages, in the Late Devonian, by contrast, arid conditions were less prevalent across the world and temperate climates were more common. The Devonian Period is formally broken into Early, Middle and Late subdivisions, the rocks corresponding to those epochs are referred to as belonging to the Lower, Middle and Upper parts of the Devonian System. Early Devonian The Early Devonian lasted from 419.2 ±2.8 to 393.3 ±2.5 and began with the Lochkovian stage, which lasted until the Pragian. It spanned from 410.8 ±2.8 to 407.6 ±2.5, and was followed by the Emsian, which lasted until the Middle Devonian began,393. 3±2.7 million years ago. Middle Devonian The Middle Devonian comprised two subdivisions, first the Eifelian, which gave way to the Givetian 387. 7±2.7 million years ago. Late Devonian Finally, the Late Devonian started with the Frasnian,382.7 ±2.8 to 372.2 ±2.5, during which the first forests took shape on land. The first tetrapods appeared in the record in the ensuing Famennian subdivision. This lasted until the end of the Devonian,358. 9±2.5 million years ago, the Devonian was a relatively warm period, and probably lacked any glaciers
7.
Carboniferous
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The Carboniferous is a geologic period and system that spans 60 million years from the end of the Devonian Period 358.9 million years ago, to the beginning of the Permian Period,298.9 Mya. The name Carboniferous means coal-bearing and derives from the Latin words carbō and ferō, and was coined by geologists William Conybeare and William Phillips in 1822. Based on a study of the British rock succession, it was the first of the system names to be employed. The Carboniferous is often treated in North America as two periods, the earlier Mississippian and the later Pennsylvanian. Terrestrial life was established by the Carboniferous period. Amphibians were the dominant land vertebrates, of one branch would eventually evolve into amniotes. Arthropods were also common, and many were much larger than those of today. Vast swaths of forest covered the land, which would eventually be laid down, the atmospheric content of oxygen also reached their highest levels in geological history during the period, 35% compared with 21% today, allowing terrestrial invertebrates to evolve to great size. A major marine and terrestrial extinction event, the Carboniferous rainforest collapse, occurred in the middle of the period, the later half of the period experienced glaciations, low sea level, and mountain building as the continents collided to form Pangaea. In the United States the Carboniferous is usually broken into Mississippian and Pennsylvanian subperiods, the Silesian is roughly contemporaneous with the late Mississippian Serpukhovian plus the Pennsylvanian. In Britain the Dinantian is traditionally known as the Carboniferous Limestone, the Namurian as the Millstone Grit, and the Westphalian as the Coal Measures and Pennant Sandstone. There was also a drop in south polar temperatures, southern Gondwanaland was glaciated throughout the period and these conditions apparently had little effect in the deep tropics, where lush swamps, later to become coal, flourished to within 30 degrees of the northernmost glaciers. Mid-Carboniferous, a drop in sea level precipitated a major extinction, one that hit crinoids. This sea level drop and the unconformity in North America separate the Mississippian subperiod from the Pennsylvanian subperiod. This happened about 323 million years ago, at the onset of the Permo-Carboniferous Glaciation, the Carboniferous was a time of active mountain-building, as the supercontinent Pangaea came together. The southern continents remained tied together in the supercontinent Gondwana, which collided with North America–Europe along the present line of eastern North America, in the same time frame, much of present eastern Eurasian plate welded itself to Europe along the line of the Ural mountains. Most of the Mesozoic supercontinent of Pangea was now assembled, although North China, the Late Carboniferous Pangaea was shaped like an O. There were two major oceans in the Carboniferous—Panthalassa and Paleo-Tethys, which was inside the O in the Carboniferous Pangaea, other minor oceans were shrinking and eventually closed - Rheic Ocean, the small, shallow Ural Ocean and Proto-Tethys Ocean
8.
Permian
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The Permian is a geologic period and system which spans 46.7 million years from the end of the Carboniferous Period 298.9 million years ago, to the beginning of the Triassic Period 252.2 Mya. It is the last period of the Paleozoic Era, the following Triassic Period belongs to the Mesozoic Era, the concept of the Permian was introduced in 1841 by geologist Sir Roderick Murchison, who named it after the city of Perm. The Permian witnessed the diversification of the early amniotes into the groups of the mammals, turtles, lepidosaurs. The world at the time was dominated by two known as Pangaea and Siberia, surrounded by a global ocean called Panthalassa. The Carboniferous rainforest collapse left behind vast regions of desert within the continental interior, amniotes, who could better cope with these drier conditions, rose to dominance in place of their amphibian ancestors. The Permian ended with the Permian–Triassic extinction event, the largest mass extinction in Earths history, in which nearly 90% of marine species and it would take well into the Triassic for life to recover from this catastrophe. Recovery from the Permian-Triassic extinction event was protracted, on land, the term Permian was introduced into geology in 1841 by Sir R. I. Murchison, president of the Geological Society of London, who identified typical strata in extensive Russian explorations undertaken with Edouard de Verneuil, the region now lies in the Perm Krai of Russia. This could have in part caused the extinctions of marine species at the end of the period by severely reducing shallow coastal areas preferred by many marine organisms. During the Permian, all the Earths major landmasses were collected into a supercontinent known as Pangaea. The Cimmeria continent rifted away from Gondwana and drifted north to Laurasia, a new ocean was growing on its southern end, the Tethys Ocean, an ocean that would dominate much of the Mesozoic Era. Large continental landmass interiors experience climates with extreme variations of heat and cold, deserts seem to have been widespread on Pangaea. Such dry conditions favored gymnosperms, plants with seeds enclosed in a cover, over plants such as ferns that disperse spores in a wetter environment. The first modern trees appeared in the Permian, the climate in the Permian was quite varied. At the start of the Permian, the Earth was still in an Ice Age, glaciers receded around the mid-Permian period as the climate gradually warmed, drying the continents interiors. In the late Permian period, the drying continued although the temperature cycled between warm and cool cycles, Permian marine deposits are rich in fossil mollusks, echinoderms, and brachiopods. By the close of the Permian, trilobites and a host of other groups became extinct. Terrestrial life in the Permian included diverse plants, fungi, arthropods, the period saw a massive desert covering the interior of Pangaea
9.
Triassic
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The Triassic is a geologic period and system which spans 50.9 million years from the end of the Permian Period 252.17 million years ago, to the beginning of the Jurassic Period 201.3 Mya. The Triassic is the first period of the Mesozoic Era, both the start and end of the period are marked by major extinction events. Therapsids and archosaurs were the terrestrial vertebrates during this time. A specialized subgroup of archosaurs, called dinosaurs, first appeared in the Late Triassic, the vast supercontinent of Pangaea existed until the mid-Triassic, after which it began to gradually rift into two separate landmasses, Laurasia to the north and Gondwana to the south. The global climate during the Triassic was mostly hot and dry, however, the climate shifted and became more humid as Pangaea began to drift apart. The end of the period was marked by yet another mass extinction, the Triassic-Jurassic extinction event. The Triassic is usually separated into Early, Middle, and Late Triassic Epochs, from the east, along the equator, the Tethys sea penetrated Pangaea, causing the Paleo-Tethys Ocean to be closed. Later in the mid-Triassic a similar sea penetrated along the equator from the west, the remaining shores were surrounded by the world-ocean known as Panthalassa. All the deep-ocean sediments laid down during the Triassic have disappeared through subduction of oceanic plates, thus, the supercontinent Pangaea was rifting during the Triassic—especially late in that period—but had not yet separated. In North America, for example, marine deposits are limited to a few exposures in the west, thus Triassic stratigraphy is mostly based on organisms that lived in lagoons and hypersaline environments, such as Estheria crustaceans. At the beginning of the Mesozoic Era, Africa was joined with Earths other continents in Pangaea, Africa shared the supercontinents relatively uniform fauna which was dominated by theropods, prosauropods and primitive ornithischians by the close of the Triassic period. Late Triassic fossils are found throughout Africa, but are common in the south than north. The time boundary separating the Permian and Triassic marks the advent of an event with global impact. At Paleorrota geopark, located in Rio Grande do Sul, Brazil, in these formations, one of the earliest dinosaurs, Staurikosaurus, as well as the mammal ancestors Brasilitherium and Brasilodon have been discovered. The Triassic continental interior climate was hot and dry, so that typical deposits are red bed sandstones and evaporites. Pangaeas large size limited the effect of the global ocean, its continental climate was highly seasonal, with very hot summers. The strong contrast between the Pangea supercontinent and the global ocean triggered intense cross-equatorial monsoons, the best studied of such episodes of humid climate, and probably the most intense and widespread, was the Carnian Pluvial Event. On land, the vascular plants included the lycophytes, the dominant cycadophytes, ginkgophyta, ferns, horsetails
10.
Jurassic
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The Jurassic is a geologic period and system that spans 56.3 million years from the end of the Triassic Period 201.3 million years ago to the beginning of the Cretaceous Period 145 Mya. The Jurassic constitutes the middle period of the Mesozoic Era, also known as the Age of Reptiles, the start of the period is marked by the major Triassic–Jurassic extinction event. The Jurassic is named after the Jura Mountains within the European Alps, by the beginning of the Jurassic, the supercontinent Pangaea had begun rifting into two landmasses, Laurasia to the north and Gondwana to the south. This created more coastlines and shifted the continental climate from dry to humid, on land, the fauna transitioned from the Triassic fauna, dominated by both dinosauromorph and crocodylomorph archosaurs, to one dominated by dinosaurs alone. The first birds also appeared during the Jurassic, having evolved from a branch of theropod dinosaurs, other major events include the appearance of the earliest lizards, and the evolution of therian mammals, including primitive placentals. Crocodilians made the transition from a terrestrial to a mode of life. The oceans were inhabited by marine reptiles such as ichthyosaurs and plesiosaurs, the chronostratigraphic term Jurassic is directly linked to the Jura Mountains. The name Jura is derived from the Celtic root jor, which was Latinised into juria, the Jurassic period is divided into the Early Jurassic, Middle, and Late Jurassic epochs. The Jurassic System, in stratigraphy, is divided into the Lower Jurassic, Middle, the separation of the term Jurassic into three sections goes back to Leopold von Buch. The Jurassic North Atlantic Ocean was relatively narrow, while the South Atlantic did not open until the following Cretaceous period, the Tethys Sea closed, and the Neotethys basin appeared. Climates were warm, with no evidence of glaciation, as in the Triassic, there was apparently no land over either pole, and no extensive ice caps existed. In contrast, the North American Jurassic record is the poorest of the Mesozoic, the Jurassic was a time of calcite sea geochemistry in which low-magnesium calcite was the primary inorganic marine precipitate of calcium carbonate. Carbonate hardgrounds were thus very common, along with calcitic ooids, calcitic cements, the first of several massive batholiths were emplaced in the northern American cordillera beginning in the mid-Jurassic, marking the Nevadan orogeny. Important Jurassic exposures are found in Russia, India, South America, Japan, Australasia. As the Jurassic proceeded, larger and more groups of dinosaurs like sauropods and ornithopods proliferated in Africa. Middle Jurassic strata are well represented nor well studied in Africa. Late Jurassic strata are also poorly represented apart from the spectacular Tendaguru fauna in Tanzania, the Late Jurassic life of Tendaguru is very similar to that found in western North Americas Morrison Formation. During the Jurassic period, the primary living in the sea were fish
11.
Cretaceous
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The Cretaceous is a geologic period and system that spans 79 million years from the end of the Jurassic Period 145 million years ago to the beginning of the Paleogene Period 66 Mya. It is the last period of the Mesozoic Era, the Cretaceous Period is usually abbreviated K, for its German translation Kreide. The Cretaceous was a period with a warm climate, resulting in high eustatic sea levels that created numerous shallow inland seas. These oceans and seas were populated with now-extinct marine reptiles, ammonites and rudists, during this time, new groups of mammals and birds, as well as flowering plants, appeared. The Cretaceous ended with a mass extinction, the Cretaceous–Paleogene extinction event, in which many groups, including non-avian dinosaurs, pterosaurs. The end of the Cretaceous is defined by the abrupt Cretaceous–Paleogene boundary, the name Cretaceous was derived from Latin creta, meaning chalk. The Cretaceous is divided into Early and Late Cretaceous epochs, or Lower and Upper Cretaceous series, in older literature the Cretaceous is sometimes divided into three series, Neocomian, Gallic and Senonian. A subdivision in eleven stages, all originating from European stratigraphy, is now used worldwide, in many parts of the world, alternative local subdivisions are still in use. As with other geologic periods, the rock beds of the Cretaceous are well identified. No great extinction or burst of diversity separates the Cretaceous from the Jurassic and this layer has been dated at 66.043 Ma. A140 Ma age for the Jurassic-Cretaceous boundary instead of the usually accepted 145 Ma was proposed in 2014 based on a study of Vaca Muerta Formation in Neuquén Basin. Víctor Ramos, one of the authors of the study proposing the 140 Ma boundary age sees the study as a first step toward formally changing the age in the International Union of Geological Sciences, due to the high sea level there was extensive space for such sedimentation. Because of the young age and great thickness of the system. Chalk is a type characteristic for the Cretaceous. It consists of coccoliths, microscopically small calcite skeletons of coccolithophores, the group is found in England, northern France, the low countries, northern Germany, Denmark and in the subsurface of the southern part of the North Sea. Chalk is not easily consolidated and the Chalk Group still consists of sediments in many places. The group also has other limestones and arenites, among the fossils it contains are sea urchins, belemnites, ammonites and sea reptiles such as Mosasaurus. In southern Europe, the Cretaceous is usually a marine system consisting of competent limestone beds or incompetent marls
12.
Paleogene
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The Paleogene is a geologic period and system that spans 43 million years from the end of the Cretaceous Period 66 million years ago to the beginning of the Neogene Period 23.03 Mya. It is the beginning of the Cenozoic Era of the present Phanerozoic Eon and this period consists of the Paleocene, Eocene and Oligocene epochs. The terms Paleogene System and lower Tertiary System are applied to the rocks deposited during the Paleogene Period. By dividing the Tertiary Period into two periods instead of directly into five epochs, the periods are more comparable to the duration of periods of the preceding Mesozoic and Paleozoic Eras. The trend was caused by the formation of the Antarctic Circumpolar Current. During the Paleogene, the continued to drift closer to their current positions. India was in the process of colliding with Asia, subsequently forming the Himalayas, the Atlantic Ocean continued to widen by a few centimeters each year. Africa was moving north to meet with Europe and form the Mediterranean, inland seas retreated from North America early in the period. Australia had also separated from Antarctica and was drifting towards Southeast Asia, mammals began a rapid diversification during this period. Some of these mammals would evolve into forms that would dominate the land, while others would become capable of living in marine, specialized terrestrial. Those that took to the oceans became modern cetaceans, while those that took to the trees became primates, the group to which humans belong. Birds, which were well established by the end of the Cretaceous. In comparison to birds and mammals, most other branches of life remained unchanged during this period. As the Earth cooled, tropical plants became less numerous and were now restricted to equatorial regions, deciduous plants, which could survive through the seasonal climates the world was now experiencing, became more common. The Paleogene is notable in the context of offshore oil drilling, and especially in Gulf of Mexico oil exploration and these rock formations represent the current cutting edge of deep-water oil discovery. Lower Tertiary explorations to date include, Kaskida Oil Field Tiber Oil Field Jack 2 Paleogene Microfossils, 180+ images of Foraminifera
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Neogene
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The Neogene is a geologic period and system that spans 20.45 million years from the end of the Paleogene Period 23.03 million years ago to the beginning of the present Quaternary Period 2.58 Mya. The Neogene is sub-divided into two epochs, the earlier Miocene and the later Pliocene, some geologists assert that the Neogene cannot be clearly delineated from the modern geological period, the Quaternary. During this period, mammals and birds continued to evolve into modern forms. Early hominids, the ancestors of humans, appeared in Africa near the end of the period, some continental movement took place, the most significant event being the connection of North and South America at the Isthmus of Panama, late in the Pliocene. This cut off the ocean currents from the Pacific to the Atlantic ocean. The global climate cooled considerably over the course of the Neogene, the terms Neogene System and upper Tertiary System describe the rocks deposited during the Neogene Period. The continents in the Neogene were very close to their current positions, the Isthmus of Panama formed, connecting North and South America. The Indian subcontinent continued to collide with Asia, forming the Himalayas, sea levels fell, creating land bridges between Africa and Eurasia and between Eurasia and North America. The global climate became seasonal and continued an overall drying and cooling trend which began at the start of the Paleogene. The ice caps on both poles began to grow and thicken, and by the end of the period the first of a series of glaciations of the current Ice Age began, marine and continental flora and fauna have a modern appearance. The reptile group Choristodera became extinct in the part of the period. Mammals and birds continued to be the dominant terrestrial vertebrates, the first hominids, the ancestors of humans, appeared in Africa and spread into Eurasia. In response to the cooler, seasonal climate, tropical plant species gave way to deciduous ones, grasses therefore greatly diversified, and herbivorous mammals evolved alongside it, creating the many grazing animals of today such as horses, antelope, and bison. The Neogene traditionally ended at the end of the Pliocene Epoch, just before the definition of the beginning of the Quaternary Period. However, there was a movement amongst geologists to also include ongoing geological time in the Neogene, by dividing the Cenozoic Era into three periods instead of seven epochs, the periods are more closely comparable to the duration of periods in the Mesozoic and Paleozoic eras. The International Commission on Stratigraphy once proposed that the Quaternary be considered a sub-era of the Neogene, with a date of 2.58 Ma. In the 2004 proposal of the ICS, the Neogene would have consisted of the Miocene and Pliocene epochs, thus the Neogene Period ends bounding the succeeding Quaternary Period at 2.58 Mya. Digital Atlas of Neogene Life for the Southeastern United States — by San Jose State University via Web Archive
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Bornean orangutan
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The Bornean orangutan is a species of orangutan native to the island of Borneo. Together with the Sumatran orangutan, it belongs to the genus of great apes native to Asia. Like the other apes, orangutans are highly intelligent, displaying advanced tool use. Orangutans share approximately 97% of their DNA with humans, the Bornean orangutan is a critically endangered species, with deforestation, palm oil plantations and hunting posing a serious threat to its continued existence. The Bornean orangutan and the Sumatran orangutan diverged about 400,000 years ago, the two orangutan species were considered merely subspecies until 1996, they were elevated to species following sequencing of their mitochondrial DNA. The population currently listed as P. p. wurmbii may be closer to the Sumatran orangutan than to the Bornean orangutan, if this is confirmed, P. abelii would be a subspecies of P. wurmbii. In addition, the locality of P. pygmaeus has not been established beyond doubt. Bradon-Jones et al. considered P. morio to be a synonym of P. pygmaeus, the Bornean orangutan is the third-heaviest living primate after the two species of gorilla, and the largest truly arboreal animal alive today. Body weights broadly overlap with the considerably taller Homo sapiens, by comparison, the Sumatran orangutan is similar in size but, on average, is marginally lighter in weight. A survey of wild orangutans found that weigh on average 75 kg, ranging from 50–100 kg, and 1. 2–1.4 m long, females average 38.5 kg, ranging from 30–50 kg. While in captivity, orangutans can grow considerably overweight, up to more than 165 kg, the heaviest known male orangutan in captivity was an obese male named Andy, who weighed 204 kg in 1959 when he was 13 years old. The Bornean orangutan has a body shape with very long arms that may reach up to 1.5 metres in length. It has a coarse, shaggy, reddish coat and prehensile, grasping hands, the Bornean orangutan lives in tropical and subtropical moist broadleaf forests in the Bornean lowlands, as well as mountainous areas up to 1,500 metres above sea level. This species lives throughout the canopy of primary and secondary forests and it can be found in the two Malaysian states of Sabah and Sarawak, and three of the four Indonesian Provinces of Kalimantan. In history, orangutans ranged throughout Southeast Asia and into southern China, as well as on the island of Java and they primarily inhabit peat swamp forests, tropical health forests, and mixed dipterocarp forests. Bornean orangutan are more solitary than their Sumatran relatives, two or three orangutans with overlapping territories may interact, but only for short periods of time. Although orangutans are not territorial, adult males will display threatening behaviors upon meeting other males, males are considered the most solitary of the orangutans. The Bornean orangutan has a lifespan of 35–45 years in the wild, despite being arboreal, the Bornean orangutan travels on the ground more than its Sumatran counterpart
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Taxonomy (biology)
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Taxonomy is the science of defining groups of biological organisms on the basis of shared characteristics and giving names to those groups. The exact definition of taxonomy varies from source to source, but the core of the remains, the conception, naming. There is some disagreement as to whether biological nomenclature is considered a part of taxonomy, the broadest meaning of taxonomy is used here. The word taxonomy was introduced in 1813 by Candolle, in his Théorie élémentaire de la botanique, the term alpha taxonomy is primarily used today to refer to the discipline of finding, describing, and naming taxa, particularly species. In earlier literature, the term had a different meaning, referring to morphological taxonomy, ideals can, it may be said, never be completely realized. They have, however, a value of acting as permanent stimulants. Some of us please ourselves by thinking we are now groping in a beta taxonomy, turrill thus explicitly excludes from alpha taxonomy various areas of study that he includes within taxonomy as a whole, such as ecology, physiology, genetics, and cytology. He further excludes phylogenetic reconstruction from alpha taxonomy, thus, Ernst Mayr in 1968 defined beta taxonomy as the classification of ranks higher than species. This activity is what the term denotes, it is also referred to as beta taxonomy. How species should be defined in a group of organisms gives rise to practical and theoretical problems that are referred to as the species problem. The scientific work of deciding how to define species has been called microtaxonomy, by extension, macrotaxonomy is the study of groups at higher taxonomic ranks, from subgenus and above only, than species. While some descriptions of taxonomic history attempt to date taxonomy to ancient civilizations, earlier works were primarily descriptive, and focused on plants that were useful in agriculture or medicine. There are a number of stages in scientific thinking. Early taxonomy was based on criteria, the so-called artificial systems. Later came systems based on a complete consideration of the characteristics of taxa, referred to as natural systems, such as those of de Jussieu, de Candolle and Bentham. The publication of Charles Darwins Origin of Species led to new ways of thinking about classification based on evolutionary relationships and this was the concept of phyletic systems, from 1883 onwards. This approach was typified by those of Eichler and Engler, the advent of molecular genetics and statistical methodology allowed the creation of the modern era of phylogenetic systems based on cladistics, rather than morphology alone. Taxonomy has been called the worlds oldest profession, and naming and classifying our surroundings has likely been taking place as long as mankind has been able to communicate
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Animal
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Animals are multicellular, eukaryotic organisms of the kingdom Animalia. The animal kingdom emerged as a clade within Apoikozoa as the group to the choanoflagellates. Animals are motile, meaning they can move spontaneously and independently at some point in their lives and their body plan eventually becomes fixed as they develop, although some undergo a process of metamorphosis later in their lives. All animals are heterotrophs, they must ingest other organisms or their products for sustenance, most known animal phyla appeared in the fossil record as marine species during the Cambrian explosion, about 542 million years ago. Animals can be divided broadly into vertebrates and invertebrates, vertebrates have a backbone or spine, and amount to less than five percent of all described animal species. They include fish, amphibians, reptiles, birds and mammals, the remaining animals are the invertebrates, which lack a backbone. These include molluscs, arthropods, annelids, nematodes, flatworms, cnidarians, ctenophores, the study of animals is called zoology. The word animal comes from the Latin animalis, meaning having breath, the biological definition of the word refers to all members of the kingdom Animalia, encompassing creatures as diverse as sponges, jellyfish, insects, and humans. Aristotle divided the world between animals and plants, and this was followed by Carl Linnaeus, in the first hierarchical classification. In Linnaeuss original scheme, the animals were one of three kingdoms, divided into the classes of Vermes, Insecta, Pisces, Amphibia, Aves, and Mammalia. Since then the last four have all been subsumed into a single phylum, in 1874, Ernst Haeckel divided the animal kingdom into two subkingdoms, Metazoa and Protozoa. The protozoa were later moved to the kingdom Protista, leaving only the metazoa, thus Metazoa is now considered a synonym of Animalia. Animals have several characteristics that set apart from other living things. Animals are eukaryotic and multicellular, which separates them from bacteria and they are heterotrophic, generally digesting food in an internal chamber, which separates them from plants and algae. They are also distinguished from plants, algae, and fungi by lacking cell walls. All animals are motile, if only at life stages. In most animals, embryos pass through a stage, which is a characteristic exclusive to animals. With a few exceptions, most notably the sponges and Placozoa and these include muscles, which are able to contract and control locomotion, and nerve tissues, which send and process signals
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Chordate
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Chordates are deuterostomes, as during the embryo development stage the anus forms before the mouth. They are also bilaterally symmetric coelomates, in the case of vertebrate chordates, the notochord is usually replaced by a vertebral column during development, and they may have body plans organized via segmentation. There are also additional extinct taxa, the Vertebrata are sometimes considered as a subgroup of the clade Craniata, consisting of chordates with a skull, the Craniata and Tunicata compose the clade Olfactores. Of the more than 65,000 living species of chordates, the worlds largest and fastest animals, the blue whale and peregrine falcon respectively, are chordates, as are humans. Fossil chordates are known from at least as early as the Cambrian explosion, Hemichordata, which includes the acorn worms, has been presented as a fourth chordate subphylum, but it now is usually treated as a separate phylum. The Hemichordata, along with the Echinodermata, form the Ambulacraria, the Chordata and Ambulacraria form the superphylum Deuterostomia, composed of the deuterostomes. Attempts to work out the relationships of the chordates have produced several hypotheses. All of the earliest chordate fossils have found in the Early Cambrian Chengjiang fauna. Because the fossil record of early chordates is poor, only molecular phylogenetics offers a prospect of dating their emergence. However, the use of molecular phylogenetics for dating evolutionary transitions is controversial and it has also proved difficult to produce a detailed classification within the living chordates. Attempts to produce family trees shows that many of the traditional classes are paraphyletic. While this has been known since the 19th century, an insistence on only monophyletic taxa has resulted in vertebrate classification being in a state of flux. Although the name Chordata is attributed to William Bateson, it was already in prevalent use by 1880, ernst Haeckel described a taxon comprising tunicates, cephalochordates, and vertebrates in 1866. Though he used the German vernacular form, it is allowed under the ICZN code because of its subsequent latinization, among the vertebrate sub-group of chordates the notochord develops into the spine, and in wholly aquatic species this helps the animal to swim by flexing its tail. In fish and other vertebrates, this develops into the spinal cord, the pharynx is the part of the throat immediately behind the mouth. In fish, the slits are modified to form gills, a muscular tail that extends backwards behind the anus. This is a groove in the wall of the pharynx. In filter-feeding species it produces mucus to gather food particles, which helps in transporting food to the esophagus and it also stores iodine, and may be a precursor of the vertebrate thyroid gland
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Mammal
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Mammals are any vertebrates within the class Mammalia, a clade of endothermic amniotes distinguished from reptiles by the possession of a neocortex, hair, three middle ear bones and mammary glands. All female mammals nurse their young with milk, secreted from the mammary glands, Mammals include the largest animals on the planet, the great whales. The basic body type is a quadruped, but some mammals are adapted for life at sea, in the air, in trees. The largest group of mammals, the placentals, have a placenta, Mammals range in size from the 30–40 mm bumblebee bat to the 30-meter blue whale. With the exception of the five species of monotreme, all modern mammals give birth to live young, most mammals, including the six most species-rich orders, belong to the placental group. The largest orders are the rodents, bats and Soricomorpha, the next three biggest orders, depending on the biological classification scheme used, are the Primates, the Cetartiodactyla, and the Carnivora. Living mammals are divided into the Yinotheria and Theriiformes There are around 5450 species of mammal, in some classifications, extant mammals are divided into two subclasses, the Prototheria, that is, the order Monotremata, and the Theria, or the infraclasses Metatheria and Eutheria. The marsupials constitute the group of the Metatheria, and include all living metatherians as well as many extinct ones. Much of the changes reflect the advances of cladistic analysis and molecular genetics, findings from molecular genetics, for example, have prompted adopting new groups, such as the Afrotheria, and abandoning traditional groups, such as the Insectivora. The mammals represent the only living Synapsida, which together with the Sauropsida form the Amniota clade, the early synapsid mammalian ancestors were sphenacodont pelycosaurs, a group that produced the non-mammalian Dimetrodon. At the end of the Carboniferous period, this group diverged from the line that led to todays reptiles. Some mammals are intelligent, with some possessing large brains, self-awareness, Mammals can communicate and vocalize in several different ways, including the production of ultrasound, scent-marking, alarm signals, singing, and echolocation. Mammals can organize themselves into fission-fusion societies, harems, and hierarchies, most mammals are polygynous, but some can be monogamous or polyandrous. They provided, and continue to provide, power for transport and agriculture, as well as commodities such as meat, dairy products, wool. Mammals are hunted or raced for sport, and are used as model organisms in science, Mammals have been depicted in art since Palaeolithic times, and appear in literature, film, mythology, and religion. Defaunation of mammals is primarily driven by anthropogenic factors, such as poaching and habitat destruction, Mammal classification has been through several iterations since Carl Linnaeus initially defined the class. No classification system is accepted, McKenna & Bell and Wilson & Reader provide useful recent compendiums. Though field work gradually made Simpsons classification outdated, it remains the closest thing to a classification of mammals
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Primate
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A primate is a mammal of the order Primates. In taxonomy, primates include two distinct lineages, strepsirrhines and haplorhines, Primates arose from ancestors that lived in the trees of tropical forests, many primate characteristics represent adaptations to life in this challenging three-dimensional environment. Most primate species remain at least partly arboreal, with the exception of humans, who inhabit every continent except for Antarctica, most primates live in tropical or subtropical regions of the Americas, Africa and Asia. Based on fossil evidence, the earliest known true primates, represented by the genus Teilhardina, an early close primate relative known from abundant remains is the Late Paleocene Plesiadapis, c. Molecular clock studies suggest that the branch may be even older. The order Primates was traditionally divided into two groupings, prosimians and anthropoids. Prosimians have characteristics more like those of the earliest primates, and include the lemurs of Madagascar, lorisoids, simians include monkeys, apes and hominins. Simians are divided into two groups, catarrhine monkeys and apes of Africa and Southeast Asia and platyrrhine or New World monkeys of South, catarrhines consist of Old World monkeys, gibbons and great apes, New World monkeys include the capuchin, howler and squirrel monkeys. Humans are the only extant catarrhines to have spread successfully outside of Africa, South Asia, New primate species are still being discovered. More than 25 species were described in the decade of the 2000s. Considered generalist mammals, primates exhibit a range of characteristics. Some primates are primarily terrestrial rather than arboreal, but all species possess adaptations for climbing trees, locomotion techniques used include leaping from tree to tree, walking on two or four limbs, knuckle-walking, and swinging between branches of trees. Primates are characterized by large brains relative to other mammals, as well as a reliance on stereoscopic vision at the expense of smell. These features are developed in monkeys and apes and noticeably less so in lorises. Three-color vision has developed in some primates, most also have opposable thumbs and some have prehensile tails. Many species are dimorphic, differences include body mass, canine tooth size. Primates have slower rates of development than other similarly sized mammals and reach maturity later, depending on the species, adults may live in solitude, in mated pairs, or in groups of up to hundreds of members. The relationships among the different groups of primates were not clearly understood until relatively recently, for example, ape has been used either as an alternative for monkey or for any tailless, relatively human-like primate
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Haplorhini
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Haplorrhini is a clade containing the tarsiers and the simians. The name is also spelt Haplorrhini, the simians include catarrhines, and the platyrrhines. The extinct omomyids, which are considered to be the most basal haplorhines, are believed to be closely related to the tarsiers than to other haplorhines. Haplorhines share a number of derived features that distinguish them from the strepsirrhine wet-nosed primates, the haplorhine upper lip, which has replaced the ancestral rhinarium found in strepsirrhines, is not directly connected to their nose or gum, allowing a large range of facial expressions. Their brain-to-body mass ratio is greater than the strepsirrhines. Haplorhines have a plate, unlike the postorbital bar found in strepsirrhines. All anthropoids have a uterus, tarsiers have a bicornate uterus like the strepsirrhines. Most species typically have single births, although twins and triplets are common for marmosets, despite similar gestation periods, haplorhine newborns are relatively much larger than strepsirrhine newborns, but have a longer dependence period on their mother. This difference in size and dependence is credited to the complexity of their behavior. The taxonomic name Haplorhini derives from the Ancient Greek haploûs and rhinos and it refers to the lack of a rhinarium or wet nose, which is found in many mammals, including strepsirrhine primates. Haplorhini and its sister clade, Strepsirrhini, diverged about 65 million years ago, the fossil Archicebus may be similar to the most recent common ancestor at this time. The other major clade within Haplorhini, the simians, is divided into two parvorders, Platyrrhini and Catarrhini, the New World monkeys split from catarrhines about 40 mya, while the apes diverged from Old World monkeys about 25 mya. The available fossil evidence indicates both the hominoid and cercopithecoid clades originated in Africa. The following is the listing of the living families, and their placement in the Order Primates, ORDER PRIMATES Suborder Strepsirrhini, lemurs, lorises
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Simian
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The simians are the monkeys, cladistically including the apes, the New World monkeys or platyrrhines, and the catarrhine clade consisting of the Old World monkeys and apes. The simian line and the line diverged about 60 million years ago. Forty million years ago, simians from Africa colonized South America, the remaining simians split 25 million years ago into apes and Old World monkeys. Under modern classification, the tarsiers and simians are grouped under the suborder Haplorhini while the strepsirrhines are placed in suborder Strepsirrhini, in anthropoidea, evidences indicate that the Old and the New World primates went through parallel evolution. Primatology, paleoanthropology, and other related fields are split on their usage of the synonymous infraorder names, Simiiformes and Anthropoidea. According to Robert Hoffstetter, the term Simiiformes has priority over Anthropoidea because of the taxonomic term Simii by van der Hoeven, from which it is constructed, dates to 1833. In contrast, Anthropoidea by Mivart dates to 1864, while Simiiformes by Haeckel dates to 1866, Hoffstetter also argued that Simiiformes is also constructed like a proper infraorder name, whereas Anthropoidea ends in -oidea, which is reserved for superfamilies. He also noted that Anthropoidea is too easily confused with anthropoïdes, the simians are split into three distinct groups. The New World monkeys in parvorder Platyrrhini split from the rest of the line about 40 mya. This group split about 25 mya between the Old World monkeys and the apes
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Catarrhini
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Catarrhini is one of the two subdivisions of the simians, the other being the plathyrrhine. The Catarrhine are all native to Africa and Asia, members of this parvorder are called catarrhines. The technical distinction between the New World platyrrhines and Old World catarrhines is the shape of their noses, the platyrrhines have nostrils which face sideways. The catarrhines have nostrils that face downwards. Catarrhines also never have tails, and have flat fingernails and toenails, a tubular ectotympanic. Most catarrhine species show sexual dimorphism and do not form a pair bond. Most, but not all, species live in social groups, like the platyrrhines, the catarrhines are generally diurnal, and have grasping hands and grasping feet. The apes – in both traditional and phylogenic nomenclature – are exclusively catarrhine species, in traditional usage, ape describes any tailless, larger, and more typically ground-dwelling species of catarrhine. Ape may be found as part of the name of such species. In phylogenic usage, the term ape applies only to the superfamily Hominoidea, the apes and Old World monkeys split from their New World monkey kin about 35 million years ago. The major catarrhine division occurred about 25 Mya, with the gibbons separating from the great apes about 15-19 Mya, therefore, cladistically, apes, catarrhines and related contemporary extinct groups such as Parapithecidaea are monkeys as well, for any consistent definition of monkey. Order PrimatesSuborder Strepsirrhini, lemurs, lorises, etc, in May 2005, dozens of new primate fossils, mainly single teeth, were discovered in the Bugti Hills of Pakistan. They were recognised as belonging to three new species, the three primates called Bugtipithecus inexpectans, Phileosimias kamali, and Phileosimias brahuiorum all date back to the Oligocene some 30 Mya – when monkeys dominated only Africa. These were small lemur-like catarrhines that prospered in an ancient tropical rainforest, possibly these Asian catarrhines led nowhere in evolution, a side branch from Eosimias. Other possible new catarrhine fossils have been uncovered in China, Thailand, disotell T. R. Noviello C. M. Raaum R. L. Sterner K. N. Stewart C. Catarrhine primate divergence dates estimated from complete mitochondrial genomes, concordance with fossil and nuclear DNA evidence
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John Edward Gray
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John Edward Gray, FRS was a British zoologist. He was the brother of zoologist George Robert Gray and son of the pharmacologist and botanist Samuel Frederick Gray. Gray was Keeper of Zoology at the British Museum in London from 1840 until Christmas 1874 and he published several catalogues of the museum collections that included comprehensive discussions of animal groups as well as descriptions of new species. He improved the zoological collections to make them amongst the best in the world, Gray was born in Walsall, but his family soon moved to London, where Gray studied medicine. He assisted his father in writing The Natural Arrangement of British Plants, after being blackballed by the Linnean Society he turned his interest from botany to zoology. He began his career by volunteering to collect insects for the British Museum at age 15. He officially joined the Zoological Department in 1824 to help John George Children catalog the reptile collection, in 1840 he took over from Children as Keeper of Zoology, which he continued for 35 years, publishing well over 1000 papers. He named many species, genera, subfamilies, and families. During this period he collaborated with Benjamin Waterhouse Hawkins, the natural history artist. Knowsley Park, near Liverpool, had been founded by Edward Stanley, Gray married Maria Emma Smith in 1826. She helped him with his work, especially with her drawings. In 1833, he was a founder of what became the Royal Entomological Society of London, during his fifty years employed at the British Museum Gray wrote nearly 500 papers, including many descriptions of species new to science. He was also active in malacology, the study of molluscs, on his death he was buried at St Marys Church, Lewisham. 1821, On the natural arrangement of Vertebrose Animals,1824, A revision of the family Equidae. 1824, On the natural arrangement of the pulmonobranchous Mollusca, annals of Philosophy, new series 8, 107–109. 1825, A list and description of species of shells not taken notice of by Lamarck. 1825, An outline of an attempt at the disposition of the Mammalia into tribes and families with a list of the genera apparently appertaining to each tribe, narrative of a Survey of the Intertropical and Western Coasts of Australia. Performed between the years 1818 and 1822, with an Appendix, containing various subjects relating to hydrography and natural history
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Type species
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A similar concept is used for suprageneric groups called a type genus. In botanical nomenclature, these terms have no standing under the code of nomenclature. In botany, the type of a name is a specimen which is also the type of a species name. The species name that has that type can also be referred to as the type of the genus name, names of genus and family ranks, the various subdivisions of those ranks, and some higher-rank names based on genus names, have such types. In bacteriology, a species is assigned for each genus. Every named genus or subgenus in zoology, whether or not currently recognized as valid, is associated with a type species. In practice, however, there is a backlog of untypified names defined in older publications when it was not required to specify a type, a type species is both a concept and a practical system that is used in the classification and nomenclature of animals. The type species represents the species and thus definition for a particular genus name. In the Glossary, type species is defined as The nominal species that is the type of a nominal genus or subgenus. The type species permanently attaches a formal name to a genus by providing just one species within that genus to which the name is permanently linked. The species name in turn is fixed, in theory, to a type specimen, for example, the type species for the land snail genus Monacha is Monacha cartusiana. That genus is placed within the family Hygromiidae. The type genus for that family is the genus Hygromia, the concept of the type species in zoology was introduced by Pierre André Latreille. The International Code of Zoological Nomenclature states that the name of the type species should always be cited. It gives an example in Article 67.1, Astacus marinus Fabricius,1775 was later designated as the type species of the genus Homarus, thus giving it the name Homarus marinus. However, the species of Homarus should always be cited using its original name. 3, the type of the genus name Elodes is quoted as the type of the species name Hypericum aegypticum, Glossary of scientific naming Genetypes – genetic sequence data from type specimens. Holotype Paratype Principle of Typification Type Type genus
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Human
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Modern humans are the only extant members of Hominina tribe, a branch of the tribe Hominini belonging to the family of great apes. Several of these hominins used fire, occupied much of Eurasia and they began to exhibit evidence of behavioral modernity around 50,000 years ago. In several waves of migration, anatomically modern humans ventured out of Africa, the spread of humans and their large and increasing population has had a profound impact on large areas of the environment and millions of native species worldwide. Humans are uniquely adept at utilizing systems of communication for self-expression and the exchange of ideas. Humans create complex structures composed of many cooperating and competing groups, from families. Social interactions between humans have established a wide variety of values, social norms, and rituals. These human societies subsequently expanded in size, establishing various forms of government, religion, today the global human population is estimated by the United Nations to be near 7.5 billion. In common usage, the word generally refers to the only extant species of the genus Homo—anatomically and behaviorally modern Homo sapiens. In scientific terms, the meanings of hominid and hominin have changed during the recent decades with advances in the discovery, there is also a distinction between anatomically modern humans and Archaic Homo sapiens, the earliest fossil members of the species. The English adjective human is a Middle English loanword from Old French humain, ultimately from Latin hūmānus, the words use as a noun dates to the 16th century. The native English term man can refer to the species generally, the species binomial Homo sapiens was coined by Carl Linnaeus in his 18th century work Systema Naturae. The generic name Homo is a learned 18th century derivation from Latin homō man, the species-name sapiens means wise or sapient. Note that the Latin word homo refers to humans of either gender, the genus Homo evolved and diverged from other hominins in Africa, after the human clade split from the chimpanzee lineage of the hominids branch of the primates. The closest living relatives of humans are chimpanzees and gorillas, with the sequencing of both the human and chimpanzee genome, current estimates of similarity between human and chimpanzee DNA sequences range between 95% and 99%. The gibbons and orangutans were the first groups to split from the leading to the humans. The splitting date between human and chimpanzee lineages is placed around 4–8 million years ago during the late Miocene epoch, during this split, chromosome 2 was formed from two other chromosomes, leaving humans with only 23 pairs of chromosomes, compared to 24 for the other apes. There is little evidence for the divergence of the gorilla, chimpanzee. Each of these species has been argued to be an ancestor of later hominins
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Carl Linnaeus
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Carl Linnaeus, also known after his ennoblement as Carl von Linné, was a Swedish botanist, physician, and zoologist, who formalised the modern system of naming organisms called binomial nomenclature. He is known by the father of modern taxonomy. Many of his writings were in Latin, and his name is rendered in Latin as Carolus Linnæus, Linnaeus was born in the countryside of Småland, in southern Sweden. He received most of his education at Uppsala University. He lived abroad between 1735 and 1738, where he studied and also published a first edition of his Systema Naturae in the Netherlands and he then returned to Sweden, where he became professor of medicine and botany at Uppsala. In the 1740s, he was sent on journeys through Sweden to find and classify plants. In the 1750s and 1760s, he continued to collect and classify animals, plants, and minerals, at the time of his death, he was one of the most acclaimed scientists in Europe. The philosopher Jean-Jacques Rousseau sent him the message, Tell him I know no man on earth. The German writer Johann Wolfgang von Goethe wrote, With the exception of Shakespeare and Spinoza, Swedish author August Strindberg wrote, Linnaeus was in reality a poet who happened to become a naturalist. Among other compliments, Linnaeus has been called Princeps botanicorum, The Pliny of the North and he is also considered as one of the founders of modern ecology. In botany, the abbreviation used to indicate Linnaeus as the authority for species names is L. In older publications, sometimes the abbreviation Linn. is found, Linnæus was born in the village of Råshult in Småland, Sweden, on 23 May 1707. He was the first child of Nicolaus Ingemarsson and Christina Brodersonia and his siblings were Anna Maria Linnæa, Sofia Juliana Linnæa, Samuel Linnæus, and Emerentia Linnæa. One of a line of peasants and priests, Nils was an amateur botanist, a Lutheran minister. Christina was the daughter of the rector of Stenbrohult, Samuel Brodersonius, a year after Linnæus birth, his grandfather Samuel Brodersonius died, and his father Nils became the rector of Stenbrohult. The family moved into the rectory from the curates house, even in his early years, Linnæus seemed to have a liking for plants, flowers in particular. Whenever he was upset, he was given a flower, which calmed him. Nils spent much time in his garden and often showed flowers to Linnaeus, soon Linnæus was given his own patch of earth where he could grow plants
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10th edition of Systema Naturae
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The 10th edition of Systema Naturae is a book written by Carl Linnaeus and published in two volumes in 1758 and 1759, which marks the starting point of zoological nomenclature. In it, Linnaeus introduced binomial nomenclature for animals, something he had already done for plants in his 1753 publication of Species Plantarum, before 1758, most biological catalogues had used polynomial names for the taxa included, including earlier editions of Systema Naturae. The first work to consistently apply binomial nomenclature across the kingdom was the 10th edition of Systema Naturae. Names published before that date are unavailable, even if they would otherwise satisfy the rules, during Linnaeus lifetime, Systema Naturae was under continuous revision. The Animal Kingdom, Animals enjoy sensation by means of an organization, animated by a medullary substance, perception by nerves. They have members for the different purposes of life, organs for their different senses and they all originate from an egg. Their external and internal structure, their anatomy, habits, instincts. The list has been broken down into the six classes Linnaeus described for animals, Mammalia, Aves, Amphibia, Pisces, Insecta. These classes were created by studying the internal anatomy, as seen in his key. Warm, red blood Viviparous, Mammalia Oviparous, Aves Heart with 1 auricle,1 ventricle, cold, red blood Lungs voluntary, Amphibia External gills, Pisces Heart with 1 auricle,0 ventricles. Linnaeus described mammals as, Animals that suckle their young by means of lactiferous teats, in external and internal structure they resemble man, most of them are quadrupeds, and with man, their natural enemy, inhabit the surface of the Earth. The largest, though fewest in number, inhabit the ocean and they are areal, vocal, swift and light, and destitute of external ears, lips, teeth, scrotum, womb, bladder, epiglottis, corpus callosum and its arch, and diaphragm. They breathe by means of gills, which are united by a bony arch, swim by means of radiate fins. Many of them are without a head, and most of them without feet. They are principally distinguished by their tentacles, by the Ancients they were not improperly called imperfect animals, as being destitute of ears, nose, head, eyes and legs, and are therefore totally distinct from Insects. In addition to repeating the species he had listed in his Species Plantarum. The species from Species Plantarum were numbered sequentially, while the new species were labelled with letters, new plant species described in the 10th edition of Systema Naturae include, The original 1758 Systema Naturae Linnaeus 1758 Classification of Animals on the Taxonomicon
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Family (biology)
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In biological classification, family is one of the eight major taxonomic ranks, it is classified between order and genus. A family may be divided into subfamilies, which are intermediate ranks above the rank of genus. In vernacular usage, a family may be named one of its common members, for example, walnuts and hickory trees belong to the family Juglandaceae. What does or does not belong to a family—or whether a family should be recognized at all—are proposed and determined by practicing taxonomists. There are no rules for describing or recognizing a family. Taxonomists often take different positions about descriptions of taxa, and there may be no broad consensus across the community for some time. Some described taxa are accepted broadly and quickly, but others only rarely, if at all, the naming of families is codified by various international codes. In zoological nomenclature, the names of animals end with the suffix -idae. The concept of rank at time was not yet settled, and in the preface to the Prodromus Magnol spoke of uniting his families into larger genera. Carolus Linnaeus used the word familia in his Philosophia botanica to denote groups of plants, trees, herbs, ferns, palms. He used this term only in the section of the book. In zoology, the family as an intermediate between order and genus was introduced by Pierre André Latreille in his Précis des caractères génériques des insectes. He used families in some but not in all his orders of insects, families can be used for evolutionary, palaeontological and generic studies because they are more stable than lower taxonomic levels such as genera and species
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Proconsulidae
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The Proconsulidae are an early family of primates that lived during the Miocene epoch in Kenya, and was restricted to Africa. The family is split into two subfamilies, the name, Proconsul, was devised by Arthur Hopwood in 1933 and means before Consul, the name of a famous captive chimp in London. At the time Consul was being used as a name for performing chimpanzees. The Folies Bergère of 1903 in Paris had a performing chimpanzee named Consul. On the latters death in that year Ben Brierley wrote a commemorative poem wondering where the Missing Link between chimpanzees and men was, Hopwood in 1931 had discovered the fossils of three individuals while expeditioning with Louis Leakey in the vicinity of Lake Victoria. The Consul that he selected to use in the name was neither of the mentioned above. Consul is being used Linnaean-style to symbolize the chimpanzee, Proconsul is therefore ancestral to the Chimpanzee in Hopwoods words. He also added africanus as the species name, for example, Mary Leakeys famous find of 1948 began as africanus and was split from it to be lumped with Thomas Whitworths finds of 1951 as heseloni by Alan Walker in 1993. The family of Proconsulidae was first proposed by Louis Leakey, eleven years after he and Wilfrid Le Gros Clark had defined africanus, nyanzae and it was not immediately accepted but ultimately prevailed. The history of classification in the second half of the 20th century is sufficiently complex to warrant a few books itself. Most of the palaeoanthropologists have changed their minds at least once as new fossils have come to light and new observations have made, the classifications found in the literature of one decade are not generally the same as those of another. For example, in 1987 Peter Andrews and Lawrence Martin, established palaeontologists, took the point of view that Proconsul is not a Hominoid, but is a sister taxon to it
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Afropithecus
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Afropithecus turkanensis is a Miocene hominoid which was excavated from a small site near Lake Turkana called Kalodirr in northern Kenya in 1986 and was named by Richard Leakey and Meave Leakey. In total there are 46 recovered specimens from Kalodirr relating to Afropithecus consisting of cranial, mandible, dentition, the type specimen of Afropithecus turkanensis is KNM-WK16999. Richard Leakey and Meave Leakey first described Afropithecus turkanensis to be a large hominoid which appeared to have relatively thick enamel, a. turkanensis had a 6. 5mm diastema between its very procumbent second incisor and the canine. The thickness of the enamel on the molars is often reported when fossils are being recorded and used to make comparisons across taxa, the thickness is referred to either as thin or thick and is commonly assessed as a linear measurement of the enamel on worn or naturally fractured teeth. From enamel testing it has suggested that A. turkanensis is the oldest known thick-enamelled hominoid. Post cranial remains such as KNM-WK16901, includes a right fibula, a right proximal third metatarsal, a right fourth metatarsal lacking the head. Other post-cranial remains include, KNM-WK 17016P a large right ulna, dental metric comparisons of Morotopithecus and Afropithecus, implications for the validity of the genus Morotopithecus
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Gibbon
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Gibbons are apes in the family Hylobatidae. The family historically contained one genus, but now is split into four genera and 18 species, gibbons occur in tropical and subtropical rainforests from eastern Bangladesh and northeast India to southern China and Indonesia. Also called the smaller apes or lesser apes, gibbons differ from great apes in being smaller, exhibiting low sexual dimorphism, in certain anatomical details, they superficially more closely resemble monkeys than great apes do, but like all apes, gibbons are tailless. Gibbons also display pair-bonding, maintaining the same mate for life, gibbons are masters of their primary mode of locomotion, brachiation, swinging from branch to branch for distances up to 15 m, at speeds as high as 55 km/h. They can also make leaps up to 8 m, and walk bipedally with their arms raised for balance and they are the fastest and most agile of all tree-dwelling, nonflying mammals. Depending on species and sex, gibbons fur coloration varies from dark to light brown shades, Gibbon species include the siamang, the white-handed or lar gibbon, and the hoolock gibbons. Whole genome molecular dating analyses indicate that the gibbon lineage diverged from that of great apes around 16.8 million years ago, adaptive divergence associated with chromosomal rearrangements led to rapid radiation of the four genera 5-7 Mya. Whole genome analysis suggests divergence of Hylobates pileatus from Hylobates moloch 1. 5-3.0 Mya, the extinct Bunopithecus sericus is a gibbon or gibbon-like ape which, until recently, was thought to be closely related to the hoolock gibbons. The family is divided into four genera based on their diploid chromosome number, Hylobates, Hoolock, Nomascus and these morphological ambiguities have led to hybrids in zoos. Interspecific hybrids, hybrids within a genus, are suspected to occur in wild gibbons where their ranges overlap. However, no records exist of fertile hybrids between different gibbon genera, either in the wild or in captivity, one unique aspect of a gibbons anatomy is the wrist, which functions something like a ball and socket joint, allowing for biaxial movement. This greatly reduces the amount of energy needed in the arm and torso. Gibbons also have long hands and feet, with a cleft between the first and second digits of their hands. Their fur is black, gray, or brownish, often with white markings on hands, feet. Some species have a throat sac, which inflates and serves as a resonating chamber when the animals call. This structure can become large in some species, sometimes equaling the size of the animals head. Their voices are more powerful than that of any human singer. Gibbon skulls and teeth resemble those of the apes
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Clade
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A clade is a group of organisms that consists of a common ancestor and all its lineal descendants, and represents a single branch on the tree of life. The common ancestor may be an individual, a population, a species, clades are nested, one in another, as each branch in turn splits into smaller branches. These splits reflect evolutionary history as populations diverged and evolved independently, over the last few decades, the cladistic approach has revolutionized biological classification and revealed surprising evolutionary relationships among organisms. Increasingly, taxonomists try to avoid naming taxa that are not clades, the term clade was coined in 1957 by the biologist Julian Huxley to refer to the result of cladogenesis, a concept Huxley borrowed from Bernhard Rensch. Many commonly named groups are clades, for example, rodents or insects, because in each case, their name refers to a common ancestor with all its descendant branches. Rodents, for example, are a branch of mammals that split off after the end of the period when the clade Dinosauria stopped being the dominant terrestrial vertebrates 66 million years ago, the original population and all its descendants are a clade. The rodent clade corresponds to the order Rodentia, and insects to the class Insecta and these clades include smaller clades, such as chipmunk or ant, each of which consists of even smaller clades. The clade rodent is in included in the mammal, vertebrate. The phenomenon of convergent evolution is, however, responsible for cases where there are misleading similarities in the morphology of groups that evolved from different lineages. The publication of Darwins theory of evolution in 1859 gave this view increasing weight, thomas Henry Huxley, an early advocate of evolutionary theory, proposed a revised taxonomy based on clades. For example, he grouped birds with reptiles, based on fossil evidence, german biologist Emil Hans Willi Hennig is considered to be the founder of cladistics. Taxonomists have increasingly worked to make the taxonomic system reflect evolution, when it comes to naming, however, this principle is not always compatible with the traditional rank-based nomenclature. For these and other reasons, phylogenetic nomenclature has been developed, a clade is by definition monophyletic, meaning that it contains one ancestor and all its descendants. The ancestor can be known or unknown, any and all members of a clade can be extant or extinct, the science that tries to reconstruct phylogenetic trees and thus discover clades is called phylogenetics or cladistics, the latter term coined by Ernst Mayr, derived from clade. The results of analyses are tree-shaped diagrams called cladograms, they. Three methods of defining clades are featured in phylogenetic nomenclature, node-, stem-, the relationship between clades can be described in several ways, A clade located within a clade is said to be nested within that clade. In the diagram, the clade, i. e. the apes. Two clades are sisters if they have a common ancestor
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Old World
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The Old World consists of Africa, Europe, and Asia, regarded collectively as the part of the world known to Europeans before contact with the Americas. It is used in the context of, and contrast with and these regions were connected via the Silk Road trade route, and they have a pronounced Iron Age period following the Bronze Age. The concept of the three continents in the Old World, viz. Asia, Africa, and Europe, goes back to classical antiquity and their boundaries as defined by Ptolemy and other geographers of antiquity were drawn along the Nile and Don rivers. This definition remained influential throughout the Middle Ages and the Early Modern period, the Old world has areas of North Africa, Eurasia, parts of Papua New Guinea and Solomon Islands. The mainland of Afro-Eurasia has been referred to as the World Island, the term may have been coined by Sir Halford John Mackinder in The Geographical Pivot of History. The equivalent of the Old World had names in some of its ancient cultures, including Midgard in Germanic cosmology, synoptic table of the principal old world prehistoric cultures Afro-Eurasia
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Africa
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Africa is the worlds second-largest and second-most-populous continent. At about 30.3 million km² including adjacent islands, it covers 6% of Earths total surface area and 20.4 % of its land area. With 1.2 billion people as of 2016, it accounts for about 16% of the human population. The continent includes Madagascar and various archipelagos and it contains 54 fully recognized sovereign states, nine territories and two de facto independent states with limited or no recognition. Africas population is the youngest amongst all the continents, the age in 2012 was 19.7. Algeria is Africas largest country by area, and Nigeria by population, afarensis, Homo erectus, H. habilis and H. ergaster – with the earliest Homo sapiens found in Ethiopia being dated to circa 200,000 years ago. Africa straddles the equator and encompasses numerous climate areas, it is the continent to stretch from the northern temperate to southern temperate zones. Africa hosts a diversity of ethnicities, cultures and languages. In the late 19th century European countries colonized most of Africa, Africa also varies greatly with regard to environments, economics, historical ties and government systems. However, most present states in Africa originate from a process of decolonization in the 20th century, afri was a Latin name used to refer to the inhabitants of Africa, which in its widest sense referred to all lands south of the Mediterranean. This name seems to have referred to a native Libyan tribe. The name is connected with Hebrew or Phoenician ʿafar dust. The same word may be found in the name of the Banu Ifran from Algeria and Tripolitania, under Roman rule, Carthage became the capital of the province of Africa Proconsularis, which also included the coastal part of modern Libya. The Latin suffix -ica can sometimes be used to denote a land, the later Muslim kingdom of Ifriqiya, modern-day Tunisia, also preserved a form of the name. According to the Romans, Africa lay to the west of Egypt, while Asia was used to refer to Anatolia, as Europeans came to understand the real extent of the continent, the idea of Africa expanded with their knowledge. 25,4, whose descendants, he claimed, had invaded Libya, isidore of Seville in Etymologiae XIV.5.2. Suggests Africa comes from the Latin aprica, meaning sunny, massey, in 1881, stated that Africa is derived from the Egyptian af-rui-ka, meaning to turn toward the opening of the Ka. The Ka is the double of every person and the opening of the Ka refers to a womb or birthplace
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Southeast Asia
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Southeast Asia or Southeastern Asia is a subregion of Asia, consisting of the countries that are geographically south of China, east of India, west of New Guinea and north of Australia. The region lies near the intersection of geological plates, with seismic and volcanic activity. Southeast Asia consists of two regions, Mainland Southeast Asia, also known historically as Indochina, comprising Vietnam, Laos, Cambodia, Thailand, Myanmar. Maritime Southeast Asia, comprising Indonesia, East Malaysia, Singapore, Philippines, East Timor, Brunei, Cocos Islands, definitions of Southeast Asia vary, but most definitions include the area represented by the countries listed below. All of the states are members of the Association of Southeast Asian Nations, the area, together with part of South Asia, was widely known as the East Indies or simply the Indies until the 20th century. Sovereignty issues exist over some territories in the South China Sea, Papua New Guinea has stated that it might join ASEAN, and is currently an observer. Southeast Asia is geographically divided into two subregions, namely Mainland Southeast Asia and Maritime Southeast Asia, Mainland Southeast Asia includes, Maritime Southeast Asia includes, The Andaman and Nicobar Islands of India are geographically considered part of Southeast Asia. Eastern Bangladesh and the Seven Sister States of India are culturally part of Southeast Asia, the eastern half of Indonesia and East Timor are considered to be biogeographically part of Oceania. Homo sapiens reached the region by around 45,000 years ago, homo floresiensis also lived in the area up until 12,000 years ago, when they became extinct. Austronesian people, who form the majority of the population in Indonesia, Malaysia, Brunei, East Timor. Solheim and others have shown evidence for a Nusantao maritime trading network ranging from Vietnam to the rest of the archipelago as early as 5000 BC to 1 AD. The peoples of Southeast Asia, especially those of Austronesian descent, have been seafarers for thousands of years and their vessels, such as the vinta, were ocean-worthy. Magellans voyage records how much more manoeuvrable their vessels were, as compared to the European ships, Passage through the Indian Ocean aided the colonisation of Madagascar by the Austronesian people, as well as commerce between West Asia and Southeast Asia. Gold from Sumatra is thought to have reached as far west as Rome and this was later replaced by Hinduism. Theravada Buddhism soon followed in 525, in the 15th century, Islamic influences began to enter. This forced the last Hindu court in Indonesia to retreat to Bali, in Mainland Southeast Asia, Burma, Cambodia and Thailand retained the Theravada form of Buddhism, brought to them from Sri Lanka. This type of Buddhism was fused with the Hindu-influenced Khmer culture, very little is known about Southeast Asian religious beliefs and practices before the advent of Indian merchants and religious influences from the 2nd century BCE onwards. Prior to the 13th century CE, Hinduism and Buddhism were the religions in Southeast Asia
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Old World monkey
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The Old World monkeys or Cercopithecidae are a family of catarrhines, the only family in the superfamily Cercopithecoidea in the clade of Catarrhini. However, a group of monkeys still survives in Gibraltar to this day. Old World monkeys include many of the most familiar species of primates, such as baboons. Old World monkeys are medium to large in size, and range from arboreal forms, such as the monkeys, to fully terrestrial forms. By superficial appearance, Old World monkeys are unlike apes in that most have tails and, unlike the New World monkeys, technically, the distinction of catarrhines from platyrrhines depends on the structure of the nose, and the distinction of Old World monkeys from apes depends on dentition. In platyrrhines, the nostrils face sideways, while in catarrhines, for example, the colobus monkeys have stubs for thumbs to assist with their arboreal movement, the proboscis monkey has an extraordinary nose, while the snub-nosed monkeys have almost no nose at all. The male mandrills penis is red and the scrotum is lilac, the coloration is more pronounced in dominant males. Most Old World monkeys are at least partially omnivorous, but all prefer plant matter, the Barbary macaques diet consists mostly of leaves and roots, though it will also eat insects and uses cedar trees as a water source. Gestation in the Old World monkeys lasts between five and seven months, births are usually single, although, as with humans, twins occur from time to time. The young are born relatively well-developed, and are able to cling onto their mothers fur with their hands from birth, compared with most other mammals, they take a long time to reach sexual maturity, with four to six years being typical of most species. In most species, daughters remain with their mothers for life, males leave the group on reaching adolescence, and find a new troop to join. In many species, only an adult male lives with each group, driving off all rivals. Group sizes are variable, even within species, depending on the availability of food. Therefore, cladistically, apes, catarrhines and related contemporary extinct groups such as Parapithecidaea are monkeys as well, for any consistent definition of monkey
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Brachiation
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Brachiation, or arm swinging, is a form of arboreal locomotion in which primates swing from tree limb to tree limb using only their arms. During brachiation, the body is supported under each forelimb. This form of locomotion is the means of locomotion for the small gibbons. Some New World monkeys such as monkeys and muriquis are semibrachiators and move through the trees with a combination of leaping. Some New World species also practice suspensory behaviors by using their prehensile tail, some traits that allow primates to brachiate include a short spine, short fingernails, long curved fingers, reduced thumbs, long forelimbs and freely rotating wrists. Modern humans retain many characteristics that suggest a brachiator ancestor, including flexible shoulder joints. In lesser apes, these characteristics were adaptations for brachiation, although great apes do not normally brachiate, our human anatomy suggests that brachiation may be an exaptation to bipedalism, and healthy modern humans are still capable of brachiating. Some childrens parks include monkey bars which children play on by brachiating, as well as shaping the evolution of gibbon body structure, brachiation has influenced the style and order of their behaviour. For example, unlike other primates who carry infants on their back and it also affects their play activities, copulation, and fighting. It is thought that gibbons gain evolutionary advantages through brachiation and being suspended by both hands when feeding, another theory postulates that brachiation is a quieter and less obvious mode of locomotion than quadrupedal jumping and climbing thereby more successfully avoiding predators
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Lar gibbon
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The lar gibbon, also known as the white-handed gibbon, is a primate in the gibbon family, Hylobatidae. It is one of the gibbons and is often seen in zoos. The hands and feet are white-colored, likewise a ring of white hair surrounds the black face, both males and females can be all color variants, and the sexes also hardly differ in size. Gibbons are true brachiators, propelling themselves through the forest by swinging under the branches using their arms, as with all apes, the number of caudal vertebrae has been reduced drastically, resulting in the loss of a functional tail. Gibbons have tough, bony padding on their buttocks, known as the ischial callosities, lar gibbons have the greatest north-south range of any of the gibbon species. They are found in Indonesia, Laos, Malaysia, Myanmar and their range historically extended from southwest China to Thailand and Burma south to the whole Malay Peninsula in primary and secondary tropical rain forests. It is also present in the northwest portion of the island of Sumatra, in recent decades, especially, the continental range has been reduced and fragmented. Lar gibbons are likely extinct in China, but if they still exist, they would only be found in southwest Yunnan and they are not usually found higher than 1200 meters above sea level. The gibbon genus is highly allopatric, usually separated by large rivers, as a result, their range extends through southern and eastern Myanmar, but only east of the Salween River. They are found through the Malay Peninsula, lar gibbons also exist west of the Mekong River in northwestern Laos and northern Sumatra. The lar gibbon is considered frugivorous with fruit constituting 50% of its diet and its dental formula is 2.1.2. 3/2.1.2.3, the generalized formula for Old World monkeys and apes. The dental arcade is U-shaped, and the mandible is thin, the incisors are broad and flat, while the molars have low, rounded cusps with thick enamel. The most noticeable characteristic of the dentition of Hylobates lar is the presence of large and these canines are not sexually dimorphic. Lar gibbons are diurnal and arboreal, inhabiting rain forests, lar gibbons are usually active for an average of 8.7 hours per day, leaving their sleeping sites right around sunrise and entering sleeping trees an average of 3.4 hours before sunset. They rarely come to the ground, instead using their arms to brachiate through the trees. With their hooked hands, they can move swiftly with great momentum, although they rarely come to the ground naturally, while there, they walk bipedally with arms raised above their heads for balance. Their social organization is dominated by monogamous family pairs, with one breeding male, when a juvenile reaches sexual maturity, it is expelled from the family unit. However, this conception has come under scrutiny
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Siamang
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The siamang is an arboreal black-furred gibbon native to the forests of Malaysia, Thailand, and Sumatra. The largest of the gibbons, the siamang can be twice the size of other gibbons, reaching 1 m in height, the siamang is the only species in the genus Symphalangus. The siamang is distinctive for two reasons, the first is that two digits on each foot are partially joined by a membrane—hence the name syndactylus, from the Ancient Greek sun-, united + daktulos, finger. The second is the gular sac, which is a throat pouch that can be inflated to the size of the siamangs head, allowing it to make loud. Two subspecies of the siamang may exist, if so, they are the nominate Sumatran siamang and the Malaysian siamang. Otherwise, the Malaysian individuals are only a population, the siamang occurs sympatrically with other gibbons, its two ranges are entirely within the combined ranges of the agile gibbon and the lar gibbon. The siamang can live to around 40 years in captivity, while the illegal pet trade takes a toll on wild populations, the principal threat to the siamang is habitat loss in both Malaysia and Sumatra. The palm oil industry is clearing large swaths of forest, reducing the habitat of the siamang, along with that of other species. The siamang lives in groups of up to six individuals with a home range of 23 hectares. Their day ranges are smaller than those of sympatric Hylobates species. The siamangs melodious singing breaks the silence in the early morning after the agile gibbons or lar gibbons calls. The siamang in Sumatra and the Malay Peninsula are similar in appearance, the siamang has long, dense, shaggy hair which is the darkest shade of all gibbons. The ape has long, gangling arms that are longer than its legs, the average length of a siamang is 90 cm, but the largest they have ever grown is 1 m 50 cm. The face of this large gibbon is mostly hairless apart from a thin mustache, the siamang eats mainly various parts of plants. The Sumatran siamang is more frugivorous than its Malayan relative, with making up to 60% of its diet. The siamang eats at least 160 species of plants, from vines to woody plants and its major food is figs (Ficus spp. The siamang prefers to eat ripe rather than fruit. It eats flowers and a few animals, mostly insects, when the siamang eats large flowers, it eats only the corollae, but it eats all parts of smaller flowers, with the small fruit collected in its hand before being consumed
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Asia
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Asia covers an area of 44,579,000 square kilometres, about 30% of Earths total land area and 8. 7% of the Earths total surface area. The continent, which has long been home to the majority of the population, was the site of many of the first civilizations. Asia is notable for not only its large size and population. In general terms, Asia is bounded on the east by the Pacific Ocean, on the south by the Indian Ocean, the western boundary with Europe is a historical and cultural construct, as there is no clear physical and geographical separation between them. The most commonly accepted boundaries place Asia to the east of the Suez Canal, the Ural River, and the Ural Mountains, and south of the Caucasus Mountains, China and India alternated in being the largest economies in the world from 1 to 1800 A. D. The accidental discovery of America by Columbus in search for India demonstrates this deep fascination, the Silk Road became the main East-West trading route in the Asian hitherland while the Straits of Malacca stood as a major sea route. Asia has exhibited economic dynamism as well as robust population growth during the 20th century, given its size and diversity, the concept of Asia—a name dating back to classical antiquity—may actually have more to do with human geography than physical geography. Asia varies greatly across and within its regions with regard to ethnic groups, cultures, environments, economics, historical ties, the boundary between Asia and Africa is the Red Sea, the Gulf of Suez, and the Suez Canal. This makes Egypt a transcontinental country, with the Sinai peninsula in Asia, the border between Asia and Europe was historically defined by European academics. In Sweden, five years after Peters death, in 1730 Philip Johan von Strahlenberg published a new atlas proposing the Urals as the border of Asia, the Russians were enthusiastic about the concept, which allowed them to keep their European identity in geography. Tatishchev announced that he had proposed the idea to von Strahlenberg, the latter had suggested the Emba River as the lower boundary. Over the next century various proposals were made until the Ural River prevailed in the mid-19th century, the border had been moved perforce from the Black Sea to the Caspian Sea into which the Ural River projects. The border between the Black Sea and the Caspian is usually placed along the crest of the Caucasus Mountains, the border between Asia and the loosely defined region of Oceania is usually placed somewhere in the Malay Archipelago. The terms Southeast Asia and Oceania, devised in the 19th century, have had several different geographic meanings since their inception. The chief factor in determining which islands of the Malay Archipelago are Asian has been the location of the possessions of the various empires there. Lewis and Wigen assert, The narrowing of Southeast Asia to its present boundaries was thus a gradual process, Asia is larger and more culturally diverse than Europe. It does not exactly correspond to the borders of its various types of constituents. From the time of Herodotus a minority of geographers have rejected the three-continent system on the grounds there is no or is no substantial physical separation between them
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Bipedalism
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Bipedalism is a form of terrestrial locomotion where an organism moves by means of its two rear limbs or legs. An animal or machine that moves in a bipedal manner is known as a biped /ˈbaɪpɛd/. Types of bipedal movement include walking, running, or hopping, few modern species are habitual bipeds whose normal method of locomotion is two-legged. A larger number of modern species intermittently or briefly use a bipedal gait, several non-archosaurian lizard species move bipedally when running, usually to escape from threats. Many primate and bear species will adopt a bipedal gait in order to reach food or explore their environment, several arboreal primate species, such as gibbons and indriids, exclusively walk on two legs during the brief periods they spend on the ground. Many animals rear up on their hind legs whilst fighting or copulating, some animals commonly stand on their hind legs, in order to reach food, to keep watch, to threaten a competitor or predator, or to pose in courtship, but do not move bipedally. The word is derived from the Latin words bi two and ped- foot, as contrasted with quadruped four feet, limited and exclusive bipedalism can offer a species several advantages. While upright, non-locomotory limbs become free for other uses, including manipulation, flight, digging, even though bipedalism is slower at first, over long distances, it has allowed humans to outrun most other animals according to the endurance running hypothesis. Bipedality in kangaroo rats has been hypothesized to improve locomotor performance, zoologists often label behaviors, including bipedalism, as facultative or obligate. Even this distinction is not completely clear-cut — for example, humans other than infants normally walk and run in biped fashion, but almost all can crawl on hands and knees when necessary. Even if one ignores exceptions caused by some kind of injury or illness, there are many cases, including the fact that normal humans can crawl on hands. This article therefore avoids the terms facultative and obligate, and focuses on the range of styles of locomotion used by various groups of animals. There are a number of states of movement commonly associated with bipedalism, in most bipeds this is an active process, requiring constant adjustment of balance. One foot in front of another, with at least one foot on the ground at any time, one foot in front of another, with periods where both feet are off the ground. Moving by a series of jumps with both feet moving together, the great majority of living terrestrial vertebrates are quadrupeds, with bipedalism exhibited by only a handful of living groups. Humans, gibbons and large birds walk by raising one foot at a time, on the other hand, most macropods, smaller birds, lemurs and bipedal rodents move by hopping on both legs simultaneously. Tree kangaroos are able to walk or hop, most commonly alternating feet when moving arboreally, there are no known living or fossil bipedal amphibians. Many species of lizards become bipedal during high-speed, sprint locomotion, including the worlds fastest lizard, the first known biped is the bolosaurid Eudibamus whose fossils date from 290 million years ago
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Orangutan
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The orangutans are the two exclusively Asian species of extant great apes. Native to Indonesia and Malaysia, orangutans are found in only the rainforests of Borneo. Classified in the genus Pongo, orangutans were considered to be one species, since 1996, they have been divided into two species, the Bornean orangutan and the Sumatran orangutan. In addition, the Bornean species is divided three subspecies. Based on genome sequencing, the two extant orangutan species evidently diverged around 400,000 years ago, the ancestors of the Ponginae subfamily split from the main ape line in Africa 16 to 19 million years ago and spread into Asia. Orangutans are the most arboreal of the apes and spend most of their time in trees. Their hair is typically reddish-brown, instead of the brown or black hair typical of chimpanzees, males and females differ in size and appearance. Dominant adult males have distinctive cheek pads and produce long calls that attract females, younger males do not have these characteristics and resemble adult females. Orangutans are the most solitary of the apes, with social bonds occurring primarily between mothers and their dependent offspring, who stay together for the first two years. Fruit is the most important component of a diet, however. They can live over 30 years in both the wild and captivity, orangutans are among the most intelligent primates, they use a variety of sophisticated tools and construct elaborate sleeping nests each night from branches and foliage. The apes have been studied for their learning abilities. There may even be distinctive cultures within populations, field studies of the apes were pioneered by primatologist Birutė Galdikas. Both orangutan species are considered to be critically endangered, human activities have caused severe declines in the populations and ranges of both species. Threats to wild populations include poaching, habitat destruction. Several conservation and rehabilitation organisations are dedicated to the survival of orangutans in the wild, the name orangutan is derived from the Malay and Indonesian words orang meaning person and hutan meaning forest, thus person of the forest. Orang Hutan was originally not used to refer to apes, the Malay words used to refer specifically to the ape are maias and mawas, but it is unclear if those words refer to just orangutans, or to all apes in general. The word appeared in several German-language descriptions of Indonesian zoology in the 17th century, the likely origin of the word comes specifically from the Banjarese variety of Malay
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Gorilla
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Gorillas are ground-dwelling, predominantly herbivorous apes that inhabit the forests of central Africa. The eponymous genus Gorilla is divided into two species, the gorillas and the western gorillas, and either four or five subspecies. They are the largest living primates, the DNA of gorillas is highly similar to that of humans, from 95–99% depending on what is counted, and they are the next closest living relatives to humans after the chimpanzees and bonobos. Gorillas natural habitats cover tropical or subtropical forests in Africa, although their range covers a small percentage of Africa, gorillas cover a wide range of elevations. The mountain gorilla inhabits the Albertine Rift montane cloud forests of the Virunga Volcanoes, ranging in altitude from 2, 200–4,300 metres. The word gorilla comes from the history of Hanno the Navigator, members of the expedition encountered a savage people, the greater part of whom were women, whose bodies were hairy, and who our interpreters called Gorillae. The word was later used as the species name, though it is unknown whether what these ancient Carthaginians encountered were truly gorillas, another species of ape or monkeys. The American physician and missionary Thomas Staughton Savage and naturalist Jeffries Wyman first described the gorilla in 1847 from specimens obtained in Liberia. The name was derived from Ancient Greek Γόριλλαι, meaning tribe of hairy women, the closest relatives of gorillas are the other two Homininae species, chimpanzees and humans, all of them having diverged from a common ancestor about 7 million years ago. Human gene sequences differ only 1. 6% on average from the sequences of corresponding gorilla genes, until recently, gorillas were considered to be a single species, with three subspecies, the western lowland gorilla, the eastern lowland gorilla and the mountain gorilla. There is now agreement that there are two species, each with two subspecies, more recently, a third subspecies has been claimed to exist in one of the species. The separate species and subspecies developed from a type of gorilla during the Ice Age. Primatologists continue to explore the relationships between various gorilla populations, the species and subspecies listed here are the ones upon which most scientists agree. The proposed third subspecies of Gorilla beringei, which has not yet received a trinomen, is the Bwindi population of the mountain gorilla, some variations that distinguish the classifications of gorilla include varying density, size, hair colour, length, culture, and facial widths. Now, over 100,000 western lowland gorillas are thought to exist in the wild, with 4,000 in zoos, mountain gorillas are the most severely endangered, with an estimated population of about 880 left in the wild and none in zoos. Population genetics of the lowland gorillas suggest that the western and eastern lowland populations diverged ~261 thousand years ago, gorillas move around by knuckle-walking, although they sometimes walk bipedally for short distances while carrying food or in defensive situations. Wild male gorillas weigh 135 to 180 kg while females usually weigh half as much as adult males at 70–115 kg. Adult males are 1.7 to 1.8 m tall, female gorillas are shorter, with smaller arm spans
44.
Chimpanzee
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Chimpanzees are one of the two species of the genus Pan, the other being the bonobo. Together with gorillas, they are the only exclusively African species of ape that are currently extant. Native to sub-Saharan Africa, both chimpanzees and bonobos are found in the Congo jungle. In addition, P. troglodytes is divided into four subspecies, based on genome sequencing, the two extant Pan species diverged around one million years ago. The most obvious differences are that chimpanzees are somewhat larger, more aggressive and male-dominated, while the bonobos are more gracile, peaceful and their hair is typically black or brown. Males and females differ in size and appearance, both chimps and bonobos are some of the most social great apes, with social bonds occurring among individuals in large communities. Fruit is the most important component of a diet, however, they will also eat vegetation, bark, honey, insects. They can live over 30 years in both the wild and captivity, Chimpanzees and bonobos are equally humanitys closest living relatives. As such, they are among the largest-brained, and most intelligent of primates, they use a variety of sophisticated tools and construct elaborate sleeping nests each night from branches and they have both been extensively studied for their learning abilities. There may even be distinctive cultures within populations, field studies of Pan troglodytes were pioneered by primatologist Jane Goodall. Both Pan species are considered to be endangered as human activities have caused declines in the populations. Threats to wild populations include poaching, habitat destruction. Several conservation and rehabilitation organisations are dedicated to the survival of Pan species in the wild, the first use of the name chimpanze is recorded in The London Magazine in 1738, glossed as meaning mockman in a language of the Angolans. The spelling chimpanzee is found in a 1758 supplement to Chambers Cyclopædia, the colloquialism chimp was most likely coined some time in the late 1870s. The common chimpanzee was named Simia troglodytes by Johann Friedrich Blumenbach in 1776, the species name troglodytes is a reference to the Troglodytae, an African people described by Greco-Roman geographers. Blumenbach first used it in his De generis humani varietate nativa liber in 1776, the genus name Pan was first introduced by Lorenz Oken in 1816. An alternative Theranthropus was suggested by Brookes 1828 and Chimpansee by Voigt 1831, troglodytes was not available, as it had been given as the name of a genus of wren in 1809. The International Commission on Zoological Nomenclature adopted Pan as the official name of the genus in 1895
45.
Tree
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In botany, a tree is a perennial plant with an elongated stem, or trunk, supporting branches and leaves in most species. In some usages, the definition of a tree may be narrower, including only woody plants with secondary growth, plants that are usable as lumber or plants above a specified height. Trees are not a group but include a variety of plant species that have independently evolved a woody trunk. In looser senses, the palms, the tree ferns, bananas. Trees tend to be long-lived, some reaching several thousand years old, the tallest known tree, a coast redwood named Hyperion, stands 115.6 m high. Trees have been in existence for 370 million years and it is estimated that there are just over 3 trillion mature trees in the world. A tree typically has many secondary branches supported clear of the ground by the trunk and this trunk typically contains woody tissue for strength, and vascular tissue to carry materials from one part of the tree to another. For most trees it is surrounded by a layer of bark which serves as a protective barrier, below the ground, the roots branch and spread out widely, they serve to anchor the tree and extract moisture and nutrients from the soil. Above ground, the divide into smaller branches and shoots. The shoots typically bear leaves, which light energy and convert it into sugars by photosynthesis, providing the food for the trees growth. Flowers and fruit may also be present, but some trees, such as conifers, instead have pollen cones and seed cones, others, such as tree ferns, trees play a significant role in reducing erosion and moderating the climate. They remove carbon dioxide from the atmosphere and store large quantities of carbon in their tissues, trees and forests provide a habitat for many species of animals and plants. Tropical rainforests are one of the most biodiverse habitats in the world, trees provide shade and shelter, timber for construction, fuel for cooking and heating, and fruit for food as well as having many other uses. In parts of the world, forests are shrinking as trees are cleared to increase the amount of available for agriculture. Because of their longevity and usefulness, trees have always revered, with sacred groves in various cultures. Although tree is a term of common parlance, there is no universally recognised precise definition of what a tree is, either botanically or in common language. In its broadest sense, a tree is any plant with the form of an elongated stem, or trunk. Trees are also defined by height, with smaller plants from 0.5 to 10 m being called shrubs
46.
Frugivore
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A frugivore /fruːdʒᵻvɔːr/ is a fruit eater. It can be any type of herbivore or omnivore where fruit is a preferred food type, because approximately 20% of all mammalian herbivores also eat fruit, frugivory is common among mammals. Since frugivores eat a lot of fruit, they are dependent on the abundance. Frugivores can either benefit fruit-producing plants by dispersing seeds, or they can hinder plants by digesting seeds along with the fruits, when both the fruit-producing plant and the frugivore species benefit by fruit-eating behavior, their interaction is called mutualism. Seed dispersal is important for plants because it allows their progeny to move away from their parents over time, the advantages of seed dispersal may have led to the evolution of fleshy fruits, which entice animals to eat the fruits and move the plants seeds from place to place. While many fruit-producing plant species would not disperse far without frugivores, many types of animals are seed dispersers. Mammal and bird species represent the majority of seed-dispersing species, however, frugivorous tortoises, lizards, amphibians, and even fish also disperse seeds. For example, cassowaries are a species because they spread fruit through digestion. While frugivores and fruit-producing plant species are present worldwide, there is evidence that tropical forests have more frugivore seed dispersers than the temperate zone. Frugivore seed dispersal is a phenomenon in many ecosystems. However, it is not a specific type of plant–animal interaction. For example, a species of frugivorous bird may disperse fruits from several species of plants. This lack of specialization could be because fruit availability varies by season and year, furthermore, different seed dispersers tend to disperse seeds to different habitats, at different abundances, and distances, depending on their behavior and numbers. There are a number of characteristics that seem to be adaptive characteristics to attract frugivores. Many animal-dispersed fruits advertise their palatability to animals with bright colors, Fruit pulp is generally rich in water and carbohydrates and low in protein and lipids. However, the nutritional composition of fruits varies widely. The seeds of animal-dispersed fruits are often adapted to survive digestion by frugivores, for example, seeds can become more permeable to water after passage through an animals gut. This leads to higher germination rates, some mistletoe seeds even germinate inside the dispersers intestine
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Chisholm, Hugh, ed. (1911). "Ape". Encyclopædia Britannica. 2 (11th ed.). Cambridge University Press. p. 160.
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Chisholm, Hugh, ed. (1911). "
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