1.
Taxonomy (biology)
–
Taxonomy is the science of defining groups of biological organisms on the basis of shared characteristics and giving names to those groups. The exact definition of taxonomy varies from source to source, but the core of the remains, the conception, naming. There is some disagreement as to whether biological nomenclature is considered a part of taxonomy, the broadest meaning of taxonomy is used here. The word taxonomy was introduced in 1813 by Candolle, in his Théorie élémentaire de la botanique, the term alpha taxonomy is primarily used today to refer to the discipline of finding, describing, and naming taxa, particularly species. In earlier literature, the term had a different meaning, referring to morphological taxonomy, ideals can, it may be said, never be completely realized. They have, however, a value of acting as permanent stimulants. Some of us please ourselves by thinking we are now groping in a beta taxonomy, turrill thus explicitly excludes from alpha taxonomy various areas of study that he includes within taxonomy as a whole, such as ecology, physiology, genetics, and cytology. He further excludes phylogenetic reconstruction from alpha taxonomy, thus, Ernst Mayr in 1968 defined beta taxonomy as the classification of ranks higher than species. This activity is what the term denotes, it is also referred to as beta taxonomy. How species should be defined in a group of organisms gives rise to practical and theoretical problems that are referred to as the species problem. The scientific work of deciding how to define species has been called microtaxonomy, by extension, macrotaxonomy is the study of groups at higher taxonomic ranks, from subgenus and above only, than species. While some descriptions of taxonomic history attempt to date taxonomy to ancient civilizations, earlier works were primarily descriptive, and focused on plants that were useful in agriculture or medicine. There are a number of stages in scientific thinking. Early taxonomy was based on criteria, the so-called artificial systems. Later came systems based on a complete consideration of the characteristics of taxa, referred to as natural systems, such as those of de Jussieu, de Candolle and Bentham. The publication of Charles Darwins Origin of Species led to new ways of thinking about classification based on evolutionary relationships and this was the concept of phyletic systems, from 1883 onwards. This approach was typified by those of Eichler and Engler, the advent of molecular genetics and statistical methodology allowed the creation of the modern era of phylogenetic systems based on cladistics, rather than morphology alone. Taxonomy has been called the worlds oldest profession, and naming and classifying our surroundings has likely been taking place as long as mankind has been able to communicate
Taxonomy (biology)
–
Title page of
Systema Naturae, Leiden, 1735
Taxonomy (biology)
–
Evolution of the
vertebrates at class level, width of spindles indicating number of families. Spindle diagrams are typical for
Evolutionary taxonomy
Taxonomy (biology)
–
The same relationship, expressed as a
cladogram typical for
cladistics
Taxonomy (biology)
–
Type specimen for
Nepenthes smilesii, a tropical
pitcher plant.
2.
Plantae
–
Plants are mainly multicellular, predominantly photosynthetic eukaryotes of the kingdom Plantae. The term is generally limited to the green plants, which form an unranked clade Viridiplantae. This includes the plants, conifers and other gymnosperms, ferns, clubmosses, hornworts, liverworts, mosses and the green algae. Green plants have cell walls containing cellulose and obtain most of their energy from sunlight via photosynthesis by primary chloroplasts and their chloroplasts contain chlorophylls a and b, which gives them their green color. Some plants are parasitic and have lost the ability to produce amounts of chlorophyll or to photosynthesize. Plants are characterized by sexual reproduction and alternation of generations, although reproduction is also common. There are about 300–315 thousand species of plants, of which the great majority, green plants provide most of the worlds molecular oxygen and are the basis of most of Earths ecologies, especially on land. Plants that produce grains, fruits and vegetables form humankinds basic foodstuffs, Plants play many roles in culture. They are used as ornaments and, until recently and in variety, they have served as the source of most medicines. The scientific study of plants is known as botany, a branch of biology, Plants are one of the two groups into which all living things were traditionally divided, the other is animals. The division goes back at least as far as Aristotle, who distinguished between plants, which generally do not move, and animals, which often are mobile to catch their food. Much later, when Linnaeus created the basis of the system of scientific classification. Since then, it has become clear that the plant kingdom as originally defined included several unrelated groups, however, these organisms are still often considered plants, particularly in popular contexts. When the name Plantae or plant is applied to a group of organisms or taxon. The evolutionary history of plants is not yet settled. Those which have been called plants are in bold, the way in which the groups of green algae are combined and named varies considerably between authors. Algae comprise several different groups of organisms which produce energy through photosynthesis, most conspicuous among the algae are the seaweeds, multicellular algae that may roughly resemble land plants, but are classified among the brown, red and green algae. Each of these groups also includes various microscopic and single-celled organisms
Plantae
–
Green algae from
Ernst Haeckel 's
Kunstformen der Natur, 1904.
Plantae
–
Had'n
Plantae
–
Dicksonia antarctica, a species of
tree fern
Plantae
–
A petrified log in
Petrified Forest National Park.
3.
Angiosperms
–
The flowering plants, also known as Angiospermae or Magnoliophyta, are the most diverse group of land plants, with 416 families, approx. 13,164 known genera and a total of c.295,383 known species, etymologically, angiosperm means a plant that produces seeds within an enclosure, in other words, a fruiting plant. The term angiosperm comes from the Greek composite word meaning enclosed seeds, the ancestors of flowering plants diverged from gymnosperms in the Triassic Period, during the range 245 to 202 million years ago, and the first flowering plants are known from 160 mya. They diversified extensively during the Lower Cretaceous, became widespread by 120 mya, angiosperms differ from other seed plants in several ways, described in the table. These distinguishing characteristics taken together have made the angiosperms the most diverse and numerous land plants, the amount and complexity of tissue-formation in flowering plants exceeds that of gymnosperms. The vascular bundles of the stem are arranged such that the xylem and phloem form concentric rings, in the dicotyledons, the bundles in the very young stem are arranged in an open ring, separating a central pith from an outer cortex. In each bundle, separating the xylem and phloem, is a layer of meristem or active formative tissue known as cambium, the soft phloem becomes crushed, but the hard wood persists and forms the bulk of the stem and branches of the woody perennial. Among the monocotyledons, the bundles are more numerous in the stem and are scattered through the ground tissue. They contain no cambium and once formed the stem increases in diameter only in exceptional cases, the characteristic feature of angiosperms is the flower. Flowers show remarkable variation in form and elaboration, and provide the most trustworthy external characteristics for establishing relationships among angiosperm species, the function of the flower is to ensure fertilization of the ovule and development of fruit containing seeds. The floral apparatus may arise terminally on a shoot or from the axil of a leaf, occasionally, as in violets, a flower arises singly in the axil of an ordinary foliage-leaf. There are two kinds of cells produced by flowers. Microspores, which divide to become pollen grains, are the male cells and are borne in the stamens. The female cells called megaspores, which divide to become the egg cell, are contained in the ovule. The flower may consist only of parts, as in willow. Usually, other structures are present and serve to protect the sporophylls, the individual members of these surrounding structures are known as sepals and petals. The outer series is usually green and leaf-like, and functions to protect the rest of the flower, the inner series is, in general, white or brightly colored, and is more delicate in structure. It functions to attract insect or bird pollinators, attraction is effected by color, scent, and nectar, which may be secreted in some part of the flower
Angiosperms
Angiosperms
–
Bud of a pink
rose
Angiosperms
–
Flowers of
Malus sylvestris (crab apple)
Angiosperms
–
Flowers and leaves of
Oxalis pes-caprae (Bermuda buttercup)
4.
Eudicots
–
The eudicots, Eudicotidae or eudicotyledons are a monophyletic clade of flowering plants that had been called tricolpates or non-magnoliid dicots by previous authors. The close relationships among flowering plants with tricolpate pollen grains was initially seen in studies of shared derived characters. These plants have a trait in their pollen grains of exhibiting three colpi or grooves paralleling the polar axis. Later molecular evidence confirmed the basis for the evolutionary relationships among flowering plants with tricolpate pollen grains. The term means true dicotyledons, as it contains the majority of plants that have been considered dicots and have characteristics of the dicots, the term eudicots has subsequently been widely adopted in botany to refer to one of the two largest clades of angiosperms, monocots being the other. The remaining angiosperms are sometimes referred to as basal angiosperms or paleodicots, the other name for the eudicots is tricolpates, a name which refers to the grooved structure of the pollen. Members of the group have tricolpate pollen, or forms derived from it and these pollens have three or more pores set in furrows called colpi. In contrast, most of the seed plants produce monosulcate pollen. The name tricolpates is preferred by some botanists to avoid confusion with the dicots, numerous familiar plants are eudicots, including many common food plants, trees, and ornamentals. Most leafy trees of midlatitudes also belong to eudicots, with exceptions being magnolias and tulip trees which belong to magnoliids, and Ginkgo biloba. The name eudicots is used in the APG system, of 1998 and it is applied to a clade, a monophyletic group, which includes most of the dicots. The eudicots can be divided into two groups, the basal eudicots and the core eudicots, basal eudicot is an informal name for a paraphyletic group. The core eudicots are a monophyletic group, a 2010 study suggested the core eudicots can be divided into two clades, Gunnerales and a clade called Pentapetalae, comprising all the remaining core eudicots
Eudicots
–
Eudicots Temporal range:
Early Cretaceous - recent
Eudicots
–
Arabis pollen has three colpi.
5.
Asterids
–
In the APG IV system for the classification of flowering plants, the name asterids denotes a clade. Most of the taxa belonging to this clade had been referred to the Asteridae in the Cronquist system, the name asterids resembles the earlier botanical name but is intended to be the name of a clade rather than a formal ranked name, in the sense of the ICBN. The phylogenetic tree presented hereafter has been proposed by the APG IV project, genetic analysis carried out after APG II maintains that the sister to all other asterids are the Cornales. A second order that split from the base of the asterids are the Ericales, the remaining orders cluster into two clades, the lamiids and the campanulids. The structure of both of these clades has changed in APG III, in APG III system, the following clades were renamed, euasterids I → lamiids euasterids II → campanulids Asterids in Stevens, P. F. Angiosperm Phylogeny Website
Asterids
–
Asterids
Asterids
–
Oregano from Lamiales
6.
Ericales
–
The Ericales are a large and diverse order of dicotyledons, including, for example, tea, persimmon, blueberry, Brazil nut, and azalea. The order includes trees, bushes, lianas, and herbaceous plants, together with ordinary autophytic plants, the Ericales include chlorophyll-deficient mycoheterotrophic plants and carnivorous plants. Many species have five petals, often grown together, fusion of the petals as a trait was traditionally used to place the order in the subclass Sympetalae. Mycorrhiza is a property, frequently associated with the Ericales. Indeed, symbiosis with fungi is quite common among the order representatives. In addition, some families among the order are notable for their ability to accumulate aluminum. Areas of distribution of families vary largely - while some are restricted to tropics, the entire order contains over 8,000 species, of which the Ericaceae account for 2, 000-4,000 species. The most commercially used plant in the order is tea from the Theaceae family, the order also includes some edible fruits, including kiwifruit, persimmon, blueberry, huckleberry, cranberry, Brazil nut, and Mamey sapote. The order also includes shea, which is the dietary lipid source for millions of sub-Saharan Africans. Many Ericales species are cultivated for their flowers, well-known examples are azalea, rhododendron, camellia, heather, polyanthus, cyclamen, phlox. These families are typical of newer classifications and those marked with an asterisk are recognized in the APG III system. Analyse des Familles de Plantes, avec lindication des principaux genres qui sy rattachent,28, S. Jansen, T. Watanabe, P. Caris, K. Geuten, F. Lens, N. Pyck, E. Smets. The Distribution and Phylogeny of Aluminium Accumulating Plants in the Ericales, W. S. Judd, C. S. Campbell, E. A. Kellogg, P. F. Stevens, M. J. Donoghue. Plant Systematics, A Phylogenetic Approach, 2nd edition, in, Nature Encyclopedia of Life Sciences. Arne A. Anderberg, Bertil Stahl, Mari Kallersjo, maesaceae, a New Primuloid Family in the Order Ericales s. l
Ericales
–
Ericales
Ericales
–
Impatiens balsamina from the
Balsaminaceae
Ericales
–
Primula rosea from the
Primulaceae
7.
Ericaceae
–
The Ericaceae are a family of flowering plants, commonly known as the heath or heather family, found most commonly in acid and infertile growing conditions. The family is large, with c.4250 known species spread across 124 genera, the many well-known and economically important members of the Ericaceae include the cranberry, blueberry, huckleberry, rhododendron, and various common heaths and heathers. The Ericaceae contain a diverse range of taxa, including herbs, dwarf shrubs, shrubs. Their leaves are alternate or whorled, simple and without stipules. Their flowers are hermaphrodite and show considerable variability, the petals are often fused with shapes ranging from narrowly tubular to funnelform or widely bowl-shaped. The corollas are usually symmetrical, but many flowers of the genus Rhododendron are somewhat bilaterally symmetrical. Adanson used the term Vaccinia to describe a family. The name comes from the type genus Erica, which appears to be derived from the Greek word ereike, the exact meaning is difficult to interpret, but some sources show it as meaning heather. Historically, the Ericaceae included both subfamilies and tribes, within tribe Rhodoreae, five genera were described, Rhododendron L. Therorhodion Small, Ledum L. Tsusiophyllum Max. Menziesia J. E. Smith, that were transferred into Rhododendron. The move significantly increased the morphological and geographical range found within the group, the Ericaceae have a nearly worldwide distribution. They are absent from continental Antarctica, parts of the high Arctic, central Greenland, northern and central Australia, the family is largely composed of plants that can tolerate acidic, infertile conditions. Like other stress-tolerant plants, many Ericaceae have mycorrhizal fungi to assist with extracting nutrients from infertile soils and this trait is not found in the Clethraceae and Cyrillaceae, the two families most closely related to the Ericaceae. Most Ericaceae form a distinctive accumulation of mycorrhizae, in which grow in and around the roots. The Pyroloideae are mixotrophic and gain sugars from the mycorrhizae, as well as nutrients, in many parts of the world, a heath or heathland is an environment characterised by an open dwarf-shrub community found on low-quality acidic soils, generally dominated by plants in the Ericaceae. A common example is Erica tetralix and this plant family is also typical of peat bogs and blanket bogs, examples include Rhododendron groenlandicum and Kalmia polifolia. In eastern North America, members of this family grow in association with an oak canopy. Some evidence suggests eutrophic rainwater can convert ericoid heaths with species such as Erica tetralix to grasslands, nitrogen is particularly suspect in this regard, and may be causing measurable changes to the distribution and abundance of some ericaceous species
Ericaceae
–
Ericaceae
8.
Arbutus
–
Arbutus is a genus of 11 accepted species of flowering plants in the family Ericaceae, native to warm temperate regions of the Mediterranean, western Europe, the Canary Islands and North America. The name is borrowed from Latin, where it referred to A. unedo, Arbutus are small trees or shrubs with red flaking bark and edible red berries. Fruit development is delayed for five months after pollination, so that flowers appear while the previous years fruit are ripening. Members of the genus are called madrones or madronas in the United States, in Canada—in British Columbia, where the species is common—arbutus is commonly used or, rarely and locally, tick tree. All refer to the species, Arbutus menziesii, native to the Pacific Northwest. It is Canadas only native broadleaved evergreen tree, some species in the genera Epigaea, Arctostaphylos and Gaultheria were formerly classified in Arbutus. As a result of its past classification, Epigaea repens has a common name of trailing arbutus. Arbutus andrachne L. – Greek Strawberry Tree Arbutus canariensis Duhamel – Canary Madrone Arbutus pavarii Pampan, Arbutus unedo L. – Strawberry Tree Arbutus arizonica Sarg. – Arizona Madrone Arbutus glandulosa Mart and it is also likely that either or both of A. canariensis and A. andrachne conferred not only structural size but leaf size and smoothness of bark. The parent species likely experienced different blooming times, as the hybrid blooms nearly continuously, Arbutus × andrachnoides Link Arctostaphylos tomentosa Lindl. Comarostaphylis discolor Diggs Gaultheria phillyreifolia Sleumer Arbutus species are used as food plants by some Lepidoptera species including emperor moth, several species are widely cultivated as ornamental plants outside of their natural ranges, though cultivation is often difficult due to their intolerance of root disturbance. The hybrid Arbutus Marina is much more adaptable and thrives under garden conditions, the Arbutus unedo tree makes up part of the coat of arms of the city of Madrid, Spain. In the center of the city there is a statue of a bear eating the fruit of the madroño tree, the image appears on city crests, taxi cabs, man-hole covers, and other city infrastructure. The tree also figured into certain myths of the Straits Salish, the fruit is edible but has minimal flavour and is not widely eaten. In Portugal, the fruit is distilled into a potent brandy known as medronho. In Madrid, the fruit is distilled into madroño, a sweet, Arbutus is a great fuelwood tree since it burns hot and long. Also, according to the Great Flood legends of several bands in the northwest, because of this the Saanich people do not burn madrone out of thanks for saving them. Vasey, Michael C. & Thomas Parker, V.2001, phylogeny and Biogeography of the Arbutoideae, Implications for the Madrean-Tethyan Hypothesis
Arbutus
–
Arbutus
Arbutus
–
The bear and the tree at
Puerta del Sol, Madrid
Arbutus
–
Arbutus menziesii —full tree
Arbutus
–
Arbutus menziesii —flowers
9.
Binomial nomenclature
–
Such a name is called a binomial name, a binomen, binominal name or a scientific name, more informally it is also called a Latin name. The first part of the name identifies the genus to which the species belongs, for example, humans belong to the genus Homo and within this genus to the species Homo sapiens. The formal introduction of system of naming species is credited to Carl Linnaeus. But Gaspard Bauhin, in as early as 1623, had introduced in his book Pinax theatri botanici many names of genera that were adopted by Linnaeus. Although the general principles underlying binomial nomenclature are common to these two codes, there are differences, both in the terminology they use and in their precise rules. Similarly, both parts are italicized when a binomial name occurs in normal text, thus the binomial name of the annual phlox is now written as Phlox drummondii. In scientific works, the authority for a name is usually given, at least when it is first mentioned. In zoology Patella vulgata Linnaeus,1758, the original name given by Linnaeus was Fringilla domestica, the parentheses indicate that the species is now considered to belong in a different genus. The ICZN does not require that the name of the person who changed the genus be given, nor the date on which the change was made, in botany Amaranthus retroflexus L. – L. is the standard abbreviation used in botany for Linnaeus. – Linnaeus first named this bluebell species Scilla italica, Rothmaler transferred it to the genus Hyacinthoides, the ICN does not require that the dates of either publication be specified. Prior to the adoption of the binomial system of naming species. Together they formed a system of polynomial nomenclature and these names had two separate functions. First, to designate or label the species, and second, to be a diagnosis or description, such polynomial names may sometimes look like binomials, but are significantly different. For example, Gerards herbal describes various kinds of spiderwort, The first is called Phalangium ramosum, Branched Spiderwort, is aptly termed Phalangium Ephemerum Virginianum, Soon-Fading Spiderwort of Virginia. The Latin phrases are short descriptions, rather than identifying labels, the Bauhins, in particular Caspar Bauhin, took some important steps towards the binomial system, by pruning the Latin descriptions, in many cases to two words. The adoption by biologists of a system of binomial nomenclature is due to Swedish botanist and physician Carl von Linné. It was in his 1753 Species Plantarum that he first began using a one-word trivial name together with a generic name in a system of binomial nomenclature. This trivial name is what is now known as an epithet or specific name
Binomial nomenclature
–
Carl Linnaeus (1707–1778), a Swedish botanist, invented the modern system of binomial nomenclature.
10.
Carl Linnaeus
–
Carl Linnaeus, also known after his ennoblement as Carl von Linné, was a Swedish botanist, physician, and zoologist, who formalised the modern system of naming organisms called binomial nomenclature. He is known by the father of modern taxonomy. Many of his writings were in Latin, and his name is rendered in Latin as Carolus Linnæus, Linnaeus was born in the countryside of Småland, in southern Sweden. He received most of his education at Uppsala University. He lived abroad between 1735 and 1738, where he studied and also published a first edition of his Systema Naturae in the Netherlands and he then returned to Sweden, where he became professor of medicine and botany at Uppsala. In the 1740s, he was sent on journeys through Sweden to find and classify plants. In the 1750s and 1760s, he continued to collect and classify animals, plants, and minerals, at the time of his death, he was one of the most acclaimed scientists in Europe. The philosopher Jean-Jacques Rousseau sent him the message, Tell him I know no man on earth. The German writer Johann Wolfgang von Goethe wrote, With the exception of Shakespeare and Spinoza, Swedish author August Strindberg wrote, Linnaeus was in reality a poet who happened to become a naturalist. Among other compliments, Linnaeus has been called Princeps botanicorum, The Pliny of the North and he is also considered as one of the founders of modern ecology. In botany, the abbreviation used to indicate Linnaeus as the authority for species names is L. In older publications, sometimes the abbreviation Linn. is found, Linnæus was born in the village of Råshult in Småland, Sweden, on 23 May 1707. He was the first child of Nicolaus Ingemarsson and Christina Brodersonia and his siblings were Anna Maria Linnæa, Sofia Juliana Linnæa, Samuel Linnæus, and Emerentia Linnæa. One of a line of peasants and priests, Nils was an amateur botanist, a Lutheran minister. Christina was the daughter of the rector of Stenbrohult, Samuel Brodersonius, a year after Linnæus birth, his grandfather Samuel Brodersonius died, and his father Nils became the rector of Stenbrohult. The family moved into the rectory from the curates house, even in his early years, Linnæus seemed to have a liking for plants, flowers in particular. Whenever he was upset, he was given a flower, which calmed him. Nils spent much time in his garden and often showed flowers to Linnaeus, soon Linnæus was given his own patch of earth where he could grow plants
Carl Linnaeus
–
Carl von Linné,
Alexander Roslin, 1775. Oil painting in the portrait collection at
Gripsholm Castle
Carl Linnaeus
–
The
coat of arms of Carl von Linné.
Carl Linnaeus
–
Birthplace of Linnæus at
Råshult
Carl Linnaeus
–
Statue of Linnaeus as a University student in
Lund
11.
Synonym (taxonomy)
–
For example, Linnaeus was the first to give a scientific name to the Norway spruce, which he called Pinus abies. This name is no longer in use, it is now a synonym of the current scientific name which is Picea abies, unlike synonyms in other contexts, in taxonomy a synonym is not interchangeable with the name of which it is a synonym. In taxonomy, synonyms are not equals, but have a different status, for any taxon with a particular circumscription, position, and rank, only one scientific name is considered to be the correct one at any given time. A synonym cannot exist in isolation, it is always an alternative to a different scientific name, given that the correct name of a taxon depends on the taxonomic viewpoint used a name that is one taxonomists synonym may be another taxonomists correct name. Synonyms may arise whenever the same taxon is described and named more than once, independently. They may also arise when existing taxa are changed, as when two taxa are joined to one, a species is moved to a different genus. To the general user of scientific names, in such as agriculture, horticulture, ecology, general science. A synonym is a name that was used as the correct scientific name but which has been displaced by another scientific name. Thus Oxford Dictionaries Online defines the term as a name which has the same application as another. In handbooks and general texts, it is useful to have mentioned as such after the current scientific name. Synonyms used in this way may not always meet the strict definitions of the synonym in the formal rules of nomenclature which govern scientific names. Changes of scientific name have two causes, they may be taxonomic or nomenclatural, a name change may be caused by changes in the circumscription, position or rank of a taxon, representing a change in taxonomic, scientific insight. A name change may be due to purely nomenclatural reasons, that is, based on the rules of nomenclature, the earliest such name is called the senior synonym, while the later name is the junior synonym. One basic principle of zoological nomenclature is that the earliest correctly published name, synonyms are important because if the earliest name cannot be used, then the next available junior synonym must be used for the taxon. Objective synonyms refer to taxa with the type and same rank. For example, John Edward Gray published the name Antilocapra anteflexa in 1855 for a species of pronghorn, however, it is now commonly accepted that his specimen was an unusual individual of the species Antilocapra americana published by George Ord in 1815. Ords name thus takes precedence, with Antilocapra anteflexa being a subjective synonym. Objective synonyms are common at the level of genera, because for various reasons two genera may contain the type species, these are objective synonyms
Synonym (taxonomy)
–
The
Latin Caudata and
Greek Urodela both mean "tailed" and have been used as a scientific name at the rank of order for the
salamanders (as opposed to the tail-less
frogs). Thus they are synonyms.
Synonym (taxonomy)
–
The common dandelion
Taraxacum officinale sensu lato is an extremely widespread group of
apomictic lineages, and some scientists apply the "biological species concept" to divide it into many distinct species; other scientists regard all the names for those independent species as synonyms.
12.
Johann Centurius Hoffmannsegg
–
Johann Centurius Hoffmann Graf von Hoffmannsegg was a German botanist, entomologist and ornithologist. Hoffmannsegg was born at Rammenau and studied at Leipzig and Göttingen and he travelled through Europe acquiring vast collections of plants and animals. He visited Hungary, Austria and Italy in 1795–1796 and Portugal from 1797 to 1801 and he sent his collections to Johann Karl Wilhelm Illiger, then in Brunswick, so that he could study them. Hoffmannsegg worked in Berlin from 1804 to 1816, and was elected a member of the Academy of Science of the city in 1815 and he was the founder of the zoological museum of Berlin in 1809. Hoffmannsegg proposed Illiger for the position of curator, and all the Hoffmannseggs collections were transferred to Berlin. The legume genus Hoffmannseggia is named for him, regarding personal names, Until 1919, Graf was a title, translated as Count, not a first or middle name. In Germany since 1919, it part of family names
Johann Centurius Hoffmannsegg
–
Johann Centurius Hoffmannsegg
13.
Evergreen
–
In botany, an evergreen is a plant that has leaves throughout the year, always green. This contrasts with deciduous plants, which lose their foliage during the winter or dry season. There are many different kinds of plants, both trees and shrubs. Deciduous trees shed their leaves, usually as an adaptation to a cold or dry/wet season, evergreen trees do lose leaves, but each tree loses its leaves gradually and not all at once. Most warm temperate climate plants are also evergreen, in cool temperate climates, fewer plants are evergreen, with a predominance of conifers, as few evergreen broadleaf plants can tolerate severe cold below about −30 °C. In areas where there is a reason for being deciduous, being evergreen is usually an adaptation to low nutrient levels, deciduous trees lose nutrients whenever they lose their leaves. In warmer areas, species such as pines and cypresses grow on poor soils. In Rhododendron, a genus with many broadleaf evergreens, several species grow in forests but are usually found on highly acidic soil where the nutrients are less available to plants. In taiga or boreal forests, it is too cold for the matter in the soil to decay rapidly, so the nutrients in the soil are less easily available to plants. These conditions favour the growth of more evergreens and make it difficult for deciduous plants to persist. In addition, the shelter provided by existing evergreen plants can make it easier for younger evergreen plants to survive cold and/or drought
Evergreen
–
A
Silver Fir shoot showing three successive years of retained leaves.
Evergreen
–
A
Southern Live Oak in winter.
14.
Shrub
–
A shrub or bush is a small to medium-sized woody plant. It is distinguished from a tree by its multiple stems and shorter height, plants of many species may grow either into shrubs or trees, depending on their growing conditions. Small, low shrubs, generally less than 2 m tall, such as lavender, periwinkle, an area of cultivated shrubs in a park or a garden is known as a shrubbery. When clipped as topiary, suitable species or varieties of shrubs develop dense foliage, many shrubs respond well to renewal pruning, in which hard cutting back to a stool results in long new stems known as canes. Other shrubs respond better to selective pruning to reveal their structure, shrubs in common garden practice are generally considered broad-leaved plants, though some smaller conifers such as mountain pine and common juniper are also shrubby in structure. Species that grow into a shrubby habit may be deciduous or evergreen
Shrub
–
A
broom shrub in flower
Shrub
–
A
rhododendron shrubbery in
Sheringham Park
15.
Tree
–
In botany, a tree is a perennial plant with an elongated stem, or trunk, supporting branches and leaves in most species. In some usages, the definition of a tree may be narrower, including only woody plants with secondary growth, plants that are usable as lumber or plants above a specified height. Trees are not a group but include a variety of plant species that have independently evolved a woody trunk. In looser senses, the palms, the tree ferns, bananas. Trees tend to be long-lived, some reaching several thousand years old, the tallest known tree, a coast redwood named Hyperion, stands 115.6 m high. Trees have been in existence for 370 million years and it is estimated that there are just over 3 trillion mature trees in the world. A tree typically has many secondary branches supported clear of the ground by the trunk and this trunk typically contains woody tissue for strength, and vascular tissue to carry materials from one part of the tree to another. For most trees it is surrounded by a layer of bark which serves as a protective barrier, below the ground, the roots branch and spread out widely, they serve to anchor the tree and extract moisture and nutrients from the soil. Above ground, the divide into smaller branches and shoots. The shoots typically bear leaves, which light energy and convert it into sugars by photosynthesis, providing the food for the trees growth. Flowers and fruit may also be present, but some trees, such as conifers, instead have pollen cones and seed cones, others, such as tree ferns, trees play a significant role in reducing erosion and moderating the climate. They remove carbon dioxide from the atmosphere and store large quantities of carbon in their tissues, trees and forests provide a habitat for many species of animals and plants. Tropical rainforests are one of the most biodiverse habitats in the world, trees provide shade and shelter, timber for construction, fuel for cooking and heating, and fruit for food as well as having many other uses. In parts of the world, forests are shrinking as trees are cleared to increase the amount of available for agriculture. Because of their longevity and usefulness, trees have always revered, with sacred groves in various cultures. Although tree is a term of common parlance, there is no universally recognised precise definition of what a tree is, either botanically or in common language. In its broadest sense, a tree is any plant with the form of an elongated stem, or trunk. Trees are also defined by height, with smaller plants from 0.5 to 10 m being called shrubs
Tree
–
Common ash (
Fraxinus excelsior), a
deciduous broad-leaved (
angiosperm) tree
Tree
–
European larch (
Larix decidua), a
coniferous tree which is also
deciduous
Tree
–
Lepidodendron, an extinct
lycophyte tree
Tree
–
Diagram of
secondary growth in a
eudicot or
coniferous tree showing idealised vertical and horizontal sections. A new layer of
wood is added in each growing season, thickening the stem, existing branches and roots.
16.
Family (biology)
–
In biological classification, family is one of the eight major taxonomic ranks, it is classified between order and genus. A family may be divided into subfamilies, which are intermediate ranks above the rank of genus. In vernacular usage, a family may be named one of its common members, for example, walnuts and hickory trees belong to the family Juglandaceae. What does or does not belong to a family—or whether a family should be recognized at all—are proposed and determined by practicing taxonomists. There are no rules for describing or recognizing a family. Taxonomists often take different positions about descriptions of taxa, and there may be no broad consensus across the community for some time. Some described taxa are accepted broadly and quickly, but others only rarely, if at all, the naming of families is codified by various international codes. In zoological nomenclature, the names of animals end with the suffix -idae. The concept of rank at time was not yet settled, and in the preface to the Prodromus Magnol spoke of uniting his families into larger genera. Carolus Linnaeus used the word familia in his Philosophia botanica to denote groups of plants, trees, herbs, ferns, palms. He used this term only in the section of the book. In zoology, the family as an intermediate between order and genus was introduced by Pierre André Latreille in his Précis des caractères génériques des insectes. He used families in some but not in all his orders of insects, families can be used for evolutionary, palaeontological and generic studies because they are more stable than lower taxonomic levels such as genera and species
Family (biology)
–
The hierarchy of
biological classification 's eight major
taxonomic ranks. An
order contains one or more families. Intermediate minor rankings are not shown.
17.
Mediterranean region
–
As a rule of thumb, the Mediterranean Basin is the Old World region where olive trees grow. The Mediterranean basin covers portions of three continents Africa, Asia, and Europe and it has a varied and contrasting topography. Contrary to the sandy beach images portrayed in most tourist brochures. Mountains can be seen from almost anywhere, the Mediterranean Basin extends into Western Asia, covering the western and southern portions of the peninsula of Turkey, excluding the temperate-climate mountains of central Turkey. It includes the Mediterranean climate Levant at the end of the Mediterranean, bounded on the east and south by the Syrian. The northern portion of the Maghreb region of northwestern Africa has a Mediterranean climate, separated from the Sahara Desert, Europe lies to the north, and three large Southern European peninsulas, the Iberian Peninsula, Italian Peninsula, and the Balkan Peninsula, extend into the Mediterranean-climate zone. The Mediterranean Basin was shaped by the ancient collision of the northward-moving African-Arabian continent with the stable Eurasian continent, as Africa-Arabia moved north, it closed the former Tethys Sea, which formerly separated Eurasia from the ancient super continent of Gondwana, of which Africa was part. At about the time,170 mya in the Jurassic period. The collision pushed up a vast system of mountains, extending from the Pyrenees in Spain to the Zagros Mountains in Iran and this episode of mountain building, known as the Alpine orogeny, occurred mostly during the Oligocene and Miocene epochs. The Neotethys became larger during these collisions and associated folding and subduction, about 6 mya during the late Miocene, the Mediterranean was closed at its western end by drifting Africa, which caused the entire sea to evaporate. There followed several episodes of sea drawdown and re-flooding known as the Messinian Salinity Crisis, recent studies, however, show that repeated desiccation and re-flooding is unlikely from a geodynamic point of view. The end of the Miocene also marked a change in the Mediterranean Basins climate, fossil evidence shows that the Mediterranean Basin had a relatively humid subtropical climate with summer rainfall during the Miocene, which supported laurel forests. The shift to a Mediterranean climate occurred within the last 3. 2–2.8 million years, during the Pliocene epoch, much of these forests and shrublands have been altered beyond recognition by thousands of years of human habitation. There are now very few relatively intact natural areas in what was once a wooded region. The Mediterranean Region was first proposed by German botanist August Grisebach in the late 19th century, drosophyllaceae, recently segregated from Droseraceae, is the only plant family endemic to the region. Among the endemic plant genera are, The genera Aubrieta, Sesamoides, Cynara, Dracunculus, moreover, many plant taxa are shared with one of the four neighboring floristic regions only. The Mediterranean Basin is the largest of the worlds five Mediterranean forests, woodlands and it is home to a number of plant communities, which vary with rainfall, elevation, latitude, and soils. Scrublands occur in the driest areas, especially areas near the seacoast where wind, low, soft-leaved scrublands around the Mediterranean are known as garrigar in Catalan, garrigue in French, phrygana in Greek, tomillares in Spanish, and batha in Hebrew
Mediterranean region
–
Potential distribution over the Mediterranean Basin of the olive tree—one of the best biological indicators of the Mediterranean Region (Oteros, 2014)
Mediterranean region
–
Political Map of the Mediterranean Basin
18.
Killarney
–
Killarney is a town in County Kerry, southwestern Ireland. Its natural heritage, history and location on the Ring of Kerry make Killarney a popular tourist destination, Killarney won the Best Kept Town award in 2007, in a cross-border competition jointly organised by the Department of the Environment and the Northern Ireland Amenity Council. In 2011, it was named Irelands tidiest town and the cleanest town in the country by Irish Business Against Litter, Killarney has featured prominently in early Irish history, with religious settlements playing an important part of its recorded history. Its first significantly historical settlement was the monastery on nearby Innisfallen Island founded in 640 by St. Finian the Leper, Innisfallen or Inishfallen is an island in Lough Leane, one of the three Lakes of Killarney in County Kerry, Ireland. It is home to the ruins of Innisfallen Abbey, one of the most impressive archaeological remains dating from the early Christian period found in the Killarney National Park, the monastery was founded in 640 by St. Finian the Leper, and was occupied for approximately 850 years. Over a period of about 300 of these, the monks wrote the Annals of Innisfallen, the monks were dispossessed of the abbey on 18 August 1594, by Elizabeth I. The location of the monastery on the island is thought to have given rise to the name Lough Leane, according to tradition the Irish High King Brian Boru received his education at Innisfallen under Maelsuthain OCarroll. Maelsuthain has been credited as the originator of the Annals. It is possible for tourists to visit the island during the summer months, Aghadoe, the local townland which overlooks present day Killarney, may have begun as a pagan religious site. The site has also associated with the 5th century missionary St. Abban. According to legend, St. Finian founded a monastery at Aghadoe in the 6th or 7th century, the first written record of a monastery dates from 939 AD in the Annals of Innisfallen where the Aghadoe monastery is referred to as the Old Abbey. Following the Anglo-Norman invasion of Ireland in 1169, the Normans built Parkavonear Castle, the castle was perhaps intended as an early warning outpost due to its views of the entire Killarney valley and lakes region. Ross Castle was built on the shore in the late 15th century by local ruling clan the ODonoghues Mor. Ownership of the castle changed hands during the Desmond Rebellions of the 1580s to the Mac Carty Mor, Muckross Abbey was founded in 1448 as a Franciscan friary for the Observantine Franciscans by Donal McCarthy Mor. The abbey was burned down by Cromwellian forces under General Ludlow in 1654, Killarney was heavily involved in the Irish War of Independence. The town, and indeed the county, had strong republican ties. The Great Southern Hotel, was for a while taken over by the British, One notable event during the war was the Headford Ambush when the IRA attacked a railway train a few miles from town. A day after the Ballyseedy Massacre, five Republican prisoners were murdered in Killarney in retaliation, killarneys tourism history goes back at least to the mid 18th century, when Thomas, fourth Viscount Kenmare, began to attract visitors and new residents to the town
Killarney
–
St Mary's Cathedral
Killarney
–
Plaque commemorating the coming of the Railway to Killarney
Killarney
–
Tourists near Ross Castle.
Killarney
–
Ross Castle
19.
Species Plantarum
–
Species Plantarum is a book by Carl Linnaeus, originally published in 1753, which lists every species of plant known at the time, classified into genera. It is the first work to consistently apply binomial names and was the point for the naming of plants. Species Plantarum was published on 1 May 1753 by Laurentius Salvius in Stockholm, a second edition was published in 1762–1763, and a third edition in 1764, although this scarcely differed from the second. Species Plantarum was the first botanical work to apply the binomial nomenclature system of naming to any large group of organisms. The use of names had originally been developed as a kind of shorthand in a student project about the plants eaten by cattle. After the specific epithet, Linnaeus gave a description of each species. Because it is the first work in which binomial nomenclature was consistently applied, Species Plantarum contained descriptions of the thousands of plant species known to Linnaeus at the time. In the first edition, there were 5,940 names, in his introduction, Linnaeus estimated that there were fewer than 10,000 plant species in existence, there are now thought to be around 400,000 species of flowering plants alone. The species were arranged in around a thousand genera, which were grouped into 24 classes, 1–2 at Biodiversity Heritage Library facsimile Species Plantarum, vol. 1, vol.2 at Botanicus Species Plantarum at Project Guttenberg I-III IV-V VI-X XI-XIII text Linnaeus Link Union Catalogue
Species Plantarum
–
Cover page of first edition
Species Plantarum
–
Prior to Species Plantarum, this plant was referred to as " Plantago foliis ovato-lanceolatis pubescentibus, spica cylindrica, scapo tereti "; Linnaeus renamed it
Plantago media.
20.
Ribosomal DNA
–
Ribosomal DNA is a DNA sequence that codes for ribosomal RNA. Ribosomes are assemblies of proteins and rRNA molecules that translate mRNA molecules to produce proteins. As shown in the figure, rDNA of eukaryotes consists of a repeat of a unit segment, an operon, composed of NTS, ETS, 18S, ITS1,5. 8S, ITS2. RDNA has another gene, coding for 5S rRNA, located in the genome in most eukaryotes, 5S rDNA is also present in tandem repeats as in Drosophila. In the nucleus, the region of the chromosome is visualized as a nucleolus which forms expanded chromosomal loops with rDNA. These rDNA regions are also called nucleolus organizer regions, as they rise to the nucleolus. In the human genome there are 5 chromosomes with nucleolus organizer regions, in Bacteria, Archaea and chloroplasts the rRNA is composed of different units, the large ribosomal RNA, 16S ribosomal RNA and 5S rRNA. The 16S rRNA is widely used for phylogenetic studies, in the large rDNA array, polymorphisms between rDNA repeat units are very low, indicating that rDNA tandem arrays are evolving through concerted evolution. They could occur by slippage of the newly synthesized strand during DNA replication or by gene conversion, the rDNA transcription tracts have low rate of polymorphism among species, which allows interspecific comparison to elucidate phylogenetic relationship using only a few specimens. Coding regions of rDNA are highly conserved among species but ITS regions are due to insertions, deletions. Between remote species as human and frog comparison of sequences at ITS tracts is not appropriate, conserved sequences at coding regions of rDNA allow comparisons of remote species, even between yeast and human. Human 5. 8S rRNA has 75% identity with yeast 5. 8S rRNA, in cases for sibling species, comparison of the rDNA segment including ITS tracts among species and phylogenetic analysis are made satisfactorily. The different coding regions of the rDNA repeats usually show distinct evolutionary rates, as a result, this DNA can provide phylogenetic information of species belonging to wide systematic levels
Ribosomal DNA
–
The gene segment of eukaryotic rDNA contains 18S, 5.8S, and 28S tracts and forms a tandem repetitive cluster; the 5S rDNA is coded separately. NTS, nontranscribed spacer, ETS, external transcribed spacer, ITS,
internal transcribed spacers 1 and 2, numbered from 5' end.
21.
Paraphyletic
–
In taxonomy, a group is paraphyletic if it consists of the groups last common ancestor and all descendants of that ancestor excluding a few—typically only one or two—monophyletic subgroups. The group is said to be paraphyletic with respect to the excluded subgroups, the term is commonly used in phylogenetics and in linguistics. The term was coined to apply to well-known taxa like reptiles which, as commonly named and traditionally defined, is paraphyletic with respect to mammals and birds. Reptilia contains the last common ancestor of reptiles and all descendants of that all extant reptiles as well as the extinct synapsids—except for mammals. Other commonly recognized paraphyletic groups include fish, monkeys and lizards, if many subgroups are missing from the named group, it is said to be polyparaphyletic. A paraphyletic group cannot be a clade, which is a monophyletic group, groups that include all the descendants of a common ancestor are said to be monophyletic. A paraphyletic group is a group from which one or more subsidiary clades is excluded to form a separate group. Ereshefsky has argued that paraphyletic taxa are the result of anagenesis in the group or groups. For example, dinosaurs are paraphyletic with respect to birds because birds possess many features that dinosaurs lack, a group whose identifying features evolved convergently in two or more lineages is polyphyletic. More broadly, any taxon that is not paraphyletic or monophyletic can be called polyphyletic and these terms were developed during the debates of the 1960s and 70s accompanying the rise of cladistics. The prokaryotes, because they exclude the eukaryotes, a descendant group, bacteria and Archaea are prokaryotes, but archaea and eukaryotes share a common ancestor that is not ancestral to the bacteria. The prokaryote/eukaryote distinction was proposed by Edouard Chatton in 1937 and was accepted after being adopted by Roger Stanier. The botanical code abandoned consideration of bacterial nomenclature in 1975, currently, dicotyledons are paraphyletic because the group excludes monocotyledons. Dicotyledon has not been used as an ICBN classification for decades, phylogenetic analysis indicates that the monocots are a development from a dicot ancestor. Excluding monocots from the dicots makes the latter a paraphyletic group, the order Artiodactyla, because it excludes Cetaceans. In the ICZN Code, the two taxa are orders of equal rank, molecular studies, however, have shown that the Cetacea descend from the Artiodactyl ancestors, although the precise phylogeny within the order remains uncertain. Without the Cetacean descendants the Artiodactyls must be paraphyletic, the class Reptilia as traditionally defined, because it excludes birds and mammals. In the ICZN Code, the three taxa are classes of equal rank, however, mammals hail from the synapsids and birds are descended from the dinosaurs, both of which are reptiles
Paraphyletic
–
The traditional class
Reptilia (green field) is a paraphyletic group comprising all
amniotes (Amniota) other than the mammals and birds. In contrast, the
clade Amniota itself is
monophyletic.
22.
Mediterranean Basin
–
As a rule of thumb, the Mediterranean Basin is the Old World region where olive trees grow. The Mediterranean basin covers portions of three continents Africa, Asia, and Europe and it has a varied and contrasting topography. Contrary to the sandy beach images portrayed in most tourist brochures. Mountains can be seen from almost anywhere, the Mediterranean Basin extends into Western Asia, covering the western and southern portions of the peninsula of Turkey, excluding the temperate-climate mountains of central Turkey. It includes the Mediterranean climate Levant at the end of the Mediterranean, bounded on the east and south by the Syrian. The northern portion of the Maghreb region of northwestern Africa has a Mediterranean climate, separated from the Sahara Desert, Europe lies to the north, and three large Southern European peninsulas, the Iberian Peninsula, Italian Peninsula, and the Balkan Peninsula, extend into the Mediterranean-climate zone. The Mediterranean Basin was shaped by the ancient collision of the northward-moving African-Arabian continent with the stable Eurasian continent, as Africa-Arabia moved north, it closed the former Tethys Sea, which formerly separated Eurasia from the ancient super continent of Gondwana, of which Africa was part. At about the time,170 mya in the Jurassic period. The collision pushed up a vast system of mountains, extending from the Pyrenees in Spain to the Zagros Mountains in Iran and this episode of mountain building, known as the Alpine orogeny, occurred mostly during the Oligocene and Miocene epochs. The Neotethys became larger during these collisions and associated folding and subduction, about 6 mya during the late Miocene, the Mediterranean was closed at its western end by drifting Africa, which caused the entire sea to evaporate. There followed several episodes of sea drawdown and re-flooding known as the Messinian Salinity Crisis, recent studies, however, show that repeated desiccation and re-flooding is unlikely from a geodynamic point of view. The end of the Miocene also marked a change in the Mediterranean Basins climate, fossil evidence shows that the Mediterranean Basin had a relatively humid subtropical climate with summer rainfall during the Miocene, which supported laurel forests. The shift to a Mediterranean climate occurred within the last 3. 2–2.8 million years, during the Pliocene epoch, much of these forests and shrublands have been altered beyond recognition by thousands of years of human habitation. There are now very few relatively intact natural areas in what was once a wooded region. The Mediterranean Region was first proposed by German botanist August Grisebach in the late 19th century, drosophyllaceae, recently segregated from Droseraceae, is the only plant family endemic to the region. Among the endemic plant genera are, The genera Aubrieta, Sesamoides, Cynara, Dracunculus, moreover, many plant taxa are shared with one of the four neighboring floristic regions only. The Mediterranean Basin is the largest of the worlds five Mediterranean forests, woodlands and it is home to a number of plant communities, which vary with rainfall, elevation, latitude, and soils. Scrublands occur in the driest areas, especially areas near the seacoast where wind, low, soft-leaved scrublands around the Mediterranean are known as garrigar in Catalan, garrigue in French, phrygana in Greek, tomillares in Spanish, and batha in Hebrew
Mediterranean Basin
–
Potential distribution over the Mediterranean Basin of the olive tree—one of the best biological indicators of the Mediterranean Region (Oteros, 2014)
Mediterranean Basin
–
Political Map of the Mediterranean Basin
23.
Arbutus andrachne
–
Arbutus andrachne, commonly called the Greek strawberry tree, is an evergreen shrub or small tree in the family Ericaceae, native to the Mediterranean region, the Middle East, and southwestern Asia. Arbutus andrachne can reach a height of about 12 meters, the smooth bark is exfoliating during the summer, leaving a layer with a pistachio green color, which changes gradually to a beautiful orange brown. The flowers bloom in Spring and are white or yellowish green, arbutus andrachne was reported by Peter Collinson as having flowered first in England in Dr John Fothergills extensive botanical garden and greenhouses at Upton House, Essex, in 1765. There was on the road a small tree bearing a fruit somewhat bigger than our largest cherries, and of the shape and taste of strawberries and it is pleasant to eat, but, if a great quantity be eaten, it mounts to the head, and intoxicates. It is ripe in November and December, editors footnote, From the description, it seems to be the arbutus Andrachne
Arbutus andrachne
–
Greek Strawberry Tree
Arbutus andrachne
–
Plant of Arbutus andrachne
Arbutus andrachne
–
Flowers and leaves
Arbutus andrachne
–
Fruits
24.
Hybrid (biology)
–
In biology, a hybrid, also known as a cross breed, is the result of combining, through sexual reproduction, the qualities of two animals or plants of different breeds, varieties, species or genera. Hybrids are not always intermediate between their parents, but can show heterosis or hybrid vigour, often growing larger or taller than either parent, some act before fertilization, others after it. A few animal species and many plant species, however, are the result of hybrid speciation, doubling the number of chromosomes to create polyploids is important in hybrid speciation, because homoploid hybrids are rarely fertile, the polyploid hybrids are allopolyploids. Human impact on the environment has greatly increased the mixing of species, with introduced species worldwide, the genetic mixing may threaten many species with extinction, while genetic erosion in crop plants may be damaging the gene pools of many species for future breeding. Many commercially useful fruits, flowers, garden herbs and trees have been produced by hybridization, one flower, Oenothera lamarckiana, was central to early genetics research into polyploidy. The term hybrid is derived from Latin hybrida, used for such as of a tame sow and a wild boar, or the child of a freeman. The term came into use in English in the 19th century. Conspicuous hybrids are named with portmanteau words, starting in the 1920s with the breeding of tiger-lion hybrids. The cross between two different homozygous lines produces an F1 hybrid that is heterozygous, having two alleles, one contributed by each parent and typically one is dominant and the other recessive. Typically, the F1 generation is also phenotypically homogeneous, producing offspring that are all similar to each other, double cross hybrids result from the cross between two different F1 hybrids. Three-way cross hybrids result from the cross between an F1 hybrid and an inbred line, triple cross hybrids result from the crossing of two different three-way cross hybrids. Top cross hybrids result from the crossing of a top quality or pure-bred male, population hybrids result from the crossing of plants or animals in a population with those of another population. These include interspecific hybrids or crosses between different breeds. e, from the point of view of genetics, different kinds of hybrid can be distinguished as follows, A genetic hybrid carries two different alleles of the same gene. A structural hybrid results from the fusion of gametes that have differing structure in at least one chromosome, a numerical hybrid results from the fusion of gametes having different haploid numbers of chromosomes. A permanent hybrid results when only the heterozygous genotype occurs, as in Oenothera lamarckiana, from the point of view of taxonomy, hybrids differ according to their parentage, Hybrids between different subspecies are called intra-specific hybrids. Offspring resulting from interspecies mating, are called interspecific hybrids, these result in hybrid speciation. Intergeneric hybrids result from matings between different genera, such as sheep and goats. Interfamilial hybrids such as chickens and guineafowl or pheasants are reliably described but extremely rare
Hybrid (biology)
–
Hercules, a "
liger ", a lion/tiger hybrid.
Hybrid (biology)
–
A "
zonkey ", a zebra/donkey hybrid.
Hybrid (biology)
–
A "
jaglion ", a jaguar/lion hybrid.
Hybrid (biology)
–
A
mule, a domestic canary/goldfinch hybrid.
25.
Bayda, Libya
–
Bayda - Elbeida, also spelled Az Zāwiyah al Bayḑā’, Zāwiyat al Bayḑā’, Beida El Beida and known as Beda Littoria under Italian colonial rule, is a commercial and industrial city in eastern Libya. It is located in the northern Cyrenaica, with a population 250,000 people, Bayda is the fourth-largest city in Libya. It is the city of the Jabal al Akhdar district. Baydas history stretches back to Ancient Greece, when it was known as Balagrae, the 2, 000-year-old ruins of the ancient Greek colony Cyrene are located nearby in Shahad. One of the greatest attractions in the city is the tomb of a companion of the Islamic prophet Muhammad. For that reason, the city was known as Sidi Rafaa after him, after the arrival of Sayyid Muhammad ibn Ali as-Senussi in the area in the 19th century, and the construction of a zawiya, the city was renamed Az Zawiya Al Bayda. The modern city was built in the 1950s and it was originally intended to be the new capital of Libya and most of the necessary government buildings were constructed there. Eventually, the plan to move the capital from Tripoli to Bayda was dropped, Bayda is the administrative seat for Jabal al Akhdar district. The town became known as az-Zawiya al-Bayḍāʾ because of the white painted zawiya, within time, the word zawiya was dropped and the city became known simply as Bayda. When the settlement was first founded by the Greeks, it was known as Balagrae, during Italian rule, the city was known as Beda Littoria. The zawiya still exists, but it is in a neglected state, the city is famous for the valleys and forests nearby, which are not found in the other cities of the Green Mountains—Akhdar Mountains. It is named the Green Mountains because they are covered by forests and woodlands, of which the best known are the Hamri, Alpellnj. The highest point in the Akhdar Mountains is around 850 metres, the Akhdar mountain range lies within the Mediterranean dry woodlands and steppe ecoregion. The native flora has many closely related to those found in similar Greek, Italian. There are about 1,800 species of Mediterranean forests, woodlands, and scrub trees, plants, the forests are characterized by many oak species, junipers, pines, carobs, Albarbc, cypress, hawthorn, Chammari, and olive trees. There is a network of springs in most parts of the mountains. The forested areas and springs provide habitat supportive of a diversity of birds, beaches stretch along the chain for about 350 kilometres, in a diverse terrain with many coastal bays and inlets. Bayda is famous for the locale of Susa, a seaside resort on the Mediterranean Sea
Bayda, Libya
–
Clockwise from top:View of traditional town area in night, Headquarter in General Directorate of National Commercial Bank of Libya, Al-Ooba Street, Former Libyan Parliament Hall, Alsells Tower
Bayda, Libya
–
The ruins of the ancient Greek city of Balagrae, in Bayda.
Bayda, Libya
–
A snowy road on the outskirts of the city
26.
Leaf
–
A leaf is an organ of a vascular plant and is the principal lateral appendage of the stem. The leaves and stem together form the shoot, Leaves are collectively referred to as foliage, as in autumn foliage. Although leaves can be seen in different shapes, sizes and textures, typically a leaf is a thin, dorsiventrally flattened organ, borne above ground. Most leaves have distinctive upper surface and lower surface that differ in colour, hairiness, broad, flat leaves with complex venation are known as megaphylls and the species that bear them, the majority, as broad-leaved or megaphyllous plants. In others, such as the clubmosses, with different evolutionary origins, some leaves, such as bulb scales are not above ground, and in many aquatic species the leaves are submerged in water. Succulent plants often have thick juicy leaves, but some leaves are without major photosynthetic function and may be dead at maturity, as in some cataphylls, furthermore, several kinds of leaf-like structures found in vascular plants are not totally homologous with them. Examples include flattened plant stems called phylloclades and cladodes, and flattened leaf stems called phyllodes which differ from both in their structure and origin. Many structures of plants, such as the phyllids of mosses and liverworts and even of some foliose lichens. Leaves are the most important organs of most vascular plants and these are then further processed by chemical synthesis into more complex organic molecules such as cellulose, the basic structural material in plant cell walls. The plant must therefore bring these three together in the leaf for photosynthesis to take place. Once sugar has been synthesized, it needs to be transported to areas of growth such as the plant shoots and roots. Vascular plants transport sucrose in a tissue called the phloem. The phloem and xylem are parallel to each other but the transport of materials is usually in opposite directions. Within the leaf these vascular systems branch to form veins which supply as much as the leaf as possible and they are arranged on the plant so as to expose their surfaces to light as efficiently as possible without shading each other, but there are many exceptions and complications. For instance plants adapted to windy conditions may have pendent leaves, such as in many willows, the flat, or laminar, shape also maximises thermal contact with the surrounding air, promoting cooling. Functionally, in addition to photosynthesis the leaf is the site of transpiration and guttation. Many gymnosperms have thin needle-like or scale-like leaves that can be advantageous in cold climates with frequent snow and these are interpreted as reduced from megaphyllous leaves of their Devonian ancestors. For xerophytes the major constraint is not light flux or intensity, some window plants such as Fenestraria species and some Haworthia species such as Haworthia tesselata and Haworthia truncata are examples of xerophytes. and Bulbine mesembryanthemoides
Leaf
–
The leaves of a
beech tree
Leaf
–
Near the ground these
Eucalyptus saplings have juvenile dorsiventral foliage from the previous year, but this season their newly sprouting foliage is isobilateral, like the mature foliage on the adult trees above
Leaf
–
A leaf shed in
autumn.
Leaf
–
Cross-section of a leaf
27.
Flower
–
A flower, sometimes known as a bloom or blossom, is the reproductive structure found in plants that are floral. The biological function of a flower is to effect reproduction, usually by providing a mechanism for the union of sperm with eggs, Flowers may facilitate outcrossing or allow selfing. Some flowers produce diaspores without fertilization, Flowers contain sporangia and are the site where gametophytes develop. Many flowers have evolved to be attractive to animals, so as to them to be vectors for the transfer of pollen. After fertilization, the ovary of the flower develops into fruit containing seeds, the essential parts of a flower can be considered in two parts, the vegetative part, consisting of petals and associated structures in the perianth, and the reproductive or sexual parts. A stereotypical flower consists of four kinds of structures attached to the tip of a short stalk, each of these kinds of parts is arranged in a whorl on the receptacle. The four main whorls are as follows, Collectively the calyx, corolla, the next whorl toward the apex, composed of units called petals, which are typically thin, soft and colored to attract animals that help the process of pollination. Androecium, the whorl, consisting of units called stamens. Stamens consist of two parts, a called a filament, topped by an anther where pollen is produced by meiosis. Gynoecium, the innermost whorl of a flower, consisting of one or more units called carpels, the carpel or multiple fused carpels form a hollow structure called an ovary, which produces ovules internally. Ovules are megasporangia and they in turn produce megaspores by meiosis which develop into female gametophytes and these give rise to egg cells. The gynoecium of a flower is described using an alternative terminology wherein the structure one sees in the innermost whorl is called a pistil. A pistil may consist of a carpel or a number of carpels fused together. The sticky tip of the pistil, the stigma, is the receptor of pollen, the supportive stalk, the style, becomes the pathway for pollen tubes to grow from pollen grains adhering to the stigma. The relationship to the gynoecium on the receptacle is described as hypogynous, perigynous, although the arrangement described above is considered typical, plant species show a wide variation in floral structure. These modifications have significance in the evolution of flowering plants and are used extensively by botanists to establish relationships among plant species, the four main parts of a flower are generally defined by their positions on the receptacle and not by their function. Many flowers lack some parts or parts may be modified into other functions and/or look like what is typically another part, in some families, like Ranunculaceae, the petals are greatly reduced and in many species the sepals are colorful and petal-like. Other flowers have modified stamens that are petal-like, the flowers of Peonies and Roses are mostly petaloid stamens
Flower
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A poster with flowers or clusters of flowers produced by twelve species of flowering plants from different families
Flower
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Main parts of a mature flower (
Ranunculus glaberrimus)
Flower
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Reproductive parts of Easter Lily (Lilium longiflorum). 1. Stigma, 2. Style, 3. Stamens, 4. Filament, 5. Petal
Flower
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Left: A normal zygomorphic Streptocarpus flower. Right: An aberrant peloric Streptocarpus flower. Both of these flowers appeared on the Streptocarpus hybrid 'Anderson's Crows' Wings'.
28.
Bees
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Bees are flying insects closely related to wasps and ants, known for their role in pollination and, in the case of the best-known bee species, the European honey bee, for producing honey and beeswax. Bees are a lineage within the superfamily Apoidea, presently considered as a clade Anthophila. There are nearly 20,000 known species of bees in seven to nine recognized families, though many are undescribed and they are found on every continent except Antarctica, in every habitat on the planet that contains insect-pollinated flowering plants. Some species including honey bees, bumblebees, and stingless bees live socially in colonies, Bees are adapted for feeding on nectar and pollen, the former primarily as an energy source and the latter primarily for protein and other nutrients. Most pollen is used as food for larvae, Bee pollination is important both ecologically and commercially, the decline in wild bees has increased the value of pollination by commercially managed hives of honey bees. The most common bees in the Northern Hemisphere are the Halictidae, or sweat bees, vertebrate predators of bees include birds such as bee-eaters, insect predators include beewolves and dragonflies. Human beekeeping or apiculture has been practised for millennia, since at least the times of Ancient Egypt, apart from honey and pollination, honey bees produce beeswax, royal jelly and propolis. Bees have appeared in mythology and folklore, again since ancient times, and they feature in works of literature as varied as Virgils Georgics, Beatrix Potters The Tale of Mrs Tittlemouse, yeatss poem The Lake Isle of Innisfree. Bee larvae are included in the Javanese dish botok tawon, where they are steamed with shredded coconut. The ancestors of bees were wasps in the family Crabronidae, which were predators of other insects. The switch from insect prey to pollen may have resulted from the consumption of insects which were flower visitors and were partially covered with pollen when they were fed to the wasp larvae. This same evolutionary scenario may have occurred within the vespoid wasps, until recently, the oldest non-compression bee fossil had been found in New Jersey amber, Cretotrigona prisca of Cretaceous age, a corbiculate bee. A bee fossil from the early Cretaceous, Melittosphex burmensis, is considered an extinct lineage of pollen-collecting Apoidea sister to the modern bees. Derived features of its place it clearly within the bees. By the Eocene there was considerable diversity among eusocial bee lineages. The highly eusocial corbiculate Apidae appeared roughly 87 Mya, and the Allodapini around 53 Mya, the Colletidae appear as fossils only from the late Oligocene to early Miocene. The Melittidae are known from Palaeomacropis eocenicus in the Early Eocene, the Megachilidae are known from trace fossils from the Middle Eocene. The Andrenidae are known from the Eocene-Oligocene boundary, around 34 Mya, the Halictidae first appear in the Early Eocene with species found in amber
Bees
Bees
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Melittosphex burmensis, a fossil bee preserved in
amber from the
Early Cretaceous of
Myanmar
Bees
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Long-tongued bees and long-tubed flowers
coevolved, like this
Amegilla cingulata on
Acanthus ilicifolius.
Bees
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The lapping
mouthparts of a honeybee, showing labium and maxillae
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