John Lindley FRS was an English botanist and orchidologist. Born in Catton, near Norwich, John Lindley was one of four children of George and Mary Lindley. George Lindley ran a commercial nursery garden. Although he had great horticultural knowledge, the undertaking was not profitable and George lived in a state of indebtedness; as a boy he would assist in the garden and collected wild flowers he found growing in the Norfolk countryside. Lindley was educated at Norwich School, he would have liked to go to university or to buy a commission in the army but the family could not afford either. He became Belgian agent for a London seed merchant in 1815. At this time Lindley became acquainted with the botanist William Jackson Hooker who allowed him to use his botanical library and who introduced him to Sir Joseph Banks who offered him employment as an assistant in his herbarium, his first publication, in 1819, a translation of the Analyse du fruit of L. C. M. Richard, was followed in 1820 by an original Monographia Rosarum, with descriptions of new species, drawings executed by himself in 1821 by Monographia Digitalium, "Observations on Pomaceae", which were both contributed to the Linnean Society.
Lindley went to work at Banks’ house in London. He concentrated on the genera “Rosa” and “Digitalis” and published the monograph “A Botanical History of Roses” which distinguished seventy-six species, describes thirteen new ones and was illustrated by nineteen coloured plates painted by himself, he became acquainted with Joseph Sabine who grew a large assortment of roses and was the Secretary of the Horticultural Society of London. His employment came to an abrupt end with the death of Banks a few months later. One of Banks’ friends, a wealthy merchant called William Cattley, paid Lindley to draw and describe new plants in his garden at Barnet, he paid for the publication of “Digitalia Monographia”. In 1820, at the age of twenty-one, Lindley was elected a fellow of the Linnean Society of London. From 1821 to 1826 he published a folio work with coloured illustrations that he had painted himself, “Collectanea botanica or Figures and botanic Illustrations of rare and curious exotic Plants”. Many of these plants came from the family Orchidaceae.
Lindley was appointed assistant secretary to the Royal Horticultural Society and its new garden at Chiswick in 1822, where he supervised the collection of plants. Assistant secretary to the Horticultural Society since 1822, in 1829 Lindley was appointed to the chair of botany at University College, which he retained until 1860, he lectured on botany from 1831 at the Royal Institution, including delivering the 1833 Royal Institution Christmas Lecture, from 1836 at the Chelsea Physic Garden, starting the society's flower show in the late 1830s. Lindley described the plants collected on Thomas Livingstone Mitchell's expeditions of 1838 and wrote an Appendix to Edwards's Botanical Register of 1839, describing plants collected by James Drummond and Georgiana Molloy of the Swan River Colony in Western Australia. According to John Ryan, Lindley’s 1840 ‘Sketch of the Vegetation of the Swan River Colony’ provided ‘the most succinct portrait to date of the flora of the Swan River Settlement’, established in 1829.
The Sketch, published during November 1839 and January 1840 in Edwards’ Botanical Register and separately on its completion, was illustrated by nine hand-coloured lithographs and four wood-cuts. He played a large part in having Charles Moore appointed as Director of the Sydney Botanical Gardens. During his professorship, he wrote many scientific and popular works as well as making significant contributions to the Botanical Register, of which he was the editor for many years, to The Gardeners' Chronicle, where he was in charge of the horticultural department from 1841, he was a fellow of the Royal and Geological Societies. He received the Royal Society's royal medal in 1857, in 1853 became a corresponding member of the Institut de France. About this time, the Horticultural Society of London, which became the Royal Horticultural Society at a date, asked Lindley to draw roses and in 1822 he became the Assistant Secretary of the Society’s garden; the Society’s historian, Harold R Fletcher described him as “ … the backbone of the Society and the greatest servant it had had.”
Now with a steady income, in 1823 he married Sarah Freestone. They rented a house in rural Acton Green, a location convenient for the Society’s garden at Turnham Green; the Secretary of the Horticultural Society of London at that time was Joseph Sabine and he authorised expenditure on large projects beyond the Society’s means. Lindley was unsuccessful in moderating his actions. By 1830, the Society had mounting debts and a committee of enquiry was set up. Sabine resigned as Secretary and Lindley defended his own position and carried the Society forward with the new Honorary Secretary, George Bentham. An eminent botanist of the time, John Claudius Loudon, sought Lindley’s collaboration on his “Encyclopedia of Plants”; this covered nearly fifteen thousand species of flowering ferns. It was a massive undertaking and Lindley was responsible for most of it. During his labour on this undertaking, completed in 1829, through arduous study of character patterns, he became convinced of the superiority of the "natural" classification system devised by Antoine Laurent de Jussieu – a system that he believed reflected the great plan of nature as distinct from the "artificial" system of Linnaeus followed in the Encyclopaedi
The flowering plants known as angiosperms, Angiospermae or Magnoliophyta, are the most diverse group of land plants, with 64 orders, 416 families 13,164 known genera and c. 369,000 known species. Like gymnosperms, angiosperms are seed-producing plants. However, they are distinguished from gymnosperms by characteristics including flowers, endosperm within the seeds, the production of fruits that contain the seeds. Etymologically, angiosperm means a plant; the term comes from the Greek words sperma. The ancestors of flowering plants diverged from gymnosperms in the Triassic Period, 245 to 202 million years ago, the first flowering plants are known from 160 mya, they diversified extensively during the Early Cretaceous, became widespread by 120 mya, replaced conifers as the dominant trees from 100 to 60 mya. Angiosperms differ from other seed plants in several ways, described in the table below; these distinguishing characteristics taken together have made the angiosperms the most diverse and numerous land plants and the most commercially important group to humans.
Angiosperm stems are made up of seven layers. The amount and complexity of tissue-formation in flowering plants exceeds that of gymnosperms; the vascular bundles of the stem are arranged such that the phloem form concentric rings. In the dicotyledons, the bundles in the young stem are arranged in an open ring, separating a central pith from an outer cortex. In each bundle, separating the xylem and phloem, is a layer of meristem or active formative tissue known as cambium. By the formation of a layer of cambium between the bundles, a complete ring is formed, a regular periodical increase in thickness results from the development of xylem on the inside and phloem on the outside; the soft phloem becomes crushed, but the hard wood persists and forms the bulk of the stem and branches of the woody perennial. Owing to differences in the character of the elements produced at the beginning and end of the season, the wood is marked out in transverse section into concentric rings, one for each season of growth, called annual rings.
Among the monocotyledons, the bundles are more numerous in the young stem and are scattered through the ground tissue. They once formed the stem increases in diameter only in exceptional cases; the characteristic feature of angiosperms is the flower. Flowers show remarkable variation in form and elaboration, provide the most trustworthy external characteristics for establishing relationships among angiosperm species; the function of the flower is to ensure fertilization of the ovule and development of fruit containing seeds. The floral apparatus may arise terminally from the axil of a leaf; as in violets, a flower arises singly in the axil of an ordinary foliage-leaf. More the flower-bearing portion of the plant is distinguished from the foliage-bearing or vegetative portion, forms a more or less elaborate branch-system called an inflorescence. There are two kinds of reproductive cells produced by flowers. Microspores, which will divide to become pollen grains, are the "male" cells and are borne in the stamens.
The "female" cells called megaspores, which will divide to become the egg cell, are contained in the ovule and enclosed in the carpel. The flower may consist only of these parts, as in willow, where each flower comprises only a few stamens or two carpels. Other structures are present and serve to protect the sporophylls and to form an envelope attractive to pollinators; the individual members of these surrounding structures are known as petals. The outer series is green and leaf-like, functions to protect the rest of the flower the bud; the inner series is, in general, white or brightly colored, is more delicate in structure. It functions to attract bird pollinators. Attraction is effected by color and nectar, which may be secreted in some part of the flower; the characteristics that attract pollinators account for the popularity of flowers and flowering plants among humans. While the majority of flowers are perfect or hermaphrodite, flowering plants have developed numerous morphological and physiological mechanisms to reduce or prevent self-fertilization.
Heteromorphic flowers have short carpels and long stamens, or vice versa, so animal pollinators cannot transfer pollen to the pistil. Homomorphic flowers may employ a biochemical mechanism called self-incompatibility to discriminate between self and non-self pollen grains. In other species, the male and female parts are morphologically separated, developing on different flowers; the botanical term "Angiosperm", from the Ancient Greek αγγείον, angeíon and σπέρμα, was coined in the form Angiospermae by Paul Hermann in 1690, as the name of one of his primary divisions of the plant kingdom. This included flowering plants possessing seeds enclosed in capsules, distinguished from his Gymnospermae, or flowering plants with achenial or schizo-carpic fruits, the whole fruit or each of its pieces being here regarded as a seed and naked; the term and its antonym were maintained by Carl Linnaeus with the same sense, but with restricted application, in the names of the orders of his class Didynamia. Its use with any
Monocotyledons referred to as monocots, are flowering plants whose seeds contain only one embryonic leaf, or cotyledon. They constitute one of the major groups into which the flowering plants have traditionally been divided, the rest of the flowering plants having two cotyledons and therefore classified as dicotyledons, or dicots. However, molecular phylogenetic research has shown that while the monocots form a monophyletic group or clade, the dicots do not. Monocots have always been recognized as a group, but with various taxonomic ranks and under several different names; the APG III system of 2009 recognises a clade called "monocots" but does not assign it to a taxonomic rank. The monocots include about 60,000 species; the largest family in this group by number of species are the orchids, with more than 20,000 species. About half as many species belong to the true grasses, which are economically the most important family of monocots. In agriculture the majority of the biomass produced; these include not only major grains, but forage grasses, sugar cane, the bamboos.
Other economically important monocot crops include various palms and plantains, gingers and their relatives and cardamom, pineapple, water chestnut, leeks and garlic. Many houseplants are monocot epiphytes. Additionally most of the horticultural bulbs, plants cultivated for their blooms, such as lilies, irises, cannas and tulips, are monocots; the monocots or monocotyledons have, as the name implies, a single cotyledon, or embryonic leaf, in their seeds. This feature was used to contrast the monocots with the dicotyledons or dicots which have two cotyledons. From a diagnostic point of view the number of cotyledons is neither a useful characteristic, nor is it reliable; the single cotyledon is only one of a number of modifications of the body plan of the ancestral monocotyledons, whose adaptive advantages are poorly understood, but may have been related to adaption to aquatic habitats, prior to radiation to terrestrial habitats. Monocots are sufficiently distinctive that there has been disagreement as to membership of this group, despite considerable diversity in terms of external morphology.
However, morphological features that reliably characterise major clades are rare. Thus monocots are distinguishable from other angiosperms both in terms of their uniformity and diversity. On the one hand the organisation of the shoots, leaf structure and floral configuration are more uniform than in the remaining angiosperms, yet within these constraints a wealth of diversity exists, indicating a high degree of evolutionary success. Monocot diversity includes perennial geophytes such as ornamental flowers including and succulent epiphytes, all in the lilioid monocots, major cereal grains in the grass family and forage grasses as well as woody tree-like palm trees, bamboo and bromeliads, bananas and ginger in the commelinid monocots, as well as both emergent and aroids, as well as floating or submerged aquatic plants such as seagrass. Organisation and life formsThe most important distinction is their growth pattern, lacking a lateral meristem that allows for continual growth in diameter with height, therefore this characteristic is a basic limitation in shoot construction.
Although herbaceous, some arboraceous monocots reach great height and mass. The latter include agaves, palms and bamboos; this creates challenges in water transport. Some, such as species of Yucca, develop anomalous secondary growth, while palm trees utilise an anomalous primary growth form described as establishment growth; the axis undergoes primary thickening, that progresses from internode to internode, resulting in a typical inverted conical shape of the basal primary axis. The limited conductivity contributes to limited branching of the stems. Despite these limitations a wide variety of adaptive growth forms has resulted from epiphytic orchids and bromeliads to submarine Alismatales and mycotrophic Burmanniaceae and Triuridaceae. Other forms of adaptation include the climbing vines of Araceae which use negative phototropism to locate host trees, while some palms such as Calamus manan produce the longest shoots in the plant kingdom, up to 185 m long. Other monocots Poales, have adopted a therophyte life form.
LeavesThe cotyledon, the primordial Angiosperm leaf consists of a proximal leaf base or hypophyll and a distal hyperphyll. In monocots the hypophyll tends to be the dominant part in contrast to other angiosperms. From these, considerable diversity arises. Mature monocot leaves are narrow and linear, forming a sheath
Bletia is a genus of about 30 species of orchids all of which are terrestrial. It is named after pharmacist Don Luis Blet; the genus is widespread across Florida, Central America, the West Indies, South America as far south as Argentina. Accepted species as of May 2014: Bletia adenocarpa Rchb.f. Bletia amabilis C. Schweinf. Bletia antillana M. A. Diaz & Sosa Bletia campanulata Lex. Bletia candida Kraenzl. Bletia catenulata Ruiz & Pav. Bletia coccinea Lex. Bletia concolor Dressler Bletia ensifolia L. O. Williams Bletia florida R. Br. Bletia gracilis Lodd. Bletia greenmaniana L. O. Williams Bletia greenwoodiana Sosa Bletia lilacina A. Rich. & Galeotti Bletia macristhmochila Greenm. Bletia meridana Garay & Dunst. Bletia neglecta Sosa Bletia nelsonii Ames Bletia netzeri Senghas Bletia parkinsonii Hook. Bletia parviflora Ruiz & Pav. Bletia patula Hook. Bletia punctata Lex. Bletia purpurata A. Rich. & Galeotti Bletia purpurea DC. Bletia reflexa Lindl. Bletia repanda Ruiz & Pav. Bletia riparia Sosa & Palestina Bletia roezlii Rchb.f.
Bletia similis Dressler Bletia stenophylla Schltr. Bletia tenuifolia Ames & C. Schweinf. Bletia uniflora Ruiz & Pav. Bletia urbana Dressler Bletia warfordiana Sosa Media related to Bletia at Wikimedia Commons Data related to Bletia at Wikispecies
In plant systematics Epidendroideae is a subfamily of the orchid family, Orchidaceae. Epidendroideae is larger than all the other orchid subfamilies together, comprising more than 15,000 species in 576 genera. Most Epidendroid orchids are tropical epiphytes with pseudobulbs. There are, some terrestrials such as Epipactis and a few myco-heterotrophs, which are parasitic upon mycorrhizal fungi, they contain the remaining orchids with a single, fertile anther, fully incumbent to suberect. The anther form arises from early anther bending; the incumbent anther is pointed backward in many genera. Most have hard pollinia, i.e. a mass of waxy pollen or of coherent pollen grains. The pollinia are without; the stigma are three-lobed. The apical part of the middle stigma lobe forms a stipe; the ovary is unilocular. The leaves are distichous or spiraling; the Epidendroideae are difficult to classify. They have been divided in “lower epidendroids” and “higher epidendroids”. Epiphytes are plants which grow on top of other plants.
They are not parasitic. By growing on other plants, the epiphytes can reach to the light better or where they can avoid struggling for light. Many mosses and lichens are epiphytes, as are 10 per cent of all seed plants and ferns. Epiphytes are common in some groups of plants, such as ferns, mosses and algae. Over half of the 20,000 species of orchids are epiphytes. Most epiphytic seed plants and ferns are found in tropical and subtropical rainforests because they need high humidity to survive; the areas which most epiphytes grow are the montane rainforests. Epiphytic orchids are found on many positions of the host tree, depending on species requirements and size, some large species will grow in a fork, whereas some small species scramble through thin branches, other species will climb up the trunk etc. etc. The trees provide many habitats with different conditions of temperature and light. In temperate places, epiphytes are most common in moist forests, such as the rainforests in Queensland. Epiphytes are not adapted to droughts in the same way are other flora, because they don’t have access to the ground, but they still have some mechanisms to help them survive.
Some become dormant for months at a time. They contain absorptive plants that are capable at taking up water when it is available and preventing drought when water is scarcer. CAM can be impeded by higher night-time temperatures, dehydrated tissues, high saturation deficits in the surrounding air, which lower the "stomata conductance" of the epiphytes, reducing the CO2 uptake, which in turn reduces growth and reproduction and induces carbon loss. Higher temperatures, strain on evaporation, contact to light cause CAM-idling, the epiphyte closing its stomata when it becomes stressed, that brings down the range of habitats a species can inhabit. Epiphyte species work biomasses are much more sensitive to different relative moisture levels than other plants; the Epidendroideae subfamily is divided into two clades or subgroups known as the higher epidendroids and the lower epidendroids. The higher epidendroids are monophyletic and polyphyletic; the tribes are listed below: This classification has a rather ephemeral nature and is prone to frequent revision.
Changes are to occur as new morphological and genetic data become available. A phylogenetic analysis of the Orchidaceae - evidence from rbcL nucleotide sequences Orchid Tree: a phylogeny of epiphytes on the tree of life
Bletia catenulata is a species of orchid native to Ecuador, Brazil, Colombia and Paraguay. IOSPE orchid photo Brazil Plants Cuidado de Orquídeas AgrOriente, Orquídeas Amazonicas, Galería Virtual
Wikispecies is a wiki-based online project supported by the Wikimedia Foundation. Its aim is to create a comprehensive free content catalogue of all species. Jimmy Wales stated that editors are not required to fax in their degrees, but that submissions will have to pass muster with a technical audience. Wikispecies is available under the GNU Free Documentation License and CC BY-SA 3.0. Started in September 2004, with biologists across the world invited to contribute, the project had grown a framework encompassing the Linnaean taxonomy with links to Wikipedia articles on individual species by April 2005. Benedikt Mandl co-ordinated the efforts of several people who are interested in getting involved with the project and contacted potential supporters in early summer 2004. Databases were evaluated and the administrators contacted, some of them have agreed on providing their data for Wikispecies. Mandl defined two major tasks: Figure out how the contents of the data base would need to be presented—by asking experts, potential non-professional users and comparing that with existing databases Figure out how to do the software, which hardware is required and how to cover the costs—by asking experts, looking for fellow volunteers and potential sponsorsAdvantages and disadvantages were discussed by the wikimedia-I mailing list.
The board of directors of the Wikimedia Foundation voted by 4 to 0 in favor of the establishment of a Wikispecies. The project is hosted at species.wikimedia.org. It was merged to a sister project of Wikimedia Foundation on September 14, 2004. On October 10, 2006, the project exceeded 75,000 articles. On May 20, 2007, the project exceeded 100,000 articles with a total of 5,495 registered users. On September 8, 2008, the project exceeded 150,000 articles with a total of 9,224 registered users. On October 23, 2011, the project reached 300,000 articles. On June 16, 2014, the project reached 400,000 articles. On January 7, 2017, the project reached 500,000 articles. On October 30, 2018, the project reached 600,000 articles, a total of 1.12 million pages. Wikispecies comprises taxon pages, additionally pages about synonyms, taxon authorities, taxonomical publications, institutions or repositories holding type specimen. Wikispecies asks users to use images from Wikimedia Commons. Wikispecies does not allow the use of content.
All Species Foundation Catalogue of Life Encyclopedia of Life Tree of Life Web Project List of online encyclopedias The Plant List Wikispecies, The free species directory that anyone can edit Species Community Portal The Wikispecies Charter, written by Wales