The Cardueae are a tribe of flowering plants in the daisy family (Asteraceae) and the subfamily Carduoideae. Most of them are commonly known as thistles; four of the best known genera are Carduus, Cynara, Cirsium, and Onopordum.
Cardueae
Thistle is the common name of a group of flowering plants characterised by leaves with sharp prickles on the margins, mostly in the family Asteraceae. Prickles can also occur all over the plant – on the stem and on the flat parts of the leaves. These prickles protect the plant from herbivores. Typically, an involucre with a clasping shape similar to a cup or urn subtends each of a thistle's flower heads. The typically feathery pappus of a ripe thistle flower is known as thistle-down.
Milk thistle flowerhead
Cirsium arizonicum, showing arachnoid cobwebbiness on stems and leaves, with ants attending aphids that might be taking advantage of the shelter.
Thistles on Calton Hill in central Edinburgh, Scotland, 2023
Carduus nutans in the early morning light.