An inflorescence is a group or cluster of flowers arranged on a stem, composed of a main branch or a complicated arrangement of branches. Morphologically, it is the modified part of the shoot of seed plants; the modifications can involve the length and the nature of the internodes and the phyllotaxis, as well as variations in the proportions, swellings, adnations and reduction of main and secondary axes. Inflorescence can be defined as the reproductive portion of a plant that bears a cluster of flowers in a specific pattern; the stem holding the whole inflorescence is called a peduncle and the major axis holding the flowers or more branches within the inflorescence is called the rachis. The stalk of each single flower is called a pedicel. A flower, not part of an inflorescence is called a solitary flower and its stalk is referred to as a peduncle. Any flower in an inflorescence may be referred to as a floret when the individual flowers are small and borne in a tight cluster, such as in a pseudanthium.
The fruiting stage of an inflorescence is known as an infructescence. Inflorescences may be complex; the rachis may be one of several types, including single, umbel, spike or raceme. Inflorescences are described by many different characteristics including how the flowers are arranged on the peduncle, the blooming order of the flowers and how different clusters of flowers are grouped within it; these terms are general representations. Inflorescences have modified shoots foliage different from the vegetative part of the plant. Considering the broadest meaning of the term, any leaf associated with an inflorescence is called a bract. A bract is located at the node where the main stem of the inflorescence forms, joined to the main stem of the plant, but other bracts can exist within the inflorescence itself, they serve a variety of functions which include protecting young flowers. According to the presence or absence of bracts and their characteristics we can distinguish: Ebracteate inflorescences: No bracts in the inflorescence.
Bracteate inflorescences: The bracts in the inflorescence are specialised, sometimes reduced to small scales, divided or dissected. Leafy inflorescences: Though reduced in size, the bracts are unspecialised and look like the typical leaves of the plant, so that the term flowering stem is applied instead of inflorescence; this use is not technically correct, as, despite their'normal' appearance, these leaves are considered, in fact, bracts, so that'leafy inflorescence' is preferable. Leafy-bracted inflorescences: Intermediate between bracteate and leafy inflorescence. If many bracts are present and they are connected to the stem, like in the family Asteraceae, the bracts might collectively be called an involucre. If the inflorescence has a second unit of bracts further up the stem, they might be called an involucel. Plant organs can grow according to two different schemes, namely monopodial or racemose and sympodial or cymose. In inflorescences these two different growth patterns are called indeterminate and determinate and indicate whether a terminal flower is formed and where flowering starts within the inflorescence.
Indeterminate inflorescence: Monopodial growth. The terminal bud keeps forming lateral flowers. A terminal flower is never formed. Determinate inflorescence: Sympodial growth; the terminal bud forms a terminal flower and dies out. Other flowers grow from lateral buds. Indeterminate and determinate inflorescences are sometimes referred to as open and closed inflorescences respectively; the indeterminate patterning of flowers is derived from determinate flowers. It is suggested that indeterminate flowers have a common mechanism that prevents terminal flower growth. Based on phylogenetic analyses, this mechanism arose independently multiple times in different species. In an indeterminate inflorescence there is no true terminal flower and the stem has a rudimentary end. In many cases the last true flower formed by the terminal bud straightens up, appearing to be a terminal flower. A vestige of the terminal bud may be noticed higher on the stem. In determinate inflorescences the terminal flower is the first to mature, while the others tend to mature starting from the bottom of the stem.
This pattern is called acropetal maturation. When flowers start to mature from the top of the stem, maturation is basipetal, while when the central mature first, divergent; as with leaves, flowers can be arranged on the stem according to many different patterns. See'Phyllotaxis' for in-depth descriptions Similarly arrangement of leaf in bud is called Ptyxis. Metatopy is the placement of organs out of their expected position: metatopy occurs in inflorescences when unequal growth rates alter different areas of the axis and the organs attached to the axis; when a single or a cluster of flower is located at the axil of a bract, the location of the bract in relation to the stem holding the flower is indicated by the use of different terms and may be a useful diagnostic indicator. Typical placement of bracts include: Some plants have bracts that subtend the inflorescence, where the flowers are on branched stalks.
An umbel is an inflorescence that consists of a number of short flower stalks which spread from a common point, somewhat like umbrella ribs. The word was coined in botany in the 1590s, from Latin umbella "parasol, sunshade"; the arrangement can vary from being flat topped to spherical. Umbels can compound; the secondary umbels of compound umbels are known as umbellets. A small umbel is called an umbellule; the arrangement of the inflorescence in umbels is referred to as umbellate, or subumbellate. Umbels are a characteristic of plants such as carrot, parsley and fennel in the family Apiaceae. An umbel is a type of indeterminate inflorescence. A compressed cyme, a determinate inflorescence, is called umbelliform if it resembles an umbel. Hinderer, Walter. "Differentiation of metabolic pathways in the umbel of Daucus carota". Phytochemistry. 22: 2417–2420. Doi:10.1016/0031-942280131-9. ISSN 0031-9422. Toben, H.-M.. Journal of Phytopathology. 144: 169–178. Doi:10.1111/j.1439-0434.1996.tb01510.x. ISSN 0931-1785.
Peterson, L. E.. C.. "Umbel Initiation and Stem Elongation in FennelInitiated by Photoperiod". Journal of Essential Oil Research. 5: 37–43. Doi:10.1080/10412905.1993.9698168. ISSN 1041-2905
A panicle is a much-branched inflorescence. Some authors distinguish it by requiring that the flowers be pedicellate; the branches of a panicle are racemes. A panicle may have indeterminate growth; this type of inflorescence is characteristic of grasses such as oat and crabgrass, as well as other plants such as pistachio and mamoncillo. Botanists use the term paniculate in two ways: "having a true panicle inflorescence" as well as "having an inflorescence with the form but not the structure of a panicle". A corymb may have a paniculate branching structure, with the lower flowers having longer pedicels than the upper, thus giving a flattish top superficially resembling an umbel. Many species in the subfamily Amygdaloideae, such as hawthorns and rowans, produce their flowers in corymbs. Thyrse, a branched inflorescence where the main axis has indeterminate growth, the branches have determinate growth
A flower, sometimes known as a bloom or blossom, is the reproductive structure found in flowering plants. The biological function of a flower is to effect reproduction by providing a mechanism for the union of sperm with eggs. Flowers may allow selfing; some flowers produce diaspores without fertilization. Flowers are the site where gametophytes develop. Many flowers have evolved to be attractive to animals, so as to cause them to be vectors for the transfer of pollen. After fertilization, the ovary of the flower develops into fruit containing seeds. In addition to facilitating the reproduction of flowering plants, flowers have long been admired and used by humans to bring beauty to their environment, as objects of romance, religion, medicine and as a source of food; the essential parts of a flower can be considered in two parts: the vegetative part, consisting of petals and associated structures in the perianth, the reproductive or sexual parts. A stereotypical flower consists of four kinds of structures attached to the tip of a short stalk.
Each of these kinds of parts is arranged in a whorl on the receptacle. The four main whorls are as follows: Collectively the calyx and corolla form the perianth. Calyx: the outermost whorl consisting of units called sepals. Corolla: the next whorl toward the apex, composed of units called petals, which are thin and colored to attract animals that help the process of pollination. Androecium: the next whorl, consisting of units called stamens. Stamens consist of two parts: a stalk called a filament, topped by an anther where pollen is produced by meiosis and dispersed. Gynoecium: the innermost whorl of a flower, consisting of one or more units called carpels; the carpel or multiple fused carpels form a hollow structure called an ovary, which produces ovules internally. Ovules are megasporangia and they in turn produce megaspores by meiosis which develop into female gametophytes; these give rise to egg cells. The gynoecium of a flower is described using an alternative terminology wherein the structure one sees in the innermost whorl is called a pistil.
A pistil may consist of a number of carpels fused together. The sticky tip of the pistil, the stigma, is the receptor of pollen; the supportive stalk, the style, becomes the pathway for pollen tubes to grow from pollen grains adhering to the stigma. The relationship to the gynoecium on the receptacle is described as hypogynous, perigynous, or epigynous. Although the arrangement described above is considered "typical", plant species show a wide variation in floral structure; these modifications have significance in the evolution of flowering plants and are used extensively by botanists to establish relationships among plant species. The four main parts of a flower are defined by their positions on the receptacle and not by their function. Many flowers lack some parts or parts may be modified into other functions and/or look like what is another part. In some families, like Ranunculaceae, the petals are reduced and in many species the sepals are colorful and petal-like. Other flowers have modified stamens.
Flowers show great variation and plant scientists describe this variation in a systematic way to identify and distinguish species. Specific terminology is used to describe their parts. Many flower parts are fused together; when petals are fused into a tube or ring that falls away as a single unit, they are sympetalous. Connate petals may have distinctive regions: the cylindrical base is the tube, the expanding region is the throat and the flaring outer region is the limb. A sympetalous flower, with bilateral symmetry with an upper and lower lip, is bilabiate. Flowers with connate petals or sepals may have various shaped corolla or calyx, including campanulate, tubular, salverform or rotate. Referring to "fusion," as it is done, appears questionable because at least some of the processes involved may be non-fusion processes. For example, the addition of intercalary growth at or below the base of the primordia of floral appendages such as sepals, petals and carpels may lead to a common base, not the result of fusion.
Many flowers have a symmetry. When the perianth is bisected through the central axis from any point and symmetrical halves are produced, the flower is said to be actinomorphic or regular, e.g. rose or trillium. This is an example of radial symmetry; when flowers are bisected and produce only one line that produces symmetrical halves, the flower is said to be irregular or zygomorphic, e.g. snapdragon or most orchids. Flowers may be directly attached to the plant at their base; the stem or stalk subtending a flower is called a peduncle. If a peduncle supports more than o
Botany called plant science, plant biology or phytology, is the science of plant life and a branch of biology. A botanist, plant scientist or phytologist is a scientist; the term "botany" comes from the Ancient Greek word βοτάνη meaning "pasture", "grass", or "fodder". Traditionally, botany has included the study of fungi and algae by mycologists and phycologists with the study of these three groups of organisms remaining within the sphere of interest of the International Botanical Congress. Nowadays, botanists study 410,000 species of land plants of which some 391,000 species are vascular plants, 20,000 are bryophytes. Botany originated in prehistory as herbalism with the efforts of early humans to identify – and cultivate – edible and poisonous plants, making it one of the oldest branches of science. Medieval physic gardens attached to monasteries, contained plants of medical importance, they were forerunners of the first botanical gardens attached to universities, founded from the 1540s onwards.
One of the earliest was the Padua botanical garden. These gardens facilitated the academic study of plants. Efforts to catalogue and describe their collections were the beginnings of plant taxonomy, led in 1753 to the binomial system of Carl Linnaeus that remains in use to this day. In the 19th and 20th centuries, new techniques were developed for the study of plants, including methods of optical microscopy and live cell imaging, electron microscopy, analysis of chromosome number, plant chemistry and the structure and function of enzymes and other proteins. In the last two decades of the 20th century, botanists exploited the techniques of molecular genetic analysis, including genomics and proteomics and DNA sequences to classify plants more accurately. Modern botany is a broad, multidisciplinary subject with inputs from most other areas of science and technology. Research topics include the study of plant structure and differentiation, reproduction and primary metabolism, chemical products, diseases, evolutionary relationships and plant taxonomy.
Dominant themes in 21st century plant science are molecular genetics and epigenetics, which are the mechanisms and control of gene expression during differentiation of plant cells and tissues. Botanical research has diverse applications in providing staple foods, materials such as timber, rubber and drugs, in modern horticulture and forestry, plant propagation and genetic modification, in the synthesis of chemicals and raw materials for construction and energy production, in environmental management, the maintenance of biodiversity. Botany originated as the study and use of plants for their medicinal properties. Many records of the Holocene period date early botanical knowledge as far back as 10,000 years ago; this early unrecorded knowledge of plants was discovered in ancient sites of human occupation within Tennessee, which make up much of the Cherokee land today. The early recorded history of botany includes many ancient writings and plant classifications. Examples of early botanical works have been found in ancient texts from India dating back to before 1100 BC, in archaic Avestan writings, in works from China before it was unified in 221 BC.
Modern botany traces its roots back to Ancient Greece to Theophrastus, a student of Aristotle who invented and described many of its principles and is regarded in the scientific community as the "Father of Botany". His major works, Enquiry into Plants and On the Causes of Plants, constitute the most important contributions to botanical science until the Middle Ages seventeen centuries later. Another work from Ancient Greece that made an early impact on botany is De Materia Medica, a five-volume encyclopedia about herbal medicine written in the middle of the first century by Greek physician and pharmacologist Pedanius Dioscorides. De Materia Medica was read for more than 1,500 years. Important contributions from the medieval Muslim world include Ibn Wahshiyya's Nabatean Agriculture, Abū Ḥanīfa Dīnawarī's the Book of Plants, Ibn Bassal's The Classification of Soils. In the early 13th century, Abu al-Abbas al-Nabati, Ibn al-Baitar wrote on botany in a systematic and scientific manner. In the mid-16th century, "botanical gardens" were founded in a number of Italian universities – the Padua botanical garden in 1545 is considered to be the first, still in its original location.
These gardens continued the practical value of earlier "physic gardens" associated with monasteries, in which plants were cultivated for medical use. They supported the growth of botany as an academic subject. Lectures were given about the plants grown in the gardens and their medical uses demonstrated. Botanical gardens came much to northern Europe. Throughout this period, botany remained subordinate to medicine. German physician Leonhart Fuchs was one of "the three German fathers of botany", along with theologian Otto Brunfels and physician Hieronymus Bock. Fuchs and Brunfels broke away from the tradition of copying earlier works to make original observations of their own. Bock created his own system of plant classification. Physician Valerius Cordus authored a botanically and pharmacologically important herbal Historia Plantarum in 1544 and a pharmacopoeia of lasting importance, the Dispensatorium
The Ancient Greek language includes the forms of Greek used in Ancient Greece and the ancient world from around the 9th century BCE to the 6th century CE. It is roughly divided into the Archaic period, Classical period, Hellenistic period, it is succeeded by medieval Greek. Koine is regarded as a separate historical stage of its own, although in its earliest form it resembled Attic Greek and in its latest form it approaches Medieval Greek. Prior to the Koine period, Greek of the classic and earlier periods included several regional dialects. Ancient Greek was the language of Homer and of fifth-century Athenian historians and philosophers, it has contributed many words to English vocabulary and has been a standard subject of study in educational institutions of the Western world since the Renaissance. This article contains information about the Epic and Classical periods of the language. Ancient Greek was a pluricentric language, divided into many dialects; the main dialect groups are Attic and Ionic, Aeolic and Doric, many of them with several subdivisions.
Some dialects are found in standardized literary forms used in literature, while others are attested only in inscriptions. There are several historical forms. Homeric Greek is a literary form of Archaic Greek used in the epic poems, the "Iliad" and "Odyssey", in poems by other authors. Homeric Greek had significant differences in grammar and pronunciation from Classical Attic and other Classical-era dialects; the origins, early form and development of the Hellenic language family are not well understood because of a lack of contemporaneous evidence. Several theories exist about what Hellenic dialect groups may have existed between the divergence of early Greek-like speech from the common Proto-Indo-European language and the Classical period, they differ in some of the detail. The only attested dialect from this period is Mycenaean Greek, but its relationship to the historical dialects and the historical circumstances of the times imply that the overall groups existed in some form. Scholars assume that major Ancient Greek period dialect groups developed not than 1120 BCE, at the time of the Dorian invasion—and that their first appearances as precise alphabetic writing began in the 8th century BCE.
The invasion would not be "Dorian" unless the invaders had some cultural relationship to the historical Dorians. The invasion is known to have displaced population to the Attic-Ionic regions, who regarded themselves as descendants of the population displaced by or contending with the Dorians; the Greeks of this period believed there were three major divisions of all Greek people—Dorians and Ionians, each with their own defining and distinctive dialects. Allowing for their oversight of Arcadian, an obscure mountain dialect, Cypriot, far from the center of Greek scholarship, this division of people and language is quite similar to the results of modern archaeological-linguistic investigation. One standard formulation for the dialects is: West vs. non-west Greek is the strongest marked and earliest division, with non-west in subsets of Ionic-Attic and Aeolic vs. Arcadocypriot, or Aeolic and Arcado-Cypriot vs. Ionic-Attic. Non-west is called East Greek. Arcadocypriot descended more from the Mycenaean Greek of the Bronze Age.
Boeotian had come under a strong Northwest Greek influence, can in some respects be considered a transitional dialect. Thessalian had come under Northwest Greek influence, though to a lesser degree. Pamphylian Greek, spoken in a small area on the southwestern coast of Anatolia and little preserved in inscriptions, may be either a fifth major dialect group, or it is Mycenaean Greek overlaid by Doric, with a non-Greek native influence. Most of the dialect sub-groups listed above had further subdivisions equivalent to a city-state and its surrounding territory, or to an island. Doric notably had several intermediate divisions as well, into Island Doric, Southern Peloponnesus Doric, Northern Peloponnesus Doric; the Lesbian dialect was Aeolic Greek. All the groups were represented by colonies beyond Greece proper as well, these colonies developed local characteristics under the influence of settlers or neighbors speaking different Greek dialects; the dialects outside the Ionic group are known from inscriptions, notable exceptions being: fragments of the works of the poet Sappho from the island of Lesbos, in Aeolian, the poems of the Boeotian poet Pindar and other lyric poets in Doric.
After the conquests of Alexander the Great in the late 4th century BCE, a new international dialect known as Koine or Common Greek developed based on Attic Greek, but with influence from other dialects. This dialect replaced most of the older dialects, although Doric dialect has survived in the Tsakonian language, spoken in the region of modern Sparta. Doric has passed down its aorist terminations into most verbs of Demotic Greek. By about the 6th century CE, the Koine had metamorphosized into Medieval Greek. Ancient Macedonian was an Indo-European language at least related to Greek, but its exact relationship is unclear because of insufficient data: a dialect of Greek; the Macedonian dialect (or l
A pedicel is a stem that attaches a single flower to the inflorescence. In the absence of a pedicel, the flowers are described as sessile. Pedicel is applied to the stem of the infructescence; the word "pedicel" is derived from the latin pediculus, meaning "little foot". The stem or branch from the main stem of the inflorescence that holds a group of pedicels is called a peduncle. In Halloween types of pumpkin or squash plants, the shape of the pedicel has received particular attention because plant breeders are trying to optimize the size and shape of the pedicel for the best "lid" for a "jack-o'-lantern". Sessile Scape Terminology for Asteraceae