1.
Precambrian
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The Precambrian is the earliest period of Earths history, set before the current Phanerozoic Eon. The Precambrian is a supereon that is subdivided into three eons of the time scale. It spans from the formation of Earth about 4.6 billion years ago to the beginning of the Cambrian Period, about 541 million years ago, the Precambrian accounts for 89% of geologic time. Relatively little is known about the Precambrian, despite it making up roughly seven-eighths of the Earths history, the Precambrian fossil record is poorer than that of the succeeding Phanerozoic, and fossils from that time are of limited biostratigraphic use. This is because many Precambrian rocks have been metamorphosed, obscuring their origins, while others have been destroyed by erosion. A stable crust was apparently in place by 4,412 Ma, the term Precambrian is recognized by the International Commission on Stratigraphy as a general term including the Archean and Proterozoic eons. It is still used by geologists and paleontologists for general discussions not requiring the more specific eon names and it was briefly called the Cryptozoic eon. A specific date for the origin of life has not been determined, carbon found in 3.8 billion year old rocks from islands off western Greenland may be of organic origin. Well-preserved microscopic fossils of bacteria older than 3.46 billion years have found in Western Australia. Probable fossils 100 million years older have been found in the same area, there is a fairly solid record of bacterial life throughout the remainder of the Precambrian. The oldest fossil evidence from that era of such complex life comes from the Lantian formation of the Ediacarian period, a very diverse collection of soft-bodied forms is found in a variety of locations worldwide and date to between 635 and 542 Ma. These are referred to as Ediacaran or Vendian biota, hard-shelled creatures appeared toward the end of that time span, marking the beginning of the Phanerozoic era. By the middle of the following Cambrian period, a diverse fauna is recorded in the Burgess Shale. The explosion in diversity of lifeforms during the early Cambrian is called the Cambrian explosion of life, while land seems to have been devoid of plants and animals, cyanobacteria and other microbes formed prokaryotic mats that covered terrestrial areas. Evidence of the details of plate motions and other activity in the Precambrian has been poorly preserved. It is generally believed that small proto-continents existed prior to 4280 Ma, the supercontinent, known as Rodinia, broke up around 750 Ma. A number of glacial periods have been identified going as far back as the Huronian epoch, one of the best studied is the Sturtian-Varangian glaciation, around 850–635 Ma, which may have brought glacial conditions all the way to the equator, resulting in a Snowball Earth. The atmosphere of the early Earth is not well understood, most geologists believe it was composed primarily of nitrogen, carbon dioxide, and other relatively inert gases, and was lacking in free oxygen
2.
Cambrian
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The Cambrian Period was the first geological period of the Paleozoic Era, of the Phanerozoic Eon. The Cambrian lasted 55.6 million years from the end of the preceding Ediacaran Period 541 million years ago to the beginning of the Ordovician Period 485.4 mya and its subdivisions, and its base, are somewhat in flux. The period was established by Adam Sedgwick, who named it after Cambria, the Latinised form of Cymru, the Welsh name for Wales, as a result, our understanding of the Cambrian biology surpasses that of some later periods. The rapid diversification of lifeforms in the Cambrian, known as the Cambrian explosion, most of the continents were probably dry and rocky due to a lack of vegetation. Shallow seas flanked the margins of several continents created during the breakup of the supercontinent Pannotia, the seas were relatively warm, and polar ice was absent for much of the period. The United States Federal Geographic Data Committee uses a barred capital C ⟨Є⟩ character similar to the capital letter Ukrainian Ye ⟨Є⟩ to represent the Cambrian Period, the proper Unicode character is U+A792 Ꞓ LATIN CAPITAL LETTER C WITH BAR. Despite the long recognition of its distinction from younger Ordovician Period rocks and older Supereon Precambrian rocks, the base of the Cambrian lies atop a complex assemblage of trace fossils known as the Treptichnus pedum assemblage. Pedum in Namibia, Spain and Newfoundland, and possibly, in the western USA, the stratigraphic range of T. pedum overlaps the range of the Ediacaran fossils in Namibia, and probably in Spain. The Cambrian Period followed the Ediacaran Period and was followed by the Ordovician Period, the Cambrian is divided into four epochs and ten ages. Currently only two series and five stages are named and have a GSSP, because the international stratigraphic subdivision is not yet complete, many local subdivisions are still widely used. In some of these subdivisions the Cambrian is divided into three epochs with locally differing names – the Early Cambrian, Middle Cambrian and Furongian, rocks of these epochs are referred to as belonging to the Lower, Middle, or Upper Cambrian. Trilobite zones allow biostratigraphic correlation in the Cambrian, each of the local epochs is divided into several stages. The International Commission on Stratigraphy list the Cambrian period as beginning at 541 million years ago, the lower boundary of the Cambrian was originally held to represent the first appearance of complex life, represented by trilobites. The recognition of small shelly fossils before the first trilobites, and Ediacara biota substantially earlier and this formal designation allowed radiometric dates to be obtained from samples across the globe that corresponded to the base of the Cambrian. Early dates of 570 million years ago quickly gained favour, though the used to obtain this number are now considered to be unsuitable. A more precise date using modern radiometric dating yield a date of 541 ±0.3 million years ago, most continental land was clustered in the Southern Hemisphere at this time, but was drifting north. Large, high-velocity rotational movement of Gondwana appears to have occurred in the Early Cambrian, the sea levels fluctuated somewhat, suggesting there were ice ages, associated with pulses of expansion and contraction of a south polar ice cap. In Baltoscandia a Lower Cambrian transgression transformed large swathes of the Sub-Cambrian peneplain into a epicontinental sea, the Earth was generally cold during the early Cambrian, probably due to the ancient continent of Gondwana covering the South Pole and cutting off polar ocean currents
3.
Ordovician
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The Ordovician is a geologic period and system, the second of six periods of the Paleozoic Era. The Ordovician spans 41.2 million years from the end of the Cambrian Period 485.4 million years ago to the start of the Silurian Period 443.8 Mya. Lapworth recognized that the fauna in the disputed strata were different from those of either the Cambrian or the Silurian periods. It received international sanction in 1960, when it was adopted as a period of the Paleozoic Era by the International Geological Congress. Life continued to flourish during the Ordovician as it did in the earlier Cambrian period, invertebrates, namely molluscs and arthropods, dominated the oceans. The Great Ordovician Biodiversification Event considerably increased the diversity of life, fish, the worlds first true vertebrates, continued to evolve, and those with jaws may have first appeared late in the period. Life had yet to diversify on land, about 100 times as many meteorites struck the Earth during the Ordovician compared with today. The Ordovician Period began with a major extinction called the Cambrian–Ordovician extinction event and it lasted for about 42 million years and ended with the Ordovician–Silurian extinction event, about 443.8 Mya which wiped out 60% of marine genera. The dates given are recent radiometric dates and vary slightly from those found in other sources and this second period of the Paleozoic era created abundant fossils that became major petroleum and gas reservoirs. The boundary chosen for the beginning of both the Ordovician Period and the Tremadocian stage is highly significant and it correlates well with the occurrence of widespread graptolite, conodont, and trilobite species. The base of the Tremadocian allows scientists to relate these species not only to each other and this makes it easier to place many more species in time relative to the beginning of the Ordovician Period. A number of terms have been used to subdivide the Ordovician Period. In 2008, the ICS erected an international system of subdivisions. There exist Baltoscandic, British, Siberian, North American, Australian, the Ordovician Period in Britain was traditionally broken into Early, Middle and Late epochs. The corresponding rocks of the Ordovician System are referred to as coming from the Lower, Middle, the Floian corresponds to the lower Arenig, the Arenig continues until the early Darriwilian, subsuming the Dapingian. The Llanvirn occupies the rest of the Darriwilian, and terminates with it at the base of the Late Ordovician. The Sandbian represents the first half of the Caradoc, the Caradoc ends in the mid-Katian, during the Ordovician, the southern continents were collected into Gondwana. Gondwana started the period in equatorial latitudes and, as the period progressed, drifted toward the South Pole, the small continent Avalonia separated from Gondwana and began to move north towards Baltica and Laurentia, opening the Rheic Ocean between Gondwana and Avalonia
4.
Silurian
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The Silurian is a geologic period and system spanning 24.6 million years from the end of the Ordovician Period, at 443.8 million years ago, to the beginning of the Devonian Period,419.2 Mya. As with other periods, the rock beds that define the periods start and end are well identified. The base of the Silurian is set at a major Ordovician-Silurian extinction event when 60% of marine species were wiped out, a significant evolutionary milestone during the Silurian was the diversification of jawed and bony fish. However, terrestrial life would not greatly diversify and affect the landscape until the Devonian, the Silurian system was first identified by British geologist Sir Roderick Impey Murchison, who was examining fossil-bearing sedimentary rock strata in south Wales in the early 1830s. He named the sequences for a Celtic tribe of Wales, the Silures, inspired by his friend Adam Sedgwick and this naming does not indicate any correlation between the occurrence of the Silurian rocks and the land inhabited by the Silures. As it was first identified, the Silurian series when traced farther afield quickly came to overlap Sedgwicks Cambrian sequence, however, charles Lapworth resolved the conflict by defining a new Ordovician system including the contested beds. An early alternative name for the Silurian was Gotlandian after the strata of the Baltic island of Gotland, the French geologist Joachim Barrande, building on Murchisons work, used the term Silurian in a more comprehensive sense than was justified by subsequent knowledge. He divided the Silurian rocks of Bohemia into eight stages and his interpretation was questioned in 1854 by Edward Forbes, and the later stages of Barrande, F, G and H, have since been shown to be Devonian. Despite these modifications in the groupings of the strata, it is recognized that Barrande established Bohemia as a classic ground for the study of the earliest fossils. The epoch is named for the town of Llandovery in Carmarthenshire, the Wenlock, which lasted from 433.4 ±1.5 to 427.4 ±2.8 mya, is subdivided into the Sheinwoodian and Homerian ages. It is named after Wenlock Edge in Shropshire, England, during the Wenlock, the oldest known tracheophytes of the genus Cooksonia, appear. The first terrestrial animals also appear in the Wenlock, represented by air-breathing millipedes from Scotland. The Ludlow, lasting from 427.4 ±1.5 to 423 ±2.8 mya, comprises the Gorstian stage, lasting until 425.6 million years ago, and it is named for the town of Ludlow in Shropshire, England. The Pridoli, lasting from 423 ±1.5 to 419.2 ±2.8 mya, is the final and it is named after one locality at the Homolka a Přídolí nature reserve near the Prague suburb Slivenec in the Czech Republic. Přídolí is the old name of a field area. The high sea levels of the Silurian and the flat land resulted in a number of island chains. The southern continents remained united during this period, the melting of icecaps and glaciers contributed to a rise in sea level, recognizable from the fact that Silurian sediments overlie eroded Ordovician sediments, forming an unconformity. The continents of Avalonia, Baltica, and Laurentia drifted together near the equator and this event is the Caledonian orogeny, a spate of mountain building that stretched from New York State through conjoined Europe and Greenland to Norway
5.
Devonian
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The Devonian is a geologic period and system of the Paleozoic, spanning 60 million years from the end of the Silurian,419.2 million years ago, to the beginning of the Carboniferous,358.9 Mya. It is named after Devon, England, where rocks from this period were first studied, the first significant adaptive radiation of life on dry land occurred during the Devonian. Free-sporing vascular plants began to spread across dry land, forming extensive forests which covered the continents, by the middle of the Devonian, several groups of plants had evolved leaves and true roots, and by the end of the period the first seed-bearing plants appeared. Various terrestrial arthropods also became well-established, Fish reached substantial diversity during this time, leading the Devonian to often be dubbed the Age of Fish. The first ray-finned and lobe-finned bony fish appeared, while the placodermi began dominating almost every aquatic environment. The ancestors of all four-limbed vertebrates began adapting to walking on land, as their strong pectoral, in the oceans, primitive sharks became more numerous than in the Silurian and Late Ordovician. The first ammonites, species of molluscs, appeared, trilobites, the mollusk-like brachiopods and the great coral reefs, were still common. The Late Devonian extinction which started about 375 million years ago severely affected marine life, killing off all placodermi, and all trilobites, save for a few species of the order Proetida. The palaeogeography was dominated by the supercontinent of Gondwana to the south, the continent of Siberia to the north, while the rock beds that define the start and end of the Devonian period are well identified, the exact dates are uncertain. According to the International Commission on Stratigraphy, the Devonian extends from the end of the Silurian 419.2 Mya, another common term is Age of the Fishes, referring to the evolution of several major groups of fish that took place during the period. Older literature on the Anglo-Welsh basin divides it into the Downtonian, Dittonian, Breconian and Farlovian stages, in the Late Devonian, by contrast, arid conditions were less prevalent across the world and temperate climates were more common. The Devonian Period is formally broken into Early, Middle and Late subdivisions, the rocks corresponding to those epochs are referred to as belonging to the Lower, Middle and Upper parts of the Devonian System. Early Devonian The Early Devonian lasted from 419.2 ±2.8 to 393.3 ±2.5 and began with the Lochkovian stage, which lasted until the Pragian. It spanned from 410.8 ±2.8 to 407.6 ±2.5, and was followed by the Emsian, which lasted until the Middle Devonian began,393. 3±2.7 million years ago. Middle Devonian The Middle Devonian comprised two subdivisions, first the Eifelian, which gave way to the Givetian 387. 7±2.7 million years ago. Late Devonian Finally, the Late Devonian started with the Frasnian,382.7 ±2.8 to 372.2 ±2.5, during which the first forests took shape on land. The first tetrapods appeared in the record in the ensuing Famennian subdivision. This lasted until the end of the Devonian,358. 9±2.5 million years ago, the Devonian was a relatively warm period, and probably lacked any glaciers
6.
Carboniferous
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The Carboniferous is a geologic period and system that spans 60 million years from the end of the Devonian Period 358.9 million years ago, to the beginning of the Permian Period,298.9 Mya. The name Carboniferous means coal-bearing and derives from the Latin words carbō and ferō, and was coined by geologists William Conybeare and William Phillips in 1822. Based on a study of the British rock succession, it was the first of the system names to be employed. The Carboniferous is often treated in North America as two periods, the earlier Mississippian and the later Pennsylvanian. Terrestrial life was established by the Carboniferous period. Amphibians were the dominant land vertebrates, of one branch would eventually evolve into amniotes. Arthropods were also common, and many were much larger than those of today. Vast swaths of forest covered the land, which would eventually be laid down, the atmospheric content of oxygen also reached their highest levels in geological history during the period, 35% compared with 21% today, allowing terrestrial invertebrates to evolve to great size. A major marine and terrestrial extinction event, the Carboniferous rainforest collapse, occurred in the middle of the period, the later half of the period experienced glaciations, low sea level, and mountain building as the continents collided to form Pangaea. In the United States the Carboniferous is usually broken into Mississippian and Pennsylvanian subperiods, the Silesian is roughly contemporaneous with the late Mississippian Serpukhovian plus the Pennsylvanian. In Britain the Dinantian is traditionally known as the Carboniferous Limestone, the Namurian as the Millstone Grit, and the Westphalian as the Coal Measures and Pennant Sandstone. There was also a drop in south polar temperatures, southern Gondwanaland was glaciated throughout the period and these conditions apparently had little effect in the deep tropics, where lush swamps, later to become coal, flourished to within 30 degrees of the northernmost glaciers. Mid-Carboniferous, a drop in sea level precipitated a major extinction, one that hit crinoids. This sea level drop and the unconformity in North America separate the Mississippian subperiod from the Pennsylvanian subperiod. This happened about 323 million years ago, at the onset of the Permo-Carboniferous Glaciation, the Carboniferous was a time of active mountain-building, as the supercontinent Pangaea came together. The southern continents remained tied together in the supercontinent Gondwana, which collided with North America–Europe along the present line of eastern North America, in the same time frame, much of present eastern Eurasian plate welded itself to Europe along the line of the Ural mountains. Most of the Mesozoic supercontinent of Pangea was now assembled, although North China, the Late Carboniferous Pangaea was shaped like an O. There were two major oceans in the Carboniferous—Panthalassa and Paleo-Tethys, which was inside the O in the Carboniferous Pangaea, other minor oceans were shrinking and eventually closed - Rheic Ocean, the small, shallow Ural Ocean and Proto-Tethys Ocean
7.
Permian
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The Permian is a geologic period and system which spans 46.7 million years from the end of the Carboniferous Period 298.9 million years ago, to the beginning of the Triassic Period 252.2 Mya. It is the last period of the Paleozoic Era, the following Triassic Period belongs to the Mesozoic Era, the concept of the Permian was introduced in 1841 by geologist Sir Roderick Murchison, who named it after the city of Perm. The Permian witnessed the diversification of the early amniotes into the groups of the mammals, turtles, lepidosaurs. The world at the time was dominated by two known as Pangaea and Siberia, surrounded by a global ocean called Panthalassa. The Carboniferous rainforest collapse left behind vast regions of desert within the continental interior, amniotes, who could better cope with these drier conditions, rose to dominance in place of their amphibian ancestors. The Permian ended with the Permian–Triassic extinction event, the largest mass extinction in Earths history, in which nearly 90% of marine species and it would take well into the Triassic for life to recover from this catastrophe. Recovery from the Permian-Triassic extinction event was protracted, on land, the term Permian was introduced into geology in 1841 by Sir R. I. Murchison, president of the Geological Society of London, who identified typical strata in extensive Russian explorations undertaken with Edouard de Verneuil, the region now lies in the Perm Krai of Russia. This could have in part caused the extinctions of marine species at the end of the period by severely reducing shallow coastal areas preferred by many marine organisms. During the Permian, all the Earths major landmasses were collected into a supercontinent known as Pangaea. The Cimmeria continent rifted away from Gondwana and drifted north to Laurasia, a new ocean was growing on its southern end, the Tethys Ocean, an ocean that would dominate much of the Mesozoic Era. Large continental landmass interiors experience climates with extreme variations of heat and cold, deserts seem to have been widespread on Pangaea. Such dry conditions favored gymnosperms, plants with seeds enclosed in a cover, over plants such as ferns that disperse spores in a wetter environment. The first modern trees appeared in the Permian, the climate in the Permian was quite varied. At the start of the Permian, the Earth was still in an Ice Age, glaciers receded around the mid-Permian period as the climate gradually warmed, drying the continents interiors. In the late Permian period, the drying continued although the temperature cycled between warm and cool cycles, Permian marine deposits are rich in fossil mollusks, echinoderms, and brachiopods. By the close of the Permian, trilobites and a host of other groups became extinct. Terrestrial life in the Permian included diverse plants, fungi, arthropods, the period saw a massive desert covering the interior of Pangaea
8.
Triassic
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The Triassic is a geologic period and system which spans 50.9 million years from the end of the Permian Period 252.17 million years ago, to the beginning of the Jurassic Period 201.3 Mya. The Triassic is the first period of the Mesozoic Era, both the start and end of the period are marked by major extinction events. Therapsids and archosaurs were the terrestrial vertebrates during this time. A specialized subgroup of archosaurs, called dinosaurs, first appeared in the Late Triassic, the vast supercontinent of Pangaea existed until the mid-Triassic, after which it began to gradually rift into two separate landmasses, Laurasia to the north and Gondwana to the south. The global climate during the Triassic was mostly hot and dry, however, the climate shifted and became more humid as Pangaea began to drift apart. The end of the period was marked by yet another mass extinction, the Triassic-Jurassic extinction event. The Triassic is usually separated into Early, Middle, and Late Triassic Epochs, from the east, along the equator, the Tethys sea penetrated Pangaea, causing the Paleo-Tethys Ocean to be closed. Later in the mid-Triassic a similar sea penetrated along the equator from the west, the remaining shores were surrounded by the world-ocean known as Panthalassa. All the deep-ocean sediments laid down during the Triassic have disappeared through subduction of oceanic plates, thus, the supercontinent Pangaea was rifting during the Triassic—especially late in that period—but had not yet separated. In North America, for example, marine deposits are limited to a few exposures in the west, thus Triassic stratigraphy is mostly based on organisms that lived in lagoons and hypersaline environments, such as Estheria crustaceans. At the beginning of the Mesozoic Era, Africa was joined with Earths other continents in Pangaea, Africa shared the supercontinents relatively uniform fauna which was dominated by theropods, prosauropods and primitive ornithischians by the close of the Triassic period. Late Triassic fossils are found throughout Africa, but are common in the south than north. The time boundary separating the Permian and Triassic marks the advent of an event with global impact. At Paleorrota geopark, located in Rio Grande do Sul, Brazil, in these formations, one of the earliest dinosaurs, Staurikosaurus, as well as the mammal ancestors Brasilitherium and Brasilodon have been discovered. The Triassic continental interior climate was hot and dry, so that typical deposits are red bed sandstones and evaporites. Pangaeas large size limited the effect of the global ocean, its continental climate was highly seasonal, with very hot summers. The strong contrast between the Pangea supercontinent and the global ocean triggered intense cross-equatorial monsoons, the best studied of such episodes of humid climate, and probably the most intense and widespread, was the Carnian Pluvial Event. On land, the vascular plants included the lycophytes, the dominant cycadophytes, ginkgophyta, ferns, horsetails
9.
Jurassic
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The Jurassic is a geologic period and system that spans 56.3 million years from the end of the Triassic Period 201.3 million years ago to the beginning of the Cretaceous Period 145 Mya. The Jurassic constitutes the middle period of the Mesozoic Era, also known as the Age of Reptiles, the start of the period is marked by the major Triassic–Jurassic extinction event. The Jurassic is named after the Jura Mountains within the European Alps, by the beginning of the Jurassic, the supercontinent Pangaea had begun rifting into two landmasses, Laurasia to the north and Gondwana to the south. This created more coastlines and shifted the continental climate from dry to humid, on land, the fauna transitioned from the Triassic fauna, dominated by both dinosauromorph and crocodylomorph archosaurs, to one dominated by dinosaurs alone. The first birds also appeared during the Jurassic, having evolved from a branch of theropod dinosaurs, other major events include the appearance of the earliest lizards, and the evolution of therian mammals, including primitive placentals. Crocodilians made the transition from a terrestrial to a mode of life. The oceans were inhabited by marine reptiles such as ichthyosaurs and plesiosaurs, the chronostratigraphic term Jurassic is directly linked to the Jura Mountains. The name Jura is derived from the Celtic root jor, which was Latinised into juria, the Jurassic period is divided into the Early Jurassic, Middle, and Late Jurassic epochs. The Jurassic System, in stratigraphy, is divided into the Lower Jurassic, Middle, the separation of the term Jurassic into three sections goes back to Leopold von Buch. The Jurassic North Atlantic Ocean was relatively narrow, while the South Atlantic did not open until the following Cretaceous period, the Tethys Sea closed, and the Neotethys basin appeared. Climates were warm, with no evidence of glaciation, as in the Triassic, there was apparently no land over either pole, and no extensive ice caps existed. In contrast, the North American Jurassic record is the poorest of the Mesozoic, the Jurassic was a time of calcite sea geochemistry in which low-magnesium calcite was the primary inorganic marine precipitate of calcium carbonate. Carbonate hardgrounds were thus very common, along with calcitic ooids, calcitic cements, the first of several massive batholiths were emplaced in the northern American cordillera beginning in the mid-Jurassic, marking the Nevadan orogeny. Important Jurassic exposures are found in Russia, India, South America, Japan, Australasia. As the Jurassic proceeded, larger and more groups of dinosaurs like sauropods and ornithopods proliferated in Africa. Middle Jurassic strata are well represented nor well studied in Africa. Late Jurassic strata are also poorly represented apart from the spectacular Tendaguru fauna in Tanzania, the Late Jurassic life of Tendaguru is very similar to that found in western North Americas Morrison Formation. During the Jurassic period, the primary living in the sea were fish
10.
Cretaceous
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The Cretaceous is a geologic period and system that spans 79 million years from the end of the Jurassic Period 145 million years ago to the beginning of the Paleogene Period 66 Mya. It is the last period of the Mesozoic Era, the Cretaceous Period is usually abbreviated K, for its German translation Kreide. The Cretaceous was a period with a warm climate, resulting in high eustatic sea levels that created numerous shallow inland seas. These oceans and seas were populated with now-extinct marine reptiles, ammonites and rudists, during this time, new groups of mammals and birds, as well as flowering plants, appeared. The Cretaceous ended with a mass extinction, the Cretaceous–Paleogene extinction event, in which many groups, including non-avian dinosaurs, pterosaurs. The end of the Cretaceous is defined by the abrupt Cretaceous–Paleogene boundary, the name Cretaceous was derived from Latin creta, meaning chalk. The Cretaceous is divided into Early and Late Cretaceous epochs, or Lower and Upper Cretaceous series, in older literature the Cretaceous is sometimes divided into three series, Neocomian, Gallic and Senonian. A subdivision in eleven stages, all originating from European stratigraphy, is now used worldwide, in many parts of the world, alternative local subdivisions are still in use. As with other geologic periods, the rock beds of the Cretaceous are well identified. No great extinction or burst of diversity separates the Cretaceous from the Jurassic and this layer has been dated at 66.043 Ma. A140 Ma age for the Jurassic-Cretaceous boundary instead of the usually accepted 145 Ma was proposed in 2014 based on a study of Vaca Muerta Formation in Neuquén Basin. Víctor Ramos, one of the authors of the study proposing the 140 Ma boundary age sees the study as a first step toward formally changing the age in the International Union of Geological Sciences, due to the high sea level there was extensive space for such sedimentation. Because of the young age and great thickness of the system. Chalk is a type characteristic for the Cretaceous. It consists of coccoliths, microscopically small calcite skeletons of coccolithophores, the group is found in England, northern France, the low countries, northern Germany, Denmark and in the subsurface of the southern part of the North Sea. Chalk is not easily consolidated and the Chalk Group still consists of sediments in many places. The group also has other limestones and arenites, among the fossils it contains are sea urchins, belemnites, ammonites and sea reptiles such as Mosasaurus. In southern Europe, the Cretaceous is usually a marine system consisting of competent limestone beds or incompetent marls
11.
Paleogene
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The Paleogene is a geologic period and system that spans 43 million years from the end of the Cretaceous Period 66 million years ago to the beginning of the Neogene Period 23.03 Mya. It is the beginning of the Cenozoic Era of the present Phanerozoic Eon and this period consists of the Paleocene, Eocene and Oligocene epochs. The terms Paleogene System and lower Tertiary System are applied to the rocks deposited during the Paleogene Period. By dividing the Tertiary Period into two periods instead of directly into five epochs, the periods are more comparable to the duration of periods of the preceding Mesozoic and Paleozoic Eras. The trend was caused by the formation of the Antarctic Circumpolar Current. During the Paleogene, the continued to drift closer to their current positions. India was in the process of colliding with Asia, subsequently forming the Himalayas, the Atlantic Ocean continued to widen by a few centimeters each year. Africa was moving north to meet with Europe and form the Mediterranean, inland seas retreated from North America early in the period. Australia had also separated from Antarctica and was drifting towards Southeast Asia, mammals began a rapid diversification during this period. Some of these mammals would evolve into forms that would dominate the land, while others would become capable of living in marine, specialized terrestrial. Those that took to the oceans became modern cetaceans, while those that took to the trees became primates, the group to which humans belong. Birds, which were well established by the end of the Cretaceous. In comparison to birds and mammals, most other branches of life remained unchanged during this period. As the Earth cooled, tropical plants became less numerous and were now restricted to equatorial regions, deciduous plants, which could survive through the seasonal climates the world was now experiencing, became more common. The Paleogene is notable in the context of offshore oil drilling, and especially in Gulf of Mexico oil exploration and these rock formations represent the current cutting edge of deep-water oil discovery. Lower Tertiary explorations to date include, Kaskida Oil Field Tiber Oil Field Jack 2 Paleogene Microfossils, 180+ images of Foraminifera
12.
Neogene
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The Neogene is a geologic period and system that spans 20.45 million years from the end of the Paleogene Period 23.03 million years ago to the beginning of the present Quaternary Period 2.58 Mya. The Neogene is sub-divided into two epochs, the earlier Miocene and the later Pliocene, some geologists assert that the Neogene cannot be clearly delineated from the modern geological period, the Quaternary. During this period, mammals and birds continued to evolve into modern forms. Early hominids, the ancestors of humans, appeared in Africa near the end of the period, some continental movement took place, the most significant event being the connection of North and South America at the Isthmus of Panama, late in the Pliocene. This cut off the ocean currents from the Pacific to the Atlantic ocean. The global climate cooled considerably over the course of the Neogene, the terms Neogene System and upper Tertiary System describe the rocks deposited during the Neogene Period. The continents in the Neogene were very close to their current positions, the Isthmus of Panama formed, connecting North and South America. The Indian subcontinent continued to collide with Asia, forming the Himalayas, sea levels fell, creating land bridges between Africa and Eurasia and between Eurasia and North America. The global climate became seasonal and continued an overall drying and cooling trend which began at the start of the Paleogene. The ice caps on both poles began to grow and thicken, and by the end of the period the first of a series of glaciations of the current Ice Age began, marine and continental flora and fauna have a modern appearance. The reptile group Choristodera became extinct in the part of the period. Mammals and birds continued to be the dominant terrestrial vertebrates, the first hominids, the ancestors of humans, appeared in Africa and spread into Eurasia. In response to the cooler, seasonal climate, tropical plant species gave way to deciduous ones, grasses therefore greatly diversified, and herbivorous mammals evolved alongside it, creating the many grazing animals of today such as horses, antelope, and bison. The Neogene traditionally ended at the end of the Pliocene Epoch, just before the definition of the beginning of the Quaternary Period. However, there was a movement amongst geologists to also include ongoing geological time in the Neogene, by dividing the Cenozoic Era into three periods instead of seven epochs, the periods are more closely comparable to the duration of periods in the Mesozoic and Paleozoic eras. The International Commission on Stratigraphy once proposed that the Quaternary be considered a sub-era of the Neogene, with a date of 2.58 Ma. In the 2004 proposal of the ICS, the Neogene would have consisted of the Miocene and Pliocene epochs, thus the Neogene Period ends bounding the succeeding Quaternary Period at 2.58 Mya. Digital Atlas of Neogene Life for the Southeastern United States — by San Jose State University via Web Archive
13.
Taxonomy (biology)
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Taxonomy is the science of defining groups of biological organisms on the basis of shared characteristics and giving names to those groups. The exact definition of taxonomy varies from source to source, but the core of the remains, the conception, naming. There is some disagreement as to whether biological nomenclature is considered a part of taxonomy, the broadest meaning of taxonomy is used here. The word taxonomy was introduced in 1813 by Candolle, in his Théorie élémentaire de la botanique, the term alpha taxonomy is primarily used today to refer to the discipline of finding, describing, and naming taxa, particularly species. In earlier literature, the term had a different meaning, referring to morphological taxonomy, ideals can, it may be said, never be completely realized. They have, however, a value of acting as permanent stimulants. Some of us please ourselves by thinking we are now groping in a beta taxonomy, turrill thus explicitly excludes from alpha taxonomy various areas of study that he includes within taxonomy as a whole, such as ecology, physiology, genetics, and cytology. He further excludes phylogenetic reconstruction from alpha taxonomy, thus, Ernst Mayr in 1968 defined beta taxonomy as the classification of ranks higher than species. This activity is what the term denotes, it is also referred to as beta taxonomy. How species should be defined in a group of organisms gives rise to practical and theoretical problems that are referred to as the species problem. The scientific work of deciding how to define species has been called microtaxonomy, by extension, macrotaxonomy is the study of groups at higher taxonomic ranks, from subgenus and above only, than species. While some descriptions of taxonomic history attempt to date taxonomy to ancient civilizations, earlier works were primarily descriptive, and focused on plants that were useful in agriculture or medicine. There are a number of stages in scientific thinking. Early taxonomy was based on criteria, the so-called artificial systems. Later came systems based on a complete consideration of the characteristics of taxa, referred to as natural systems, such as those of de Jussieu, de Candolle and Bentham. The publication of Charles Darwins Origin of Species led to new ways of thinking about classification based on evolutionary relationships and this was the concept of phyletic systems, from 1883 onwards. This approach was typified by those of Eichler and Engler, the advent of molecular genetics and statistical methodology allowed the creation of the modern era of phylogenetic systems based on cladistics, rather than morphology alone. Taxonomy has been called the worlds oldest profession, and naming and classifying our surroundings has likely been taking place as long as mankind has been able to communicate
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Animal
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Animals are multicellular, eukaryotic organisms of the kingdom Animalia. The animal kingdom emerged as a clade within Apoikozoa as the group to the choanoflagellates. Animals are motile, meaning they can move spontaneously and independently at some point in their lives and their body plan eventually becomes fixed as they develop, although some undergo a process of metamorphosis later in their lives. All animals are heterotrophs, they must ingest other organisms or their products for sustenance, most known animal phyla appeared in the fossil record as marine species during the Cambrian explosion, about 542 million years ago. Animals can be divided broadly into vertebrates and invertebrates, vertebrates have a backbone or spine, and amount to less than five percent of all described animal species. They include fish, amphibians, reptiles, birds and mammals, the remaining animals are the invertebrates, which lack a backbone. These include molluscs, arthropods, annelids, nematodes, flatworms, cnidarians, ctenophores, the study of animals is called zoology. The word animal comes from the Latin animalis, meaning having breath, the biological definition of the word refers to all members of the kingdom Animalia, encompassing creatures as diverse as sponges, jellyfish, insects, and humans. Aristotle divided the world between animals and plants, and this was followed by Carl Linnaeus, in the first hierarchical classification. In Linnaeuss original scheme, the animals were one of three kingdoms, divided into the classes of Vermes, Insecta, Pisces, Amphibia, Aves, and Mammalia. Since then the last four have all been subsumed into a single phylum, in 1874, Ernst Haeckel divided the animal kingdom into two subkingdoms, Metazoa and Protozoa. The protozoa were later moved to the kingdom Protista, leaving only the metazoa, thus Metazoa is now considered a synonym of Animalia. Animals have several characteristics that set apart from other living things. Animals are eukaryotic and multicellular, which separates them from bacteria and they are heterotrophic, generally digesting food in an internal chamber, which separates them from plants and algae. They are also distinguished from plants, algae, and fungi by lacking cell walls. All animals are motile, if only at life stages. In most animals, embryos pass through a stage, which is a characteristic exclusive to animals. With a few exceptions, most notably the sponges and Placozoa and these include muscles, which are able to contract and control locomotion, and nerve tissues, which send and process signals
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Arthropod
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An arthropod is an invertebrate animal having an exoskeleton, a segmented body, and jointed appendages. Arthropods form the phylum Arthropoda, which includes the insects, arachnids, myriapods, arthropods are characterized by their jointed limbs and cuticle made of chitin, often mineralised with calcium carbonate. The arthropod body plan consists of segments, each with a pair of appendages, the rigid cuticle inhibits growth, so arthropods replace it periodically by moulting. Their versatility has enabled them to become the most species-rich members of all guilds in most environments. They have over a million described species, making up more than 80% of all described living species, some of which. Arthropods range in size from the microscopic crustacean Stygotantulus up to the Japanese spider crab, arthropods primary internal cavity is a hemocoel, which accommodates their internal organs, and through which their haemolymph – analogue of blood – circulates, they have open circulatory systems. Like their exteriors, the organs of arthropods are generally built of repeated segments. Their nervous system is ladder-like, with paired ventral nerve cords running through all segments and their heads are formed by fusion of varying numbers of segments, and their brains are formed by fusion of the ganglia of these segments and encircle the esophagus. The respiratory and excretory systems of arthropods vary, depending as much on their environment as on the subphylum to which they belong, arthropods also have a wide range of chemical and mechanical sensors, mostly based on modifications of the many setae that project through their cuticles. Aquatic species use internal or external fertilization. Almost all arthropods lay eggs, but scorpions give birth to live young after the eggs have hatched inside the mother, arthropod hatchlings vary from miniature adults to grubs and caterpillars that lack jointed limbs and eventually undergo a total metamorphosis to produce the adult form. The level of care for hatchlings varies from nonexistent to the prolonged care provided by scorpions. The evolutionary ancestry of arthropods dates back to the Cambrian period, the group is generally regarded as monophyletic, and many analyses support the placement of arthropods with cycloneuralians in a superphylum Ecdysozoa. Overall however, the relationships of Metazoa are not yet well resolved. Likewise, the relationships between various groups are still actively debated. Arthropods contribute to the food supply both directly as food, and more importantly indirectly as pollinators of crops. Some species are known to spread disease to humans, livestock. The word arthropod comes from the Greek ἄρθρον árthron, joint, and πούς pous, i. e. foot or leg, arthropods are invertebrates with segmented bodies and jointed limbs
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Crustacean
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Crustaceans form a large, diverse arthropod taxon which includes such familiar animals as crabs, lobsters, crayfish, shrimp, krill, woodlice and barnacles. In other words, some crustaceans are more related to insects. The 67,000 described species range in size from Stygotantulus stocki at 0.1 mm, to the Japanese spider crab with a leg span of up to 3.8 m, like other arthropods, crustaceans have an exoskeleton, which they moult to grow. Most crustaceans are free-living aquatic animals, but some are terrestrial, some are parasitic, the group has an extensive fossil record, reaching back to the Cambrian, and includes living fossils such as Triops cancriformis, which has existed apparently unchanged since the Triassic period. More than 10 million tons of crustaceans are produced by fishery or farming for human consumption, krill and copepods are not as widely fished, but may be the animals with the greatest biomass on the planet, and form a vital part of the food chain. The scientific study of crustaceans is known as carcinology, and a scientist who works in carcinology is a carcinologist, the body of a crustacean is composed of segments, which are grouped into three regions, the cephalon or head, the thorax, and the pleon or abdomen. The head and thorax may be fused together to form a cephalothorax, the crustacean body is protected by the hard exoskeleton, which must be moulted for the animal to grow. The shell around each somite can be divided into a dorsal tergum, ventral sternum, various parts of the exoskeleton may be fused together. The abdomen bears pleopods, and ends in a telson, which bears the anus, the number and variety of appendages in different crustaceans may be partly responsible for the groups success. It is unclear whether the condition is a derived state which evolved in crustaceans. Trilobites, for instance, also possessed biramous appendages, the main body cavity is an open circulatory system, where blood is pumped into the haemocoel by a heart located near the dorsum. Malacostraca have haemocyanin as the pigment, while copepods, ostracods, barnacles. The alimentary canal consists of a tube that often has a gizzard-like gastric mill for grinding food and a pair of digestive glands that absorb food. Structures that function as kidneys are located near the antennae, a brain exists in the form of ganglia close to the antennae, and a collection of major ganglia is found below the gut. In many decapods, the first pair of pleopods are specialised in the male for sperm transfer, many terrestrial crustaceans mate seasonally and return to the sea to release the eggs. Others, such as woodlice, lay their eggs on land, in most decapods, the females retain the eggs until they hatch into free-swimming larvae. Marine crustaceans are as ubiquitous in the oceans as insects are on land, some branchiurans are able to withstand rapid changes of salinity and will also switch hosts from marine to non-marine species. Krill are the layer and the most important part of the food chain in Antarctic animal communities
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Malacostraca
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Malacostraca is the largest of the six classes of crustaceans, containing about 40,000 living species, divided among 16 orders. They are abundant in all environments and have colonised freshwater. They are segmented animals, united by a body plan comprising 20 body segments, and divided into a head, thorax. The name Malacostraca was coined by a French zoologist Pierre André Latreille in 1802 and he was curator of the arthropod collection at the National Museum of Natural History in Paris. The name comes from the Greek roots μαλακός and ὄστρακον, the name is misleading, since the shell is only soft immediately after moulting, and is usually hard. Malacostracans are sometimes contrasted with entomostracans, a name applied to all crustaceans outside the Malacostraca, the class Malacostraca includes about 40,000 species, and arguably. Contains a greater diversity of forms than any other class in the animal kingdom. Malacostracans have abdominal appendages, a fact that differentiates them from all other major crustacean taxa except Remipedia, each body segment bears a pair of jointed appendages, although these may be lost secondarily. The head bears two pairs of antennae, the first of which is often biramous and the second pair bear exopods which are flattened into antennal scales known as scaphocerites. The mouthparts consist of each of mandibles, maxillules and maxillae. Usually a pair of stalked compound eyes is present, although in some taxa the eyes are unstalked, reduced or lost, up to three thoracic segments may be fused with the head to form a cephalothorax, the associated appendages turn forward and are modified as maxillipeds. A carapace may be absent, present or secondarily lost, and may cover the head, part or all of the thorax and it is variable in form and may be fused dorsally with some of the thoracic segments or occasionally be in two parts, hinged dorsally. Typically, each of the thoracic appendages is biramous and the endopods are the developed of the branches. Each endopod consist of seven articulating segments, the coxa, basis, ischium, in decapods, the claw is formed by the articulation of the dactylus against an outgrowth of the propodus. In some taxa, the exopods are lost and the appendages are uniramous, there is a clear demarcation between the thorax and the six or seven-segmented abdomen. In most taxa, each abdominal segment except the last carries a pair of biramous pleopods used for swimming, burrowing, gas exchange, the first and second abdominal pleopods may be modified in the male to form gonopods. The appendages of the last segment are typically flattened into uropods and it is the sudden flexion of this tail fan that provides the thrust for the rapid escape response of these crustaceans and the tail fan is also used in steering. In Leptostraca, the appendages on the telson instead form caudal rami, the digestive tract is straight and the foregut consists of a short oesophagus and a two-chambered stomach, the first part of which contains a gizzard-like gastric mill for grinding food
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Decapoda
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The Decapoda or decapods are an order of crustaceans within the class Malacostraca, including many familiar groups, such as crayfish, crabs, lobsters, prawns, and shrimp. The order is estimated to contain nearly 15,000 species in around 2,700 genera, nearly half of these species are crabs, with the shrimp and Anomura (including hermit crabs, porcelain crabs, squat lobsters making up the bulk of the remainder. The earliest fossil decapod is the Devonian Palaeopalaemon, as the name Decapoda implies, all decapods have what are considered ten legs, in the form of five pairs of thoracic appendages on the last five thoracic segments. Decapods can have as many as twenty apendages, arranged in one pair per body segment, the front three pairs function as mouthparts and are generally referred to as maxillipeds, the remainder are pereiopods. In many decapods, however, one pair of legs has enlarged pincers, the remaining appendages are found on the abdomen, with each segment capable of carrying a pair of biramous pleopods, the last of which form part of the tail fan and are called uropods. Classification within the order Decapoda depends on the structure of the gills and legs, the Dendrobranchiata consist of prawns, including many species colloquially referred to as shrimp, such as the white shrimp, Litopenaeus setiferus. The Pleocyemata include the groups, including true shrimp. Those groups which usually walk rather than form a clade called Reptantia. This classification to the level of superfamilies follows De Grave et al
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Pleocyemata
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Pleocyemata is a sub-order of decapod crustaceans, erected by Martin Burkenroad in 1963. Burkenroads classification replaced the earlier sub-orders of Natantia and Reptantia with the monophyletic groups Dendrobranchiata and Pleocyemata, Pleocyemata contains all the members of the Reptantia, as well as the Stenopodidea, and Caridea, which contains the true shrimp. It is this characteristic that gives the group its name, Pleocyemata also possess a lamellar gill structure as opposed to the branched found in Dendrobranchiata. The earliest fossil representative is the Devonian Palaeopalaemon, data related to Pleocyemata at Wikispecies
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Carl Linnaeus
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Carl Linnaeus, also known after his ennoblement as Carl von Linné, was a Swedish botanist, physician, and zoologist, who formalised the modern system of naming organisms called binomial nomenclature. He is known by the father of modern taxonomy. Many of his writings were in Latin, and his name is rendered in Latin as Carolus Linnæus, Linnaeus was born in the countryside of Småland, in southern Sweden. He received most of his education at Uppsala University. He lived abroad between 1735 and 1738, where he studied and also published a first edition of his Systema Naturae in the Netherlands and he then returned to Sweden, where he became professor of medicine and botany at Uppsala. In the 1740s, he was sent on journeys through Sweden to find and classify plants. In the 1750s and 1760s, he continued to collect and classify animals, plants, and minerals, at the time of his death, he was one of the most acclaimed scientists in Europe. The philosopher Jean-Jacques Rousseau sent him the message, Tell him I know no man on earth. The German writer Johann Wolfgang von Goethe wrote, With the exception of Shakespeare and Spinoza, Swedish author August Strindberg wrote, Linnaeus was in reality a poet who happened to become a naturalist. Among other compliments, Linnaeus has been called Princeps botanicorum, The Pliny of the North and he is also considered as one of the founders of modern ecology. In botany, the abbreviation used to indicate Linnaeus as the authority for species names is L. In older publications, sometimes the abbreviation Linn. is found, Linnæus was born in the village of Råshult in Småland, Sweden, on 23 May 1707. He was the first child of Nicolaus Ingemarsson and Christina Brodersonia and his siblings were Anna Maria Linnæa, Sofia Juliana Linnæa, Samuel Linnæus, and Emerentia Linnæa. One of a line of peasants and priests, Nils was an amateur botanist, a Lutheran minister. Christina was the daughter of the rector of Stenbrohult, Samuel Brodersonius, a year after Linnæus birth, his grandfather Samuel Brodersonius died, and his father Nils became the rector of Stenbrohult. The family moved into the rectory from the curates house, even in his early years, Linnæus seemed to have a liking for plants, flowers in particular. Whenever he was upset, he was given a flower, which calmed him. Nils spent much time in his garden and often showed flowers to Linnaeus, soon Linnæus was given his own patch of earth where he could grow plants
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10th edition of Systema Naturae
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The 10th edition of Systema Naturae is a book written by Carl Linnaeus and published in two volumes in 1758 and 1759, which marks the starting point of zoological nomenclature. In it, Linnaeus introduced binomial nomenclature for animals, something he had already done for plants in his 1753 publication of Species Plantarum, before 1758, most biological catalogues had used polynomial names for the taxa included, including earlier editions of Systema Naturae. The first work to consistently apply binomial nomenclature across the kingdom was the 10th edition of Systema Naturae. Names published before that date are unavailable, even if they would otherwise satisfy the rules, during Linnaeus lifetime, Systema Naturae was under continuous revision. The Animal Kingdom, Animals enjoy sensation by means of an organization, animated by a medullary substance, perception by nerves. They have members for the different purposes of life, organs for their different senses and they all originate from an egg. Their external and internal structure, their anatomy, habits, instincts. The list has been broken down into the six classes Linnaeus described for animals, Mammalia, Aves, Amphibia, Pisces, Insecta. These classes were created by studying the internal anatomy, as seen in his key. Warm, red blood Viviparous, Mammalia Oviparous, Aves Heart with 1 auricle,1 ventricle, cold, red blood Lungs voluntary, Amphibia External gills, Pisces Heart with 1 auricle,0 ventricles. Linnaeus described mammals as, Animals that suckle their young by means of lactiferous teats, in external and internal structure they resemble man, most of them are quadrupeds, and with man, their natural enemy, inhabit the surface of the Earth. The largest, though fewest in number, inhabit the ocean and they are areal, vocal, swift and light, and destitute of external ears, lips, teeth, scrotum, womb, bladder, epiglottis, corpus callosum and its arch, and diaphragm. They breathe by means of gills, which are united by a bony arch, swim by means of radiate fins. Many of them are without a head, and most of them without feet. They are principally distinguished by their tentacles, by the Ancients they were not improperly called imperfect animals, as being destitute of ears, nose, head, eyes and legs, and are therefore totally distinct from Insects. In addition to repeating the species he had listed in his Species Plantarum. The species from Species Plantarum were numbered sequentially, while the new species were labelled with letters, new plant species described in the 10th edition of Systema Naturae include, The original 1758 Systema Naturae Linnaeus 1758 Classification of Animals on the Taxonomicon
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Dromiacea
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Dromiacea is a group of crabs, ranked as a section. It contains 240 extant and nearly 300 extinct species, the larvae of Dromiacea resemble those of the Anomura more closely than those of other crabs. This may simply reflect their position in the crab phylogeny. The fossil record of Dromiacea reaches back at least as far as the Jurassic, the superfamily Eocarcinoidea, containing Eocarcinus and Platykotta, was previously considered to be a member of the Dromiacea, but has since been transferred to the Anomura
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Eubrachyura
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Eubrachyura is a group of crustaceans comprising the more advanced crabs. It is divided into two subsections, based on the position of the openings in the two sexes. In the Heterotremata, the openings are on the legs in the males, but on the sternum in females, while in the Thoracotremata and this contrasts with the situation in other decapods, in which the genital openings are always on the legs. Heterotremata is the larger of the two groups, containing the species rich superfamilies Xanthoidea and Pilumnoidea and all the freshwater crabs. The fossil record of Eubrachyura extends back to the Cretaceous, the supposed Bathonian representative of the group, Hebertides jurassica, data related to Eubrachyura at Wikispecies
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Infraorder
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In biological classification, the order is a taxonomic rank used in the classification of organisms and recognized by the nomenclature codes. Other well-known ranks are life, domain, kingdom, phylum, class, family, genus, an immediately higher rank, superorder, may be added directly above order, while suborder would be a lower rank. A taxonomic unit, a taxon, in that rank, in that case the plural is orders. Example, The Juglans and Hickory trees belong to the family Juglandaceae, what does and does not belong to each order is determined by a taxonomist, as is whether a particular order should be recognized at all. Often there is no agreement, with different taxonomists each taking a different position. There are no rules that a taxonomist needs to follow in describing or recognizing an order. Some taxa are accepted almost universally, while others are recognised only rarely, for some groups of organisms, consistent suffixes are used to denote that the rank is an order. The Latin suffix -formes meaning having the form of is used for the name of orders of birds and fishes. The suffix -ales is for the name of orders of plants, fungi, for some clades covered by the International Code of Zoological Nomenclature, a number of additional classifications are sometimes used, although not all of these are officially recognised. In their 1997 classification of mammals, McKenna and Bell used two levels between superorder and order, grandorder and mirorder. Michael Novacek inserted them at the same position, michael Benton inserted them between superorder and magnorder instead. This position was adopted by Systema Naturae 2000 and others, in botany, the ranks of subclass and suborder are secondary ranks pre-defined as respectively above and below the rank of order. Any number of further ranks can be used as long as they are clearly defined, the superorder rank is commonly used, with the ending -anae that was initiated by Armen Takhtajans publications from 1966 onwards. Carl Linnaeus was the first to apply it consistently to the division of all three kingdoms of nature in his Systema Naturae, for plants, Linnaeus orders in the Systema Naturae and the Species Plantarum were strictly artificial, introduced to subdivide the artificial classes into more comprehensible smaller groups. In French botanical publications, from Michel Adansons Familles naturelles des plantes and until the end of the 19th century, some of the plant families still retain the names of Linnaean natural orders or even the names of pre-Linnaean natural groups recognised by Linnaeus as orders in his natural classification. Such names are known as family names. In zoology, the Linnaean orders were used more consistently and that is, the orders in the zoology part of the Systema Naturae refer to natural groups. Some of his names are still in use
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Tail
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The tail is the section at the rear end of an animals body, in general, the term refers to a distinct, flexible appendage to the torso. It is the part of the body that corresponds roughly to the sacrum and coccyx in mammals, reptiles, tailed objects are sometimes referred to as caudate and the part of the body associated with or proximal to the tail are given the adjective caudal. Animal tails are used in a variety of ways and they provide a source of locomotion for fish and some other forms of marine life. Many land animals use their tails to brush away flies and other biting insects, tails are also used for social signaling. Some species tails are armored, and some, such as those of scorpions, some species of lizard can detach their tails from their bodies. This can help them to escape predators, which are either distracted by the wriggling, tails cast in this manner generally grow back over time, though the replacement is typically darker in colour than the original. Most birds tails end in long feathers called rectrices and these feathers are used as a rudder, helping the bird steer and maneuver in flight, they also help the bird to balance while it is perched. In some species—such as birds of paradise, lyrebirds, and most notably peafowl—modified tail feathers play an important role in courtship displays, the extra-stiff tail feathers of other species, including woodpeckers and woodcreepers, allow them to brace themselves firmly against tree trunks. The tails of grazing animals, such as horses, are used both to sweep away insects and positioned or moved in ways that indicate the physical or emotional state. Human embryos have a tail measures about one-sixth of the size of the embryo itself. As the embryo develops into a fetus, the tail is absorbed by the growing body, humans have a tail bone attached to the pelvis, formed of fused vertebrae, usually four, at the bottom of the vertebral column. Rump Empennage, the tail of an aircraft
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Abdomen
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The abdomen constitutes the part of the body between the thorax and pelvis, in humans and in other vertebrates. The region occupied by the abdomen is termed the abdominal cavity, in arthropods it is the posterior tagma of the body, it follows the thorax or cephalothorax. The abdomen stretches from the thorax at the diaphragm to the pelvis at the pelvic brim. The pelvic brim stretches from the joint to the pubic symphysis and is the edge of the pelvic inlet. The space above this inlet and under the thoracic diaphragm is termed the abdominal cavity, the boundary of the abdominal cavity is the abdominal wall in the front and the peritoneal surface at the rear. The abdomen contains most of the organs of the digestive tract. Hollow abdominal organs include the stomach, the intestine. Organs such as the liver, its attached gallbladder, and the function in close association with the digestive tract. The spleen, kidneys, and adrenal glands also lie within the abdomen, along with blood vessels including the aorta. Anatomists may consider the urinary bladder, uterus, fallopian tubes, finally, the abdomen contains an extensive membrane called the peritoneum. A fold of peritoneum may completely cover certain organs, whereas it may only one side of organs that usually lie closer to the abdominal wall. Anatomists call the type of organs retroperitoneal. For example, the stomach of ruminants is divided into four chambers – rumen, in vertebrates, the abdomen is a large cavity enclosed by the abdominal muscles, ventrally and laterally, and by the vertebral column dorsally. Lower ribs can also enclose ventral and lateral walls, the abdominal cavity is upper part of the pelvic cavity. It is attached to the cavity by the diaphragm. Structures such as the aorta, inferior vena cava and esophagus pass through the diaphragm, both the abdominal and pelvic cavities are lined by a serous membrane known as the parietal peritoneum. This membrane is continuous with the visceral peritoneum lining the organs, the abdomen in vertebrates contains a number of organs belonging, for instance, to the digestive tract and urinary system. There are three layers of the abdominal wall and they are, from the outside to the inside, external oblique, internal oblique, and transverse abdominal
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Greek language
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Greek is an independent branch of the Indo-European family of languages, native to Greece and other parts of the Eastern Mediterranean. It has the longest documented history of any living language, spanning 34 centuries of written records and its writing system has been the Greek alphabet for the major part of its history, other systems, such as Linear B and the Cypriot syllabary, were used previously. The alphabet arose from the Phoenician script and was in turn the basis of the Latin, Cyrillic, Armenian, Coptic, Gothic and many other writing systems. Together with the Latin texts and traditions of the Roman world, during antiquity, Greek was a widely spoken lingua franca in the Mediterranean world and many places beyond. It would eventually become the official parlance of the Byzantine Empire, the language is spoken by at least 13.2 million people today in Greece, Cyprus, Italy, Albania, Turkey, and the Greek diaspora. Greek roots are used to coin new words for other languages, Greek. Greek has been spoken in the Balkan peninsula since around the 3rd millennium BC, the earliest written evidence is a Linear B clay tablet found in Messenia that dates to between 1450 and 1350 BC, making Greek the worlds oldest recorded living language. Among the Indo-European languages, its date of earliest written attestation is matched only by the now extinct Anatolian languages, the Greek language is conventionally divided into the following periods, Proto-Greek, the unrecorded but assumed last ancestor of all known varieties of Greek. The unity of Proto-Greek would have ended as Hellenic migrants entered the Greek peninsula sometime in the Neolithic era or the Bronze Age, Mycenaean Greek, the language of the Mycenaean civilisation. It is recorded in the Linear B script on tablets dating from the 15th century BC onwards, Ancient Greek, in its various dialects, the language of the Archaic and Classical periods of the ancient Greek civilisation. It was widely known throughout the Roman Empire, after the Roman conquest of Greece, an unofficial bilingualism of Greek and Latin was established in the city of Rome and Koine Greek became a first or second language in the Roman Empire. The origin of Christianity can also be traced through Koine Greek, Medieval Greek, also known as Byzantine Greek, the continuation of Koine Greek in Byzantine Greece, up to the demise of the Byzantine Empire in the 15th century. Much of the written Greek that was used as the language of the Byzantine Empire was an eclectic middle-ground variety based on the tradition of written Koine. Modern Greek, Stemming from Medieval Greek, Modern Greek usages can be traced in the Byzantine period and it is the language used by the modern Greeks, and, apart from Standard Modern Greek, there are several dialects of it. In the modern era, the Greek language entered a state of diglossia, the historical unity and continuing identity between the various stages of the Greek language is often emphasised. Greek speakers today still tend to regard literary works of ancient Greek as part of their own rather than a foreign language and it is also often stated that the historical changes have been relatively slight compared with some other languages. According to one estimation, Homeric Greek is probably closer to demotic than 12-century Middle English is to modern spoken English, Greek is spoken by about 13 million people, mainly in Greece, Albania and Cyprus, but also worldwide by the large Greek diaspora. Greek is the language of Greece, where it is spoken by almost the entire population
28.
Thorax
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The thorax or chest is a part of the anatomy of humans and various other animals located between the neck and the abdomen. The thorax includes the thoracic cavity and the thoracic wall and it contains organs including the heart, lungs, and thymus gland, as well as muscles and various other internal structures. Many diseases may affect the chest, and one of the most common symptoms is chest pain, in humans and other hominids, the thorax is the chest region of the body between the neck and the abdomen, along with its internal organs and other contents. It is mostly protected and supported by the rib cage, spine, arteries and veins are also contained –, bones. The area exposed by open-necked shirts, the V of the chest is sometimes the location of a skin disease polymorphous light eruption. In the human body, the region of the thorax between the neck and diaphragm in the front of the body is called the chest, the corresponding area in an animal can also be referred to as the chest. The shape of the chest does not correspond to part of the thoracic skeleton that encloses the heart. All the breadth of the shoulders is due to the girdle, and contains the axillae. Level with this line the second ribs join the sternum, at the lower part of the sternum, where the seventh or last true ribs join it, the ensiform cartilage begins, and above this there is often a depression known as the pit of the stomach. The bones of the thorax, called the thoracic skeleton is a component of the axial skeleton and it consists of the ribs and sternum. The ribs of the thorax are numbered in ascending order from 1-12. 11 &12 are known as floating ribs because they have no anterior attachment point in particular the cartilage attached to the sternum, as 1-7 are, whereas ribs 8-10 are termed false ribs as their costal cartilage articulates with the rib aboves costal cartilage. The anatomy of the chest can also be described through the use of anatomical landmarks, the female nipple is surrounded for half an inch by a more or less pigmented disc, the areola. The apex of a heart is in the fifth left intercostal space. Different types of diseases or conditions that affect the chest include pleurisy, flail chest, atelectasis, and these conditions can be hereditary or caused by birth defects or trauma. Any condition that lowers the ability to breathe deeply or to cough is considered a chest disease or condition. Injury to the chest results in up to ¼ of all due to trauma in the United States. The major pathophysiologies encountered in blunt chest trauma involve derangements in the flow of air, blood, sepsis due to leakage of alimentary tract contents, as in esophageal perforations, also must be considered. Blunt trauma commonly results in chest wall injuries, the pain associated with these injuries can make breathing difficult, and this may compromise ventilation
29.
Hermit crab
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Hermit crabs are decapod crustaceans of the superfamily Paguroidea. Most of the approximately 1100 species possess an asymmetrical abdomen that is concealed in a gastropod shell carried around by the hermit crab. Most species have long, spirally curved abdomens, which are soft, unlike the hard, the vulnerable abdomen is protected from predators by a salvaged empty seashell carried by the hermit crab, into which its whole body can retract. Most frequently, hermit crabs use the shells of sea snails, the tip of the hermit crabs abdomen is adapted to clasp strongly onto the columella of the snail shell. Hermit crabs can be divided into two groups, The first group is the hermit crabs. These crabs spend most of their life underwater as aquatic animals, live in varying depths of saltwater from shallow reefs and shorelines to deep sea bottoms, as pets, several marine species of hermit crabs are common in the marine aquarium trade. They are commonly kept in fish tanks. They breathe through gills but they dont have to carry around their water to do so, most can survive briefly out of water as long as their gills are damp. However, this ability is not as developed as it is in land hermit crabs, a few species do not use a mobile home and inhabit immobile structures left by polychaete worms, vermetid gastropods, corals, and sponges. They belong to the family Coenobitidae, other species, such as Coenobita brevimanus, Coenobita rugosus, Coenobita perlatus or Coenobita cavipes, are less common but growing in availability and popularity as pets. As hermit crabs grow, they require larger shells, since suitable intact gastropod shells are sometimes a limited resource, vigorous competition often occurs among hermit crabs for shells. The availability of empty shells at any given place depends on the abundance of gastropods and hermit crabs. An equally important issue is the population of organisms that prey upon gastropods, hermit crabs kept together may fight or kill a competitor to gain access to the shell they favour. However, if the crabs vary significantly in size, the occurrence of fights over empty shells will decrease or remain nonexistent. Hermit crabs with too-small shells cannot grow as fast as those with well-fitting shells, as the hermit crab grows in size, it must find a larger shell and abandon the previous one. This habit of living in a second-hand shell gives rise to the name hermit crab. Several hermit crab species, both terrestrial and marine, have been observed forming a chain to exchange shells. When an individual finds a new empty shell it will leave its own shell
30.
King crab
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King crabs are a superfamily of crab-like decapod crustaceans chiefly found in cold seas. Because of their size and the taste of their meat, many species are widely caught and sold as food. King crabs are thought to be derived from hermit crab-like ancestors. This ancestry is supported by several anatomical peculiarities which are present only in king crabs, although some doubt still exists about this theory, king crabs are the most widely quoted example of carcinisation among the Decapoda. The evidence for this comes from the asymmetry of the king crabs abdomen, which is thought to reflect the asymmetry of hermit crabs. Although formerly classified among the hermit crabs in the superfamily Paguroidea, king crabs are now placed in a separate superfamily and its single species, G. cristatipes was originally placed in the genus Rhinolithodes. In 2011, scientists found Neolithodes yaldwini on the edge of Antarctica, probably as a result of warming, with major impacts on sediment texture, bioturbation. Red and blue king crabs are some of the most important fisheries in Alaska, however populations have fluctuated in the past 25 years, the two species are similar in size, shape and life history. Habitat is the main factor separating the range of blue and red king crabs in the Bering Sea, Matthew, St. Lawrence and the Diomede Islands. Red king crabs have an 11-month brood cycle in their first reproductive year, the red king crab, Paralithodes camtschaticus, is a very large species, sometimes reaching a carapace width of 11 in and a leg span of 6 ft. Its natural range is the Bering Sea around the Kamchatka Peninsula area and it can now also be found in the Barents Sea and the European Arctic, where it was intentionally introduced and is now becoming a pest. The blue king crab, Paralithodes platypus, lives near St. Matthew Island, the Pribilof Islands, and the Diomede Islands, Alaska, blue king crabs from the Pribilof Islands are the largest of all the king crabs, sometimes exceeding 18 lb in weight. Alaskan king crab fishing Deadliest Catch Media related to Lithodidae at Wikimedia Commons Data related to Lithodidae at Wikispecies
31.
Porcelain crab
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Porcelain crabs are decapod crustaceans in the widespread family Porcellanidae, which superficially resemble true crabs. They have flattened bodies as an adaptation for living in rock crevices and they are delicate, readily losing limbs when attacked, and use their large claws for maintaining territories. Porcelain crabs are small, usually with body widths of less than 15 mm and they share the general body plan of a squat lobster, but their bodies are more compact and flattened, an adaptation for living and hiding under rocks. Porcelain crabs are quite fragile animals, and will shed their limbs to escape predators. The lost appendage can grow back over several moults, porcelain crabs have large chelae, which are used for territorial struggles, but not for catching food. The fifth pair of pereiopods is reduced and used for cleaning, porcelain crabs are an example of carcinisation, whereby a noncrab-like animal evolves into an animal that resembles a true crab. Porcelain crabs can be distinguished from true crabs by the apparent number of walking legs, the abdomen of the porcelain crab is long and folded underneath it, free to move. Porcelain crabs live in all the oceans, except the Arctic Ocean. They are common rocks, and can often be found and observed on rocky beaches and shorelines. They feed by combing plankton and other particles from the water using long setae on the mouthparts. In Hong Kong, Petrolisthes japonicus is common
32.
Horseshoe crab
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Horseshoe crabs are marine arthropods of the family Limulidae and order Xiphosura or Xiphosurida. They are invertebrates meaning that they lack a spine, horseshoe crabs live primarily in and around shallow ocean waters on soft sandy or muddy bottoms. They occasionally come onto shore to mate and they are commonly used as bait and in fertilizer. In recent years, a decline in the population has occurred as a consequence of coastal destruction in Japan. Tetrodotoxin may be present in the roe of species inhabiting the waters of Thailand, because of their origin 450 million years ago, horseshoe crabs are considered living fossils. They belong to a separate subphylum, Chelicerata, and are related to arachnids. The earliest horseshoe crab fossils are found in strata from the late Ordovician period, the latter become functional just before the embryo hatches. Also, a pair of eyes is located near the mouth. The horseshoe crab has 5 additional eyes on top of its shell, the mouth is located in the center of the legs, whose bases are referred to as gnathobases and have the same function as jaws and help grind up food. The horseshoe crab has five pairs of legs for walking, swimming, the long, straight, rigid tail can be used to flip the animal over if turned upside down, so a horseshoe crab with a broken tail is susceptible to desiccation or predation. Behind its legs, the crab has book gills, which exchange respiratory gases. As in other arthropods, a true endoskeleton is absent, and they are more often found on the ocean floor searching for worms and molluscs, which are their main food. They may also feed on crustaceans and even small fish, females are larger than males, C. rotundicauda is the size of a human hand, while L. polyphemus can be up to 60 cm long. The juveniles grow about 33% larger with every molt until reaching adult size, during the breeding season, horseshoe crabs migrate to shallow coastal waters. A male selects a female and clings to her back, often, several males surround the female and all fertilize together which makes it easy to spot and count females as they are the large center carapace surrounded by 3-5 smaller ones. The female digs a hole in the sand and lays her eggs while the male fertilize them, the female can lay between 60,000 and 120,000 eggs in batches of a few thousand at a time. The eggs take about two weeks to hatch, shore birds eat many of them before they hatch, the larvae molt six times during the first year. Raising horseshoe crabs in captivity has proven to be difficult, some evidence indicates mating takes place only in the presence of the sand or mud in which the horseshoe crabs eggs were hatched
33.
Crab louse
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The crab louse is an insect that is an obligate ectoparasite of humans, feeding exclusively on blood. The crab louse usually is found in the pubic hair. Although the louse cannot jump, it can live in other areas of the body that are covered with coarse hair. Humans are the known hosts of the crab louse, although a closely related species, Pthirus gorillae. The human parasite diverged from Pthirus gorillae approximately 3.3 million years ago, and is distantly related to the genus Pediculus. An adult crab louse is about 1. 3–2 mm long, another distinguishing feature is that the rearmost two pairs of legs of a crab louse are much thicker than the front legs and have large claws. The eggs of the crab louse are laid usually on the hairs of the genital and perianal regions of the human body. Crab lice may also be found on areas of the body that have coarse and relatively sparse coverings of hair, such as the beard, moustache, eyelashes. They do not generally occur on the hair of the scalp. The female lays three eggs a day. The eggs take 6–8 days to hatch, and there are three stages which together take 10–17 days before the adult develops, making a total life cycle from egg to adult of 16–25 days. Adults live for up to 30 days, crab lice feed exclusively on blood, and take a blood meal 4–5 times daily. Infestations of crab lice are known as pediculosis pubis or phthiriasis pubis, infestation of the eyelashes is referred to as pediculosis ciliaris or phthiriasis palpebrarum. In some infestations, a characteristic grey-blue or slate coloration appears at the feeding site, crab lice usually infect a new host only by close contact between individuals, usually through sexual intercourse. Adults are more frequently infested than children, non-sexual transmissions may occur among family and roommates through the use of shared towels, clothing, beds or closets. They can only survive a short time away from the warmth and it has been suggested that an increasing percentage of humans removing their pubic hair has led to reduced crab louse populations in some parts of the world. Because the World Health Organization and other authorities do not record statistics for pubic louse infestation, there is virtually no hard data to support this supposition
34.
Gecarcinus quadratus
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G. quadratus is found in mangrove, sand dunes and rainforest along the Pacific coast from Mexico south to Panama. Previously it has also reported from the Pacific coast of northwestern South America. The taxonomy in relation to the Atlantic G. lateralis is disputed, with many considering G. quadratus, the carapace of G. quadratus may reach a length of 5 centimetres. This nocturnal crab digs burrows – sometimes as long as 1.5 metres – in the rainforests of Central America, and is common along the coasts of Costa Rica, Panama. It lives in the forest at least some of its adult life and it is largely herbivorous, and consumes leaf litter and seedlings. Media related to Gecarcinus quadratus at Wikimedia Commons
35.
Terrestrial crab
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A number of lineages of crabs have evolved to live predominantly on land. There is no distinction between terrestrial, semi-terrestrial, and aquatic crabs. Rather, there is a continuum of terrestriality displayed among the true crabs, some species of terrestrial crabs can be found many kilometres from the sea, but have to complete annual migrations to the sea. For example, following the Indian Ocean monsoon, the Christmas Island red crab migrates en masse, the crabs can travel up to 1.46 km in a day, and up to 4 km in total. Terrestrial crabs have evolved from freshwater crabs, since the physiological changes needed for living in fresh water are pre-adaptations for terrestrial living. On some oceanic islands, terrestrial crabs occupy the top of the energy pyramid, freshwater crab Coenobitidae, the terrestrial hermit crabs Coconut crab, a terrestrial hermit crab which is the largest terrestrial invertebrate in the world Land Crabs
36.
Central America
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Central America is the southernmost, isthmian portion of the North American continent, which connects with South America on the southeast. Central America is bordered by Mexico to the north, Colombia to the southeast, the Caribbean Sea to the east, Central America consists of seven countries, Belize, Guatemala, El Salvador, Honduras, Nicaragua, Costa Rica and Panama. The combined population of Central America is between 41,739,000 and 42,688,190, Central America is a part of the Mesoamerican biodiversity hotspot, which extends from northern Guatemala through to central Panama. Due to the presence of several active faults and the Central America Volcanic Arc. Volcanic eruptions and earthquakes occur frequently, these disasters have resulted in the loss of many lives. In the Pre-Columbian era, Central America was inhabited by the peoples of Mesoamerica to the north and west. Soon after Christopher Columbuss voyages to the Americas, the Spanish began to colonize the Americas, the seven states finally became independent autonomous states, beginning with Nicaragua, Honduras, Costa Rica, and Guatemala, followed by El Salvador, then Panama, and finally Belize. Middle America is usually thought to comprise Mexico to the north of the 7 states of Central America as well as Colombia, usually the whole of the Caribbean to the north-east and sometimes the Guyanas are also included. According to one source, the term Central America was used as a synonym for Middle America as recently as 1962, in Brazil, Central America comprises all countries between Mexico and Colombia, including those in the Caribbean. Mexico, in whole or in part, is included by British people. For the people living in the 5 countries formerly part of the Federal Republic of Central America there is a distinction between the Spanish language terms América Central and Centroamérica, in the Pre-Columbian era, the northern areas of Central America were inhabited by the indigenous peoples of Mesoamerica. Most notable among these were the Mayans, who had built numerous cities throughout the region, and the Aztecs, following Christopher Columbuss voyages to the Americas, the Spanish sent many expeditions to the region, and they began their conquest of Maya territory in 1523. Soon after the conquest of the Aztec Empire, Spanish conquistador Pedro de Alvarado commenced the conquest of northern Central America for the Spanish Empire. Beginning with his arrival in Soconusco in 1523, Alvarados forces systematically conquered and subjugated most of the major Maya kingdoms, including the Kiche, Tzutujil, Pipil, and the Kaqchikel. By 1528, the conquest of Guatemala was nearly complete, with only the Petén Basin remaining outside the Spanish sphere of influence, the last independent Maya kingdoms – the Kowoj and the Itza people – were finally defeated in 1697, as part of the Spanish conquest of Petén. In 1538, Spain established the Real Audiencia of Panama, which had jurisdiction over all land from the Strait of Magellan to the Gulf of Fonseca. This entity was dissolved in 1543, and most of the territory within Central America then fell under the jurisdiction of the Audiencia Real de Guatemala. This area included the current territories of Costa Rica, El Salvador, Guatemala, Honduras, Nicaragua, and the Mexican state of Chiapas, the president of the Audiencia, which had its seat in Antigua Guatemala, was the governor of the entire area
37.
Exoskeleton
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An exoskeleton is the external skeleton that supports and protects an animals body, in contrast to the internal skeleton of, for example, a human. In usage, some of the kinds of exoskeletons are known as shells. Examples of animals with exoskeletons include insects such as grasshoppers and cockroaches, the shells of certain sponges and the various groups of shelled molluscs, including those of snails, clams, tusk shells, chitons and nautilus, are also exoskeletons. Some animals, such as the tortoise, have both an endoskeleton and an exoskeleton, exoskeletons have a role in defense from pests and predators, support, and in providing an attachment framework for musculature. Exoskeletons contain chitin, the addition of calcium carbonate makes them harder and stronger, ingrowths of the arthropod exoskeleton known as apodemes serve as attachment sites for muscles. These structures are composed of chitin, and are six times as strong. Similar to tendons, apodemes can stretch to store energy for jumping. Many different species produce exoskeletons, which are composed of a range of materials, bone, cartilage, or dentine is used in the Ostracoderm fish and turtles. Chitin forms the exoskeleton in arthropods including insects, arachnids such as spiders, crustaceans such as crabs and lobsters, calcium carbonates constitute the shells of molluscs, brachiopods, and some tube-building polychaete worms. Silica forms the exoskeleton in the microscopic diatoms and radiolaria, one species of mollusc, the scaly-foot gastropod, even makes use of the iron sulfides greigite and pyrite. Some organisms, such as some foraminifera, agglutinate exoskeletons by sticking grains of sand, contrary to a common misconception, echinoderms do not possess an exoskeleton, as their test is always contained within a layer of living tissue. Exoskeletons have evolved many times,18 lineages evolved calcified exoskeletons alone. Further, other lineages have produced tough outer coatings analogous to an exoskeleton and this coating is constructed from bone in the armadillo, and hair in the pangolin. The armor of reptiles like turtles and Ankylosaurs is constructed of bone, crocodiles have bony scutes, since exoskeletons are rigid, they present some limits to growth. Organisms with open shells can grow by adding new material to the aperture of their shell, as is the case in snails, bivalves, a true exoskeleton, like that found in arthropods, must be shed when it is outgrown. A new exoskeleton is produced beneath the old one, as the old one is shed, the new skeleton is soft and pliable. The animal will pump itself up to expand the new shell to maximal size, when the shell has set, the empty space inside the new skeleton can be filled up as the animal eats. Failure to shed the exoskeleton once outgrown can result in the animal being suffocated within its own shell and this is the mechanism behind some insect pesticides, such as Azadirachtin
38.
Chitin
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Chitin n is a long-chain polymer of an N-acetylglucosamine, a derivative of glucose, and is found in many places throughout the natural world. The structure of chitin is comparable to the cellulose, forming crystalline nanofibrils or whiskers. In terms of function, it may be compared to the protein keratin, chitin has proved versatile for several medicinal, industrial and biotechnological purposes. The English word chitin comes from the French word chitine, which first appeared in 1821 and derived from the Greek word χιτών, a similar word, chiton, refers to a marine animal with a protective shell. The structure of chitin was solved by Albert Hofmann in 1929, chitin is a modified polysaccharide that contains nitrogen, it is synthesized from units of N-acetyl-D-glucosamine. Therefore, chitin may be described as cellulose with one group on each monomer replaced with an acetyl amine group. This allows for increased hydrogen bonding between adjacent polymers, giving the chitin-polymer matrix increased strength, in its pure, unmodified form, chitin is translucent, pliable, resilient, and quite tough. Combined with calcium carbonate, as in the shells of crustaceans and molluscs and this composite material is much harder and stiffer than pure chitin, and is tougher and less brittle than pure calcium carbonate. Another difference between pure and composite forms can be seen by comparing the body wall of a caterpillar to the stiff. The elaborate chitin gyroid construction in butterfly wings creates a model of optical devices having potential for innovations in biomimicry, scarab beetles in the genus Cyphochilus also utilize chitin to form extremely thin scales that diffusely reflect white light. These scales are networks of randomly ordered filaments of chitin with diameters on the scale of hundreds of nanometres, the multiple scattering of light is thought to play a role in the unusual whiteness of the scales. In addition, some wasps, such as Protopolybia chartergoides, orally secrete material containing predominantly chitin to reinforce the outer nest envelopes. Chitosan is produced commercially by deacetylation of chitin, chitosan is soluble in water, chitin producing organisms like protozoa, fungi, arthropods, and nematodes are often pathogens in other species. Keratinocytes in skin can also react to chitin or chitin fragments, according to in vitro studies, chitin is sensed by receptors, such as FIBCD1, KLRB1, REG3G, Toll-like receptor 2, CLEC7A, and mannose receptors. Plants also have receptors that can cause a response to chitin, namely chitin elicitor receptor kinase 1, the first chitin receptor was cloned in 2006. When the receptors are activated by chitin, genes related to plant defense are expressed, and jasmonate hormones are activated, commensal fungi that have ways to interact with the host immune response that as of 2016 were not well understood. Some pathogens produce chitin-binding proteins that mask the chitin they shed from these receptors, zymoseptoria tritici is an example of a fungal pathogen that has such blocking proteins, it is a major pest in wheat crops. Chitin was probably present in the exoskeletons of Cambrian arthropods such as trilobites, the oldest preserved chitin dates to the Oligocene, about 25 million years ago, consisting of a scorpion encased in amber
39.
Chela (organ)
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A chela /kˈiːlə/, also named claw, nipper, or pincer, is a pincer-like organ terminating certain limbs of some arthropods. The name comes from Greek through New Latin, legs bearing a chela are called chelipeds. Another name is claw because most chelae are curved and have a point like a claw
40.
Fresh water
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Fresh water is generally characterized by having low concentrations of dissolved salts and other total dissolved solids. The term specifically excludes seawater and brackish water although it does include mineral-rich waters such as chalybeate springs, the term sweet water has been used to describe fresh water in contrast to salt water. The term fresh water does not have the meaning as potable water. Much of the fresh water and ground water is unsuitable for drinking without some form of purification because of the presence of chemical or biological contaminants. It may also be in contact with the underlying underground water. The majority of water on Earth is in ice caps. The source of almost all water is precipitation from the atmosphere, in the form of mist, rain. Fresh water falling as mist, rain or snow contains materials dissolved from the atmosphere and material from the sea, in some cases this acid rain results in pollution of lakes and rivers. In coastal areas fresh water may contain significant concentrations of salts derived from the sea if windy conditions have lifted drops of seawater into the rain-bearing clouds. This can give rise to elevated concentrations of sodium, chloride, magnesium, significant quantities of iron may be transported in this way including the well-documented transfer of iron-rich rainfall falling in Brazil derived from sand-storms in the Sahara in north Africa. Water is an issue for the survival of all living organisms. Some can use water but many organisms including the great majority of higher plants. Out of all the water on Earth, saline water in oceans, seas and saline groundwater make up about 97% of it. Freshwater lakes contain about 87% of this surface water, including 29% in the African Great Lakes, 20% in Lake Baikal in Russia, 21% in the North American Great Lakes. Swamps have most of the balance only a small amount in rivers. In areas with no water on the ground surface, fresh water derived from precipitation may, because of its lower density. Most of the fresh water is frozen in ice sheets. Many areas suffer from lack of distribution of water, such as deserts
41.
Tropics
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The tropics are a region of the Earth surrounding the equator. The tropics are also referred to as the zone and the torrid zone. The tropics include all the areas on the Earth where the Sun is at a point directly overhead at least once during the solar year. The tropics are distinguished from the climatic and biomatic regions of Earth, which are the middle latitudes. Tropical is sometimes used in a sense for a tropical climate to mean warm to hot and moist year-round. Many tropical areas have a dry and wet season, the wet season, rainy season or green season, is the time of year, ranging from one or more months, when most of the average annual rainfall in a region falls. Areas with wet seasons are disseminated across portions of the tropics and subtropics, under the Köppen climate classification, for tropical climates, a wet season month is defined as a month where average precipitation is 60 millimetres or more. Tropical rainforests technically do not have dry or wet seasons, since their rainfall is distributed through the year. When the wet season occurs during the season, or summer, precipitation falls mainly during the late afternoon. The wet season is a time when air quality improves, freshwater quality improves and vegetation grows significantly, floods cause rivers to overflow their banks, and some animals to retreat to higher ground. Soil nutrients diminish and erosion increases, the incidence of malaria increases in areas where the rainy season coincides with high temperatures. Animals have adaptation and survival strategies for the wetter regime, unfortunately, the previous dry season leads to food shortages into the wet season, as the crops have yet to mature. Regions within the tropics may well not have a tropical climate, there are alpine tundra and snow-capped peaks, including Mauna Kea, Mount Kilimanjaro, and the Andes as far south as the northernmost parts of Chile and Argentina. Under the Köppen climate classification, much of the area within the tropics is classed not as tropical but as dry including the Sahara Desert. Tropical plants and animals are those native to the tropics. Tropical ecosystems may consist of rainforests, dry forests, spiny forests, desert. There are often significant areas of biodiversity, and species present, particularly in rainforests. In biogeography, the tropics are divided into Paleotropics and Neotropics, together, they are sometimes referred to as the Pantropic
42.
Pea crab
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The pea crab, Pinnotheres pisum, is a small crab in the family Pinnotheridae that lives as a parasite in oysters, clams, mussels, and other species of bivalves. Pea crabs are small crustaceans about the size of a pea or dime, with a smooth surface of the carapace. The exoskeleton of males is hard and circular and has eyes and antennae extending from their fronts, and the chelipeds are more robust in males than in females, which have more elongated chelipeds. The bodies of the female pea crabs are often translucent and show the inner organs and gonads as yellow and red, the relationship between the pea crab and its host is one of parasitism, rather than commensalism, since the host may be harmed by the crabs feeding activities. The pea crab solely relies on its host for food, safety, pea crabs have a variety of hosts, the most important of which are mollusks. The pea crab lives in the cavity of these hosts. Other hosts, in addition to oysters, include sea urchins, Pinnotheres can be found inside sand dollars, in the rectum of sea cucumbers, in the tubes of parchment worms, in the burrows of mud shrimp, or in the gills of sea squirts. The feeding process can be harmful to the crabs host when it feeds on the strings that help carry the food to the hosts mouth. A male pea crab will rub the edge of a shellfish containing a female pea crab for hours until the shellfish opens, Pinnotheres is Greek for guard of Pinna and pisum is Latin for a pea, in reference to the shape of the crab
43.
Japanese spider crab
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The Japanese spider crab, Macrocheira kaempferi, is a species of marine crab that lives in the waters around Japan. It has the largest leg span of any arthropod and it is the subject of small-scale fishery which has led to some conservation measures. The Japanese spider crab has the greatest leg span of any arthropod, the body may grow to a size of 40 cm or 16 in and the whole crab can weigh up to 19 kilograms —second only to the American lobster among all living arthropod species. The males have the longer chelipeds, females have much shorter chelipeds, apart from its outstanding size, the Japanese spider crab differs from other crabs in a number of ways. The first pleopods of males are unusually twisted, and its larvae appear primitive, the crab is orange, with white spots along the legs. It is reported to have a disposition, despite its ferocious appearance. The Japanese name for this species is taka-ashi-gani literally translating to “tall legs crab. ”Their armored exoskeletons help protect them from predators such as octopuses. The crabs bumpy carapace blends into the ocean floor. To further the illusion, a crab will adorn its shell with sponges. Japanese spider crabs are found off the southern coasts of the Japanese island of Honshū. Outlying populations have found in Iwate Prefecture and off Su-ao in Taiwan. Adults can be found at depths between 50 and 600 m and they like to inhabit vents and holes in the deeper parts of the ocean. The temperature preference of adults is unknown, but the species is regular at a depth of 300 m in Suruga Bay where the water generally is about 10 °C. Based on results from public aquariums, Japanese spider crabs tolerate between 6 and 16 °C, but most are maintained at 10–13 °C, female crabs carry the fertilized eggs attached to their abdominal appendages until they hatch into tiny planktonic larvae. Development of the larvae is temperature-dependent and takes between 54 and 72 days at 12–15 °C. During the larval stage the young crab looks nothing like its parents and it is small and transparent with a round, legless body and usually drifts as plankton at the surface of the ocean. The Japanese spider crab is an omnivore, consuming plant matter and animals. It also sometimes acts as a scavenger consuming dead animals, some have been known to scrape the ocean floor for plants and algae while others pry open the shells of mollusks
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