Eurypterids, often informally called sea scorpions, are a group of extinct arthropods that form the order Eurypterida. The earliest known eurypterids date to the Darriwilian stage of the Ordovician period 467.3 million years ago. The group is likely to have appeared first either during the Early Ordovician or Late Cambrian period. With approximately 250 species, the Eurypterida is the most diverse Paleozoic chelicerate order. Following their appearance during the Ordovician, eurypterids became major components of marine faunas during the Silurian, from which the majority of eurypterid species have been described. The Silurian genus Eurypterus accounts for more than 90% of all known eurypterid specimens. Though the group continued to diversify during the subsequent Devonian period, the eurypterids were heavily affected by the Late Devonian extinction event. They declined in numbers and diversity until becoming extinct during the Permian–Triassic extinction event 251.9 million years ago.
Illustration of Hibbertopterus, a large stylonurine (lacking swimming paddles) eurypterid
The holotype of Palmichnium kosinkiorum, containing the largest eurypterid footprints known
The supposed "gill tracts" of eurypterids have been compared to the air-breathing pseudotracheae present in the posterior legs of modern isopods, such as Oniscus (pictured).
The Silurian is a geologic period and system spanning 24.6 million years from the end of the Ordovician Period, at 443.8 million years ago (Mya), to the beginning of the Devonian Period, 419.2 Mya. The Silurian is the shortest period of the Paleozoic Era. As with other geologic periods, the rock beds that define the period's start and end are well identified, but the exact dates are uncertain by a few million years. The base of the Silurian is set at a series of major Ordovician–Silurian extinction events when up to 60% of marine genera were wiped out.
Ordovician-Silurian boundary on Hovedøya, Norway, showing brownish late Ordovician mudstone and later dark deep-water Silurian shale. The layers have been overturned by the Caledonian orogeny.
Fossils of the late Silurian sea bed
Silurian sea bed fossils collected from Wren's Nest Nature Reserve, Dudley UK
Crinoid fragments in a Silurian (Pridoli) limestone (Saaremaa, Estonia)