1.
Taxonomy (biology)
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Taxonomy is the science of defining groups of biological organisms on the basis of shared characteristics and giving names to those groups. The exact definition of taxonomy varies from source to source, but the core of the remains, the conception, naming. There is some disagreement as to whether biological nomenclature is considered a part of taxonomy, the broadest meaning of taxonomy is used here. The word taxonomy was introduced in 1813 by Candolle, in his Théorie élémentaire de la botanique, the term alpha taxonomy is primarily used today to refer to the discipline of finding, describing, and naming taxa, particularly species. In earlier literature, the term had a different meaning, referring to morphological taxonomy, ideals can, it may be said, never be completely realized. They have, however, a value of acting as permanent stimulants. Some of us please ourselves by thinking we are now groping in a beta taxonomy, turrill thus explicitly excludes from alpha taxonomy various areas of study that he includes within taxonomy as a whole, such as ecology, physiology, genetics, and cytology. He further excludes phylogenetic reconstruction from alpha taxonomy, thus, Ernst Mayr in 1968 defined beta taxonomy as the classification of ranks higher than species. This activity is what the term denotes, it is also referred to as beta taxonomy. How species should be defined in a group of organisms gives rise to practical and theoretical problems that are referred to as the species problem. The scientific work of deciding how to define species has been called microtaxonomy, by extension, macrotaxonomy is the study of groups at higher taxonomic ranks, from subgenus and above only, than species. While some descriptions of taxonomic history attempt to date taxonomy to ancient civilizations, earlier works were primarily descriptive, and focused on plants that were useful in agriculture or medicine. There are a number of stages in scientific thinking. Early taxonomy was based on criteria, the so-called artificial systems. Later came systems based on a complete consideration of the characteristics of taxa, referred to as natural systems, such as those of de Jussieu, de Candolle and Bentham. The publication of Charles Darwins Origin of Species led to new ways of thinking about classification based on evolutionary relationships and this was the concept of phyletic systems, from 1883 onwards. This approach was typified by those of Eichler and Engler, the advent of molecular genetics and statistical methodology allowed the creation of the modern era of phylogenetic systems based on cladistics, rather than morphology alone. Taxonomy has been called the worlds oldest profession, and naming and classifying our surroundings has likely been taking place as long as mankind has been able to communicate
2.
Plant
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Plants are mainly multicellular, predominantly photosynthetic eukaryotes of the kingdom Plantae. The term is generally limited to the green plants, which form an unranked clade Viridiplantae. This includes the plants, conifers and other gymnosperms, ferns, clubmosses, hornworts, liverworts, mosses and the green algae. Green plants have cell walls containing cellulose and obtain most of their energy from sunlight via photosynthesis by primary chloroplasts and their chloroplasts contain chlorophylls a and b, which gives them their green color. Some plants are parasitic and have lost the ability to produce amounts of chlorophyll or to photosynthesize. Plants are characterized by sexual reproduction and alternation of generations, although reproduction is also common. There are about 300–315 thousand species of plants, of which the great majority, green plants provide most of the worlds molecular oxygen and are the basis of most of Earths ecologies, especially on land. Plants that produce grains, fruits and vegetables form humankinds basic foodstuffs, Plants play many roles in culture. They are used as ornaments and, until recently and in variety, they have served as the source of most medicines. The scientific study of plants is known as botany, a branch of biology, Plants are one of the two groups into which all living things were traditionally divided, the other is animals. The division goes back at least as far as Aristotle, who distinguished between plants, which generally do not move, and animals, which often are mobile to catch their food. Much later, when Linnaeus created the basis of the system of scientific classification. Since then, it has become clear that the plant kingdom as originally defined included several unrelated groups, however, these organisms are still often considered plants, particularly in popular contexts. When the name Plantae or plant is applied to a group of organisms or taxon. The evolutionary history of plants is not yet settled. Those which have been called plants are in bold, the way in which the groups of green algae are combined and named varies considerably between authors. Algae comprise several different groups of organisms which produce energy through photosynthesis, most conspicuous among the algae are the seaweeds, multicellular algae that may roughly resemble land plants, but are classified among the brown, red and green algae. Each of these groups also includes various microscopic and single-celled organisms
3.
Flowering plant
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The flowering plants, also known as Angiospermae or Magnoliophyta, are the most diverse group of land plants, with 416 families, approx. 13,164 known genera and a total of c.295,383 known species, etymologically, angiosperm means a plant that produces seeds within an enclosure, in other words, a fruiting plant. The term angiosperm comes from the Greek composite word meaning enclosed seeds, the ancestors of flowering plants diverged from gymnosperms in the Triassic Period, during the range 245 to 202 million years ago, and the first flowering plants are known from 160 mya. They diversified extensively during the Lower Cretaceous, became widespread by 120 mya, angiosperms differ from other seed plants in several ways, described in the table. These distinguishing characteristics taken together have made the angiosperms the most diverse and numerous land plants, the amount and complexity of tissue-formation in flowering plants exceeds that of gymnosperms. The vascular bundles of the stem are arranged such that the xylem and phloem form concentric rings, in the dicotyledons, the bundles in the very young stem are arranged in an open ring, separating a central pith from an outer cortex. In each bundle, separating the xylem and phloem, is a layer of meristem or active formative tissue known as cambium, the soft phloem becomes crushed, but the hard wood persists and forms the bulk of the stem and branches of the woody perennial. Among the monocotyledons, the bundles are more numerous in the stem and are scattered through the ground tissue. They contain no cambium and once formed the stem increases in diameter only in exceptional cases, the characteristic feature of angiosperms is the flower. Flowers show remarkable variation in form and elaboration, and provide the most trustworthy external characteristics for establishing relationships among angiosperm species, the function of the flower is to ensure fertilization of the ovule and development of fruit containing seeds. The floral apparatus may arise terminally on a shoot or from the axil of a leaf, occasionally, as in violets, a flower arises singly in the axil of an ordinary foliage-leaf. There are two kinds of cells produced by flowers. Microspores, which divide to become pollen grains, are the male cells and are borne in the stamens. The female cells called megaspores, which divide to become the egg cell, are contained in the ovule. The flower may consist only of parts, as in willow. Usually, other structures are present and serve to protect the sporophylls, the individual members of these surrounding structures are known as sepals and petals. The outer series is usually green and leaf-like, and functions to protect the rest of the flower, the inner series is, in general, white or brightly colored, and is more delicate in structure. It functions to attract insect or bird pollinators, attraction is effected by color, scent, and nectar, which may be secreted in some part of the flower
4.
Beech
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Beech is a genus of deciduous trees in the family Fagaceae, native to temperate Europe, Asia and North America. Recent classification systems of the genus recognize ten to thirteen species in two subgenera, Engleriana and Fagus. Further differentiating characteristics include the whitish bloom on the underside of the leaves, the visible tertiary leaf veins, Fagus japonica, Fagus engleriana, and the species F. okamotoi, proposed by the botanist Chung-Fu Shen in 1992, comprise this subgenus. The better known Fagus subgenus beeches are high-branching with tall, stout trunks and this group includes Fagus sylvatica, Fagus grandifolia, Fagus crenata, Fagus lucida, Fagus longipetiolata, and Fagus hayatae. The classification of the European beech, Fagus sylvatica is complex, research suggests that beeches in Eurasia differentiated fairly late in evolutionary history, during the Miocene. The populations in this area represent a range of often overlapping morphotypes, within its family, the Fagaceae, recent research has suggested that Fagus is the evolutionarily most basal group. The southern beeches previously thought closely related to beeches, are now treated as members of a separate family and they are found in Australia, New Zealand, New Guinea, New Caledonia, Argentina and Chile. The European beech is the most commonly cultivated, although there are few important differences between species aside from detail elements such as leaf shape, the leaves of beech trees are entire or sparsely toothed, from 5–15 cm long and 4–10 cm broad. Beeches are monoecious, bearing male and female flowers on the same plant. The small flowers are unisexual, the flowers borne in pairs. They are produced in spring shortly after the new leaves appear, the bark is smooth and light grey. The fruit is a small, sharply three–angled nut 10–15 mm long, borne singly or in pairs in soft-spined husks 1. 5–2.5 cm long, known as cupules. The husk can have a variety of spine- to scale-like appendages, the nuts are edible, though bitter with a high tannin content, and are called beechnuts or beechmast. The name of the tree is of Indo-European origin, and played an important role in debates on the geographical origins of the Indo-European people. Greek φηγός is from the root, but the word was transferred to the oak tree as a result of the absence of beech trees in Greece. Beech grows on a range of soil types, acidic or basic. The tree canopy casts dense shade, and carpets the ground thickly with leaf litter, in North America, they often form beech-maple climax forests by partnering with the sugar maple. The beech blight aphid is a common pest of American beech trees, Beeches are also used as food plants by some species of Lepidoptera
5.
Oak
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An oak is a tree or shrub in the genus Quercus of the beech family, Fagaceae. There are approximately 600 extant species of oaks, the common name oak also appears in the names of species in related genera, notably Lithocarpus, as well as in those of unrelated species such as Grevillea robusta and the Casuarinaceae. North America contains the largest number of oak species, with approximately 90 occurring in the United States, the second greatest center of oak diversity is China, which contains approximately 100 species. Oaks have spirally arranged leaves, with lobate margins in many species, also, the acorns contain tannic acid, as do the leaves, which helps to guard from fungi and insects. Many deciduous species are marcescent, not dropping dead leaves until spring, in spring, a single oak tree produces both male flowers and small female flowers. The fruit is a nut called an acorn, borne in a structure known as a cupule, each acorn contains one seed and takes 6–18 months to mature. The live oaks are distinguished for being evergreen, but are not actually a distinct group, the oak tree is a flowering plant. Oaks may be divided into two genera and a number of sections, The genus Quercus is divided into the following sections, Quercus, the white oaks of Europe, Asia and North America. Styles are short, acorns mature in 6 months and taste sweet or slightly bitter, the leaves mostly lack a bristle on their lobe tips, which are usually rounded. The type species is Quercus robur, Mesobalanus, Hungarian oak and its relatives of Europe and Asia. Styles long, acorns mature in about 6 months and taste bitter, the section Mesobalanus is closely related to section Quercus and sometimes included in it. Cerris, the Turkey oak and its relatives of Europe and Asia, styles long, acorn mature in 18 months and taste very bitter. The inside of the shell is hairless. Its leaves typically have sharp tips, with bristles at the lobe tip. Protobalanus, the live oak and its relatives, in southwest United States. Styles short, acorns mature in 18 months and taste very bitter, the inside of the acorn shell appears woolly. Leaves typically have sharp tips, with bristles at the lobe tip. Lobatae, the red oaks of North America, Central America, styles long, acorns mature in 18 months and taste very bitter
6.
Deciduous
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In a more general sense, deciduous means the dropping of a part that is no longer needed or falling away after its purpose is finished. In plants it is the result of natural processes, in botany and horticulture, deciduous plants, including trees, shrubs and herbaceous perennials, are those that lose all of their leaves for part of the year. In some cases leaf loss coincides with winter—namely in temperate or polar climates, in other parts of the world, including tropical, subtropical, and arid regions, plants lose their leaves during the dry season or other seasons, depending on variations in rainfall. The converse of deciduous is coniferous, where foliage is shed on a different schedule from deciduous trees, plants that are intermediate may be called semi-deciduous, they lose old foliage as new growth begins. Other plants are semi-evergreen and lose their leaves before the growing season. Many deciduous plants flower during the period when they are leafless, the absence of leaves improves wind transmission of pollen for wind-pollinated plants and increases the visibility of the flowers to insects in insect-pollinated plants. This strategy is not without risks, as the flowers can be damaged by frost or, in dry season regions, leaf drop or abscission involves complex physiological signals and changes within plants. The process of photosynthesis steadily degrades the supply of chlorophylls in foliage, the brightest leaf colors are produced when days grow short and nights are cool, but remain above freezing. These other pigments include carotenoids that are yellow, brown, anthocyanin pigments produce red and purple colors, though they are not always present in the leaves. Rather, they are produced in the foliage in late summer, parts of the world that have showy displays of bright autumn colors are limited to locations where days become short and nights are cool. In other parts of the world, the leaves of deciduous trees simply fall off without turning the bright colors produced from the accumulation of anthocyanin pigments, the beginnings of leaf drop starts when an abscission layer is formed between the leaf petiole and the stem. This layer is formed in the spring during active new growth of the leaf, the cells are sensitive to a plant hormone called auxin that is produced by the leaf and other parts of the plant. The elongation of cells break the connection between the different cell layers, allowing the leaf to break away from the plant. It also forms a layer that seals the break, so the plant does not lose sap, in the spring, these proteins are used as a nitrogen source during the growth of new leaves or flowers. Plants with deciduous foliage have advantages and disadvantages compared to plants with evergreen foliage, evergreens suffer greater water loss during the winter and they also can experience greater predation pressure, especially when small. Losing leaves in winter may reduce damage from insects, repairing leaves, removing leaves also reduces cavitation which can damage xylem vessels in plants. This then allows deciduous plants to have xylem vessels with larger diameters, the deciduous characteristic has developed repeatedly among woody plants. Trees include maple, many oaks and nothofagus, elm, aspen, Deciduous shrubs include honeysuckle, viburnum, and many others
7.
Flower
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A flower, sometimes known as a bloom or blossom, is the reproductive structure found in plants that are floral. The biological function of a flower is to effect reproduction, usually by providing a mechanism for the union of sperm with eggs, Flowers may facilitate outcrossing or allow selfing. Some flowers produce diaspores without fertilization, Flowers contain sporangia and are the site where gametophytes develop. Many flowers have evolved to be attractive to animals, so as to them to be vectors for the transfer of pollen. After fertilization, the ovary of the flower develops into fruit containing seeds, the essential parts of a flower can be considered in two parts, the vegetative part, consisting of petals and associated structures in the perianth, and the reproductive or sexual parts. A stereotypical flower consists of four kinds of structures attached to the tip of a short stalk, each of these kinds of parts is arranged in a whorl on the receptacle. The four main whorls are as follows, Collectively the calyx, corolla, the next whorl toward the apex, composed of units called petals, which are typically thin, soft and colored to attract animals that help the process of pollination. Androecium, the whorl, consisting of units called stamens. Stamens consist of two parts, a called a filament, topped by an anther where pollen is produced by meiosis. Gynoecium, the innermost whorl of a flower, consisting of one or more units called carpels, the carpel or multiple fused carpels form a hollow structure called an ovary, which produces ovules internally. Ovules are megasporangia and they in turn produce megaspores by meiosis which develop into female gametophytes and these give rise to egg cells. The gynoecium of a flower is described using an alternative terminology wherein the structure one sees in the innermost whorl is called a pistil. A pistil may consist of a carpel or a number of carpels fused together. The sticky tip of the pistil, the stigma, is the receptor of pollen, the supportive stalk, the style, becomes the pathway for pollen tubes to grow from pollen grains adhering to the stigma. The relationship to the gynoecium on the receptacle is described as hypogynous, perigynous, although the arrangement described above is considered typical, plant species show a wide variation in floral structure. These modifications have significance in the evolution of flowering plants and are used extensively by botanists to establish relationships among plant species, the four main parts of a flower are generally defined by their positions on the receptacle and not by their function. Many flowers lack some parts or parts may be modified into other functions and/or look like what is typically another part, in some families, like Ranunculaceae, the petals are greatly reduced and in many species the sepals are colorful and petal-like. Other flowers have modified stamens that are petal-like, the flowers of Peonies and Roses are mostly petaloid stamens
8.
Catkin
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A catkin or ament is a slim, cylindrical flower cluster, with inconspicuous or no petals, usually wind-pollinated but sometimes insect-pollinated. They contain many, usually flowers, arranged closely along a central stem which is often drooping. They are found in plant families, including Betulaceae, Fagaceae, Moraceae. For some time, they were believed to be a key synapomorphy among the proposed Hamamelididae, based on molecular phylogeny work, it is now believed that Hamamelididae is a polyphyletic group. This suggests that the flower arrangement has arisen at least twice independently by convergent evolution, in Fagales. Such a convergent evolution raises questions about what the ancestral inflorescence characters might be, in many of these plants, only the male flowers form catkins, and the female flowers are single, a cone or other types. In other plants both male and female flowers are borne in catkins, catkin-bearing plants include many other trees or shrubs such as birch, willow, hickory, sweet chestnut and sweetfern. The word catkin is a loanword from the old Dutch katteken, meaning kitten, ament is from the Latin amentum, meaning thong or strap. Catkins in Fagales Catkins in Salicaceae
9.
Stipule
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In botany, stipule is a term coined by Linnaeus which refers to outgrowths borne on either side of the base of a leafstalk. A pair of stipules is considered part of the anatomy of the leaf of a flowering plant. In some older botanical writing, the term stipule was used generally to refer to any small leaves or leaf-parts. The position of stipules on a plant varies widely from species to species, stipules are most common on dicotyledons, where they appear in pairs alongside each leaf. Some monocotyledon plants display stipule-like structures, but only one per leaf. A relationship exists between the anatomy of the node and the presence or absence of stipules, most plants with trilacunar nodes have stipules, species with unilacunar nodes lack stipules. Stipules are morphologically variable and might appear as glands, scales, hairs, spines, if a single stipule goes all the way around the stem, it is known as an ochrea. The three types of stipules according to duration are caducous, deciduous and persistent, caducous stipules fall off before the leaf unfolds, while deciduous stipules fall off immediately after the leaf unfolds. Persistent stipules remain attached to the plant, stipules can be considered free lateral, adnate, interpetiolar, intrapetiolar, ochreate, foliaceous, bud scales, tendrillar or spiny. A stipule can be fused to the stem, or to the other stipule from the same node, a stipule is adnate if its fused together on part of the petiole length, but the anterior is still free. A stipule is intrapetiolar if it is located in the angle thats between a stem and a petiole, in this case, the two stipules generally form together and appear to be one stipule. A stipule is ochreate if a single stipule appears to be a tube that goes all the way around the stem. A stipule is foliaceous if it is leaf-like and these are generally used to photosynthesize. A stipule is considered a bud scale if it is hard or scaly and these generally fall off as soon as the leaf unfolds. A stipule is considered tendrillar if they are long thin tendrils, a stipule is considered spiny if they are long and pointy. These are generally used to deter animals, a stipule is considered to be abaxial, counter or leaf opposed if its located on the opposite side to where the leaf meets the stem. Stipules can be used by the plants in various ways, and it is known that foliaceous stipules are used like leaves to make energy for the plants. Sometimes stipules protect the next leaf or bud as it grows in, then falls off after the leaf unfolds, stipules can be used as climbing tendrils by climbing plants
10.
Chestnut
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The chestnut group is a genus of eight or nine species of deciduous trees and shrubs in the beech family Fagaceae, native to temperate regions of the Northern Hemisphere. The name also refers to the edible nuts they produce, Chestnuts belong to the family Fagaceae, which also includes oaks and beeches. Unrelated but externally similar species of horse chestnut are abundant around Europe, other trees commonly mistaken for chestnut trees are the chestnut oak and the American beech, both of which are also in Fagaceae. The name chestnut is derived from an earlier English term chesten nut, the name Castanea is probably derived from the old name for the sweet chestnut, either in Latin or in Ancient Greek. Another possible source of the name is the town of Kastania in Thessaly, Greece, more probable, in the Mediterranean climate zone, chestnut trees are rarer in Greece because the chalky soil is not conducive to the trees growth. Kastania is located on one of the relatively few sedimentary or siliceous outcrops and they grow so abundantly there, their presence would have determined the places name. Still others take the name as coming from the Greek name of Sardis glans – Sardis being the capital of Lydia, Asia Minor, the name is cited twice in the King James Version of the Bible. In one instance, Jacob puts peeled twigs in the troughs to promote healthy offspring of his livestock. Although it may indicate another tree, it indicates the fruit was a staple food in the early 17th century. These synonyms are or have been in use, Fagus castanea, Sardian nut, Jupiters nut, husked nut, Chestnut trees are of moderate growth rate to fast-growing for American and European species. Their mature heights vary from the smallest species of chinkapins, often shrubby, to the giant of past American forests, C. dentata that could reach 60 m. Between these extremes are found the Japanese chestnut at 10 m average, followed by the Chinese chestnut at about 15 m, when standing on their own, they spread on the sides and develop broad, rounded, dense crowns at maturity. The two latters foliage has striking yellow autumn colouring and its bark is smooth when young, of a vinous maroon or red-brown colour for the American chestnut, grey for the European chestnut. The leaves are simple, ovate or lanceolate, 10–30 cm long and 4–10 cm wide, with sharply pointed, the flowers follow the leaves, appearing in late spring or early summer or into July. They are arranged in long catkins of two kinds, with both kinds being borne on every tree, some catkins are made of only male flowers, which mature first. Each flower has eight stamens, or 10 to 12 for C. mollissima, the ripe pollen carries a heavy, sweet odour that some people find too sweet or unpleasant. Other catkins have these pollen-bearing flowers, but also carry near the twig from which these spring, two or three flowers together form a four-lobed prickly calybium, which ultimately grows completely together to make the brown hull, or husk, covering the fruits. Chestnut flowers are not self-compatible, so two trees are required for pollination, all Castanea species readily hybridize with each other
11.
Quercus suber
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Quercus suber, commonly called the cork oak, is a medium-sized, evergreen oak tree in the section Quercus sect. It is the source of cork for wine bottle stoppers and other uses. It is native to southwest Europe and northwest Africa and it grows to up to 20 m, although it is typically more stunted in its native environment. The leaves are 4 to 7 cm long, weakly lobed or coarsely toothed, dark green above, the acorns are 2 to 3 cm long, in a deep cup fringed with elongated scales. Natural stands of oak can support diverse ecosystems. In Western Europe, namely in Portugal and Spain, the oak forests are home to endangered species such as the Iberian lynx. The tree has a thick, insulating bark that may have been the cork oaks evolutionary answer to forest fires, the quick regeneration of the pyrophyte tree seems to be an advantage compared to other species that, after a fire, return to an initial stage of development. The tree forms a thick, rugged bark containing high levels of suberin, over time the cork cambium layer of bark can develop considerable thickness and can be harvested every 7 to ten years to produce cork. The harvesting of cork does not harm the tree, in fact, only the bark is extracted, and a new layer of cork regrows, making it a renewable resource. The tree is cultivated in Spain, Portugal, Algeria, Morocco, France, Italy, Cork oaks are considered to be soil builders and their fruits have been shown to have useful insecticidal properties. Cork oak forests cover approximately 25,000 square kilometres in those countries, Portugal accounts for around 50% of the world cork harvest. Q. suber is commonly grown in agroforestry systems, known as montado in Portugal and these are open woods with low tree density. In these systems, forage species are grown under the trees. Cork oaks live about 150 to 250 years, virgin cork is the first cork cut from generally 25-year-old trees. Another 9 to 12 years is required for the second harvest, Cork harvesting is done entirely without machinery, being dependent solely on human labor. Usually five people are required to harvest the bark, using a small axe. The process requires training due to the required to harvest bark without harming the tree. The European cork industry produces 300,000 tonnes of cork a year, with a value of €1.5 billion, wine corks represent 15% of cork usage by weight but 66% of revenues
12.
Nothofagus
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Nothofagus, also known as the southern beeches, is a genus of 43 species of trees and shrubs native to the Southern Hemisphere in southern South America and Australasia. The species are ecological dominants in many forests in these regions. Some species are naturalised in Germany and Great Britain. The genus has a fossil record of leaves, cupules and pollen, with fossils extending into the late Cretaceous and occurring in Australia, New Zealand, Antarctica. In the past, they were included in the family Fagaceae, but genetic tests revealed them to be distinct, and they are now included in their own family. The leaves are toothed or entire, evergreen or deciduous, the fruit is a small, flattened or triangular nut, borne in cupules containing one to seven nuts. Nothofagus species are used as food plants by the larvae of moths of the genus Aenetus, including A. eximia. Sexual reproduction has since shown to be possible. This proposed change is not taxonomically essential and has not been accepted outside New Zealand, in South America the northern genus limit can be construed as La Campana National Park and the Vizcachas Mountains in the central part of Chile. Every four to six years or so, Nothofagus produces a crop of seeds and is known as the beech mast. In New Zealand, the beech mast causes an increase in the population of introduced mammals such as mice, rats, when the rodent population collapses, the stoats begin to prey on native bird species, many of which are threatened with extinction. This phenomenon is covered in detail in the article on stoats in New Zealand. Beechwood Honeydew is a type of honey made in New Zealand from the secretions of Hemiptera aphids feeding on Nothofagus trees, cyttaria, genus of ascomycete fungi found on or associated with Nothofagus Misodendrum, specialist parasites of Nothofagus
