Graptolites are a group of colonial animals, members of the subclass Graptolithina within the class Pterobranchia. These filter-feeding organisms are known chiefly from fossils found from the Middle Cambrian through the Lower Carboniferous (Mississippian). A possible early graptolite, Chaunograptus, is known from the Middle Cambrian. Recent analyses have favored the idea that the living pterobranch Rhabdopleura represents an extant graptolite which diverged from the rest of the group in the Cambrian.
Graptolite
Diversity of graptolite colony forms
A hypothetical graptolite zooid inside its theca, reconstructed based on living pterobranchs
Pendeograptus fruticosus from the Bendigonian Australian Stage (Lower Ordovician; 477–474 mya) near Bendigo, Victoria, Australia. There are two overlapping, three-stiped rhabdosomes.
Biostratigraphy is the branch of stratigraphy which focuses on correlating and assigning relative ages of rock strata by using the fossil assemblages contained within them. The primary objective of biostratigraphy is correlation, demonstrating that a particular horizon in one geological section represents the same period of time as another horizon at a different section. Fossils within these strata are useful because sediments of the same age can look completely different, due to local variations in the sedimentary environment. For example, one section might have been made up of clays and marls, while another has more chalky limestones. However, if the fossil species recorded are similar, the two sediments are likely to have been laid down around the same time. Ideally these fossils are used to help identify biozones, as they make up the basic biostratigraphy units, and define geological time periods based upon the fossil species found within each section.
The first reef builder is a worldwide index fossil for the Lower Cambrian
Amplexograptus, a graptolite index fossil, from the Ordovician near Caney Springs, Tennessee.
Image displaying newly discovered fossil being introduced into the succession sequence.