1.
Sacred
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Objects are often considered sacred if used for spiritual purposes, such as the worship or service of gods. The property is often ascribed to objects, or places, the word sacred descends from the Latin wikt, sacer, that is consecrated, or dedicated to the gods or anything in their power, and to sacerdos and sanctum, set apart. Although there are similarities between the sacred and holy and they are sometimes used interchangeably, there are subtle differences. Holiness is generally the term used in relation to persons and relationship, while sacredness is used in relation to objects, places, thus a saint may be considered as holy, but he is not viewed as sacred. However, there are things that are holy and sacred such as the holy bible. The Scottish hale is the most complete form of this Old English root. The word holy in its modern form appears in Wycliffes Bible of 1382, in non-specialist contexts, the term holy is used in a more general way, to refer to someone or something that is associated with a divine power, such as water used for baptism. While both words denote something or someone set apart to the worship of God and therefore worthy of respect and in some cases veneration, Holiness originates in God and is communicated to things, places, times, and persons engaged in His Service. Thus holiness or sanctity is the outcome of sanctification, that Divine act by which God freely justifies a person, hierology is the study of sacred literature or lore. Mircea Eliade outlines that religion should not be interpreted only as belief in deities and he analyses the dialectic of the sacred. The sacred is presented in relation to the profane, the relation between the sacred and the profane is not of opposition, but of complementarity, as the profane is viewed as a hierophany. In Durkheims theory, the sacred represented the interests of the group, especially unity, the profane, on the other hand, involved mundane individual concerns. Durkheim explicitly stated that the dichotomy sacred/profane was not equivalent to good/evil, the sacred could be good or evil, and the profane could be either as well. In Theravada Buddhism one finds the designation of noble person or ariyapuggala, the Buddha described four grades of such person depending on their level of purity. This purity is measured by which of the ten fetters and klesha have been purified and integrated from the mindstream and these persons are called Sotāpanna, Sakadagami, Anāgāmi and Arahant. Catholicism has inherited much of the Jewish vision of the world in terms of holiness, the calendar gives shape to Catholic practice, which tends to focus on the Eucharist, in which the Real Presence of Christ is manifested. Holy days, celebrating events of the life of Christ and the lives of Catholic saints officially recognized as holy, are celebrated throughout the year, in extension of this focus on the Sacrament as holy, many objects in Catholicism are also considered holy. They are called sacramentals and are blessed by a priest
2.
Ilex aquifolium
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Ilex aquifolium, is a species of holly native to western and southern Europe, northwest Africa, and southwest Asia. I. aquifolium is the familiar in Christmas decoration, and is regarded as the type species of the genus Ilex. It is a tree or shrub found, for example, in shady areas of forests of oak. It has a capacity to adapt to different conditions and is a pioneer species that repopulates the margins of forests or clearcuts. As a tree, Christmas holly can exceed 10 m in height and it is usually found as a shrub or a small tree about 1 m tall with a straight trunk and pyramidal crown, which branches from the base. It grows slowly and does not usually fully mature due to grazing, cutting and it can live 500 years, but usually does not reach 100. The flowers are white, four-lobed, and pollinated by bees, Holly is dioecious, meaning that there are male plants and female plants. The sex cannot be determined until the plants begin flowering, usually between 4 and 12 years of age, in male specimens, the flowers are yellowish and appear in axillary groups. In the female, flowers are isolated or in groups of three and are small and white or slightly pink, and consist of four petals and four sepals partially fused at the base and they are eaten by rodents, birds and larger herbivores. Each fruit contains 3 to 4 seeds which do not germinate until the second or third spring, the fruit only appears on female plants, which require male plants nearby to fertilise them. Today, holly is found in western Asia and Europe in the undergrowth of oak forest and beech forest in particular and it requires moist, shady environments, found within forests or in shady slopes, cliffs, and mountain gorges. It has been placed on the Washington State Noxious Weed Control Boards monitor list, during the Cenozoic Era, the Mediterranean region, Europe, and northwest Africa had a wetter climate and were largely covered by laurel forests. Holly was a representative species of this biome, where many current species of the genus Ilex were present. With the drying of the Mediterranean Basin during the Pliocene, the forests gradually retreated, replaced by more drought-tolerant sclerophyll plant communities. The modern Ilex aquifolium resulted from this change, most of the last remaining laurel forests around the Mediterranean are believed to have died out approximately 10,000 years ago at the end of the Pleistocene. Holly is a pioneer species that prefers relatively moist areas. The plant is common in the garrigue and maquis and is found in deciduous forest. Pure stands of hollies can grow into a labyrinth of vaults in which thrushes and deer take refuge, after the first frost of the season, holly fruits become soft and fall to the ground serving as important food for winter birds at a time of scarce resources
3.
Taxonomy (biology)
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Taxonomy is the science of defining groups of biological organisms on the basis of shared characteristics and giving names to those groups. The exact definition of taxonomy varies from source to source, but the core of the remains, the conception, naming. There is some disagreement as to whether biological nomenclature is considered a part of taxonomy, the broadest meaning of taxonomy is used here. The word taxonomy was introduced in 1813 by Candolle, in his Théorie élémentaire de la botanique, the term alpha taxonomy is primarily used today to refer to the discipline of finding, describing, and naming taxa, particularly species. In earlier literature, the term had a different meaning, referring to morphological taxonomy, ideals can, it may be said, never be completely realized. They have, however, a value of acting as permanent stimulants. Some of us please ourselves by thinking we are now groping in a beta taxonomy, turrill thus explicitly excludes from alpha taxonomy various areas of study that he includes within taxonomy as a whole, such as ecology, physiology, genetics, and cytology. He further excludes phylogenetic reconstruction from alpha taxonomy, thus, Ernst Mayr in 1968 defined beta taxonomy as the classification of ranks higher than species. This activity is what the term denotes, it is also referred to as beta taxonomy. How species should be defined in a group of organisms gives rise to practical and theoretical problems that are referred to as the species problem. The scientific work of deciding how to define species has been called microtaxonomy, by extension, macrotaxonomy is the study of groups at higher taxonomic ranks, from subgenus and above only, than species. While some descriptions of taxonomic history attempt to date taxonomy to ancient civilizations, earlier works were primarily descriptive, and focused on plants that were useful in agriculture or medicine. There are a number of stages in scientific thinking. Early taxonomy was based on criteria, the so-called artificial systems. Later came systems based on a complete consideration of the characteristics of taxa, referred to as natural systems, such as those of de Jussieu, de Candolle and Bentham. The publication of Charles Darwins Origin of Species led to new ways of thinking about classification based on evolutionary relationships and this was the concept of phyletic systems, from 1883 onwards. This approach was typified by those of Eichler and Engler, the advent of molecular genetics and statistical methodology allowed the creation of the modern era of phylogenetic systems based on cladistics, rather than morphology alone. Taxonomy has been called the worlds oldest profession, and naming and classifying our surroundings has likely been taking place as long as mankind has been able to communicate
4.
Plant
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Plants are mainly multicellular, predominantly photosynthetic eukaryotes of the kingdom Plantae. The term is generally limited to the green plants, which form an unranked clade Viridiplantae. This includes the plants, conifers and other gymnosperms, ferns, clubmosses, hornworts, liverworts, mosses and the green algae. Green plants have cell walls containing cellulose and obtain most of their energy from sunlight via photosynthesis by primary chloroplasts and their chloroplasts contain chlorophylls a and b, which gives them their green color. Some plants are parasitic and have lost the ability to produce amounts of chlorophyll or to photosynthesize. Plants are characterized by sexual reproduction and alternation of generations, although reproduction is also common. There are about 300–315 thousand species of plants, of which the great majority, green plants provide most of the worlds molecular oxygen and are the basis of most of Earths ecologies, especially on land. Plants that produce grains, fruits and vegetables form humankinds basic foodstuffs, Plants play many roles in culture. They are used as ornaments and, until recently and in variety, they have served as the source of most medicines. The scientific study of plants is known as botany, a branch of biology, Plants are one of the two groups into which all living things were traditionally divided, the other is animals. The division goes back at least as far as Aristotle, who distinguished between plants, which generally do not move, and animals, which often are mobile to catch their food. Much later, when Linnaeus created the basis of the system of scientific classification. Since then, it has become clear that the plant kingdom as originally defined included several unrelated groups, however, these organisms are still often considered plants, particularly in popular contexts. When the name Plantae or plant is applied to a group of organisms or taxon. The evolutionary history of plants is not yet settled. Those which have been called plants are in bold, the way in which the groups of green algae are combined and named varies considerably between authors. Algae comprise several different groups of organisms which produce energy through photosynthesis, most conspicuous among the algae are the seaweeds, multicellular algae that may roughly resemble land plants, but are classified among the brown, red and green algae. Each of these groups also includes various microscopic and single-celled organisms
5.
Flowering plant
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The flowering plants, also known as Angiospermae or Magnoliophyta, are the most diverse group of land plants, with 416 families, approx. 13,164 known genera and a total of c.295,383 known species, etymologically, angiosperm means a plant that produces seeds within an enclosure, in other words, a fruiting plant. The term angiosperm comes from the Greek composite word meaning enclosed seeds, the ancestors of flowering plants diverged from gymnosperms in the Triassic Period, during the range 245 to 202 million years ago, and the first flowering plants are known from 160 mya. They diversified extensively during the Lower Cretaceous, became widespread by 120 mya, angiosperms differ from other seed plants in several ways, described in the table. These distinguishing characteristics taken together have made the angiosperms the most diverse and numerous land plants, the amount and complexity of tissue-formation in flowering plants exceeds that of gymnosperms. The vascular bundles of the stem are arranged such that the xylem and phloem form concentric rings, in the dicotyledons, the bundles in the very young stem are arranged in an open ring, separating a central pith from an outer cortex. In each bundle, separating the xylem and phloem, is a layer of meristem or active formative tissue known as cambium, the soft phloem becomes crushed, but the hard wood persists and forms the bulk of the stem and branches of the woody perennial. Among the monocotyledons, the bundles are more numerous in the stem and are scattered through the ground tissue. They contain no cambium and once formed the stem increases in diameter only in exceptional cases, the characteristic feature of angiosperms is the flower. Flowers show remarkable variation in form and elaboration, and provide the most trustworthy external characteristics for establishing relationships among angiosperm species, the function of the flower is to ensure fertilization of the ovule and development of fruit containing seeds. The floral apparatus may arise terminally on a shoot or from the axil of a leaf, occasionally, as in violets, a flower arises singly in the axil of an ordinary foliage-leaf. There are two kinds of cells produced by flowers. Microspores, which divide to become pollen grains, are the male cells and are borne in the stamens. The female cells called megaspores, which divide to become the egg cell, are contained in the ovule. The flower may consist only of parts, as in willow. Usually, other structures are present and serve to protect the sporophylls, the individual members of these surrounding structures are known as sepals and petals. The outer series is usually green and leaf-like, and functions to protect the rest of the flower, the inner series is, in general, white or brightly colored, and is more delicate in structure. It functions to attract insect or bird pollinators, attraction is effected by color, scent, and nectar, which may be secreted in some part of the flower
6.
Eudicots
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The eudicots, Eudicotidae or eudicotyledons are a monophyletic clade of flowering plants that had been called tricolpates or non-magnoliid dicots by previous authors. The close relationships among flowering plants with tricolpate pollen grains was initially seen in studies of shared derived characters. These plants have a trait in their pollen grains of exhibiting three colpi or grooves paralleling the polar axis. Later molecular evidence confirmed the basis for the evolutionary relationships among flowering plants with tricolpate pollen grains. The term means true dicotyledons, as it contains the majority of plants that have been considered dicots and have characteristics of the dicots, the term eudicots has subsequently been widely adopted in botany to refer to one of the two largest clades of angiosperms, monocots being the other. The remaining angiosperms are sometimes referred to as basal angiosperms or paleodicots, the other name for the eudicots is tricolpates, a name which refers to the grooved structure of the pollen. Members of the group have tricolpate pollen, or forms derived from it and these pollens have three or more pores set in furrows called colpi. In contrast, most of the seed plants produce monosulcate pollen. The name tricolpates is preferred by some botanists to avoid confusion with the dicots, numerous familiar plants are eudicots, including many common food plants, trees, and ornamentals. Most leafy trees of midlatitudes also belong to eudicots, with exceptions being magnolias and tulip trees which belong to magnoliids, and Ginkgo biloba. The name eudicots is used in the APG system, of 1998 and it is applied to a clade, a monophyletic group, which includes most of the dicots. The eudicots can be divided into two groups, the basal eudicots and the core eudicots, basal eudicot is an informal name for a paraphyletic group. The core eudicots are a monophyletic group, a 2010 study suggested the core eudicots can be divided into two clades, Gunnerales and a clade called Pentapetalae, comprising all the remaining core eudicots
7.
Asterids
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In the APG IV system for the classification of flowering plants, the name asterids denotes a clade. Most of the taxa belonging to this clade had been referred to the Asteridae in the Cronquist system, the name asterids resembles the earlier botanical name but is intended to be the name of a clade rather than a formal ranked name, in the sense of the ICBN. The phylogenetic tree presented hereafter has been proposed by the APG IV project, genetic analysis carried out after APG II maintains that the sister to all other asterids are the Cornales. A second order that split from the base of the asterids are the Ericales, the remaining orders cluster into two clades, the lamiids and the campanulids. The structure of both of these clades has changed in APG III, in APG III system, the following clades were renamed, euasterids I → lamiids euasterids II → campanulids Asterids in Stevens, P. F. Angiosperm Phylogeny Website
8.
Augustin Pyramus de Candolle
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Augustin Pyramus de Candolle also spelled Augustin Pyrame de Candolle was a Swiss botanist. René Louiche Desfontaines launched de Candolles botanical career by recommending him at an herbarium, within a couple of years de Candolle had established a new genus, and he went on to document hundreds of plant families and create a new natural plant classification system. Although de Candolles main focus was botany, he contributed to related fields such as phytogeography, agronomy, paleontology, medical botany. Candolle originated the idea of Natures war, which influenced Charles Darwin, de Candolle recognized that multiple species may develop similar characteristics that did not appear in a common evolutionary ancestor, this was later termed analogy. During his work with plants, de Candolle noticed that plant leaf movements follow a cycle in constant light. Though many scientists doubted de Candolles findings, experiments over a century later demonstrated that ″the internal biological clock″ indeed exists, Candolles descendants continued his work on plant classification. Alphonse de Candolle and Casimir Pyrame de Candolle contributed to the Prodromus Systematis Naturalis Regni Vegetabilis, Augustin Pyramus de Candolle was born on 4 February 1778 in Geneva, Switzerland, to Augustin de Candolle, a former official, and his wife, Louise Eléonore Brière. His family descended from one of the ancient families of Provence in France, at age seven de Candolle contracted of a severe case of hydrocephalus, which significantly affected his childhood. Candolles initiation to botany happened in 1794 thanks to Pierre Etienne Vaucher in Geneva and he spent then four years at the Geneva Academy, studying science and law according to his fathers wishes. In 1798, he moved to Paris after Geneva had been annexed to the French Republic and his botanical career began with the help of René Louiche Desfontaines, who recommended de Candolle for work in the herbarium of Charles Louis LHéritier de Brutelle during the summer of 1798. The position elevated de Candolles reputation and also led to valuable instruction from Desfontaines himself, the premise of de Candolles method is that taxa do not fall along a linear scale, they are discrete, not continuous. In 1804, de Candolle published his Essai sur les propriétés médicales des plantes and was granted a doctor of medicine degree by the faculty of Paris. In 1807 he was appointed professor of botany in the faculty of the University of Montpellier. While in Montpellier, de Candolle published his Théorie élémentaire de la botanique, which introduced a new classification system and the word taxonomy. Candolle moved back to Geneva in 1816 and in the year was invited by the government of the Canton of Geneva to fill the newly created chair of natural history. De Candolle spent the rest of his life in an attempt to elaborate and complete his natural system of botanical classification, de Candolle published initial work in his Regni vegetabillis systema naturale, but after two volumes he realized he could not complete the project on such a large scale. Consequently, he began his less extensive Prodromus Systematis Naturalis Regni Vegetabilis in 1824, however, he was able to finish only seven volumes, or two-thirds of the whole. Even so, he was able to characterize over one hundred families of plants, although de Candolles main focus was botany, throughout his career he also dabbled in fields related to botany, such as phytogeography, agronomy, paleontology, medical botany, and economic botany
9.
Achille Richard
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Achille Richard was a French botanist and physician. He was son of a notable botanist, Louis-Claude Marie Richard, pharmacist in the French fleet and member of several well-known societies of their time. Achille Richard was one of the leaders of his time. He entered on 24 February 1834, as member of the French Academy of Sciences and he was also member of the French National Academy of Medicine. Achille Richard studied and described several genera of orchids that take his abbreviation in the generic name, monographie du genre Hydrocotyle de la famille des ombellifères. - Claude Richard 1784–1868 Other botanists called Richard are, Louis-Claude Marie Richard, joseph Herve Pierre Richard Books by Achille Richard at the Biodiversity Library
10.
Carl Linnaeus
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Carl Linnaeus, also known after his ennoblement as Carl von Linné, was a Swedish botanist, physician, and zoologist, who formalised the modern system of naming organisms called binomial nomenclature. He is known by the father of modern taxonomy. Many of his writings were in Latin, and his name is rendered in Latin as Carolus Linnæus, Linnaeus was born in the countryside of Småland, in southern Sweden. He received most of his education at Uppsala University. He lived abroad between 1735 and 1738, where he studied and also published a first edition of his Systema Naturae in the Netherlands and he then returned to Sweden, where he became professor of medicine and botany at Uppsala. In the 1740s, he was sent on journeys through Sweden to find and classify plants. In the 1750s and 1760s, he continued to collect and classify animals, plants, and minerals, at the time of his death, he was one of the most acclaimed scientists in Europe. The philosopher Jean-Jacques Rousseau sent him the message, Tell him I know no man on earth. The German writer Johann Wolfgang von Goethe wrote, With the exception of Shakespeare and Spinoza, Swedish author August Strindberg wrote, Linnaeus was in reality a poet who happened to become a naturalist. Among other compliments, Linnaeus has been called Princeps botanicorum, The Pliny of the North and he is also considered as one of the founders of modern ecology. In botany, the abbreviation used to indicate Linnaeus as the authority for species names is L. In older publications, sometimes the abbreviation Linn. is found, Linnæus was born in the village of Råshult in Småland, Sweden, on 23 May 1707. He was the first child of Nicolaus Ingemarsson and Christina Brodersonia and his siblings were Anna Maria Linnæa, Sofia Juliana Linnæa, Samuel Linnæus, and Emerentia Linnæa. One of a line of peasants and priests, Nils was an amateur botanist, a Lutheran minister. Christina was the daughter of the rector of Stenbrohult, Samuel Brodersonius, a year after Linnæus birth, his grandfather Samuel Brodersonius died, and his father Nils became the rector of Stenbrohult. The family moved into the rectory from the curates house, even in his early years, Linnæus seemed to have a liking for plants, flowers in particular. Whenever he was upset, he was given a flower, which calmed him. Nils spent much time in his garden and often showed flowers to Linnaeus, soon Linnæus was given his own patch of earth where he could grow plants
11.
Species
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In biology, a species is the basic unit of biological classification and a taxonomic rank. A species is defined as the largest group of organisms in which two individuals can produce fertile offspring, typically by sexual reproduction. While this definition is often adequate, looked at more closely it is problematic, for example, with hybridisation, in a species complex of hundreds of similar microspecies, or in a ring species, the boundaries between closely related species become unclear. Other ways of defining species include similarity of DNA, morphology, all species are given a two-part name, a binomial. The first part of a binomial is the genus to which the species belongs, the second part is called the specific name or the specific epithet. For example, Boa constrictor is one of four species of the Boa genus, Species were seen from the time of Aristotle until the 18th century as fixed kinds that could be arranged in a hierarchy, the great chain of being. In the 19th century, biologists grasped that species could evolve given sufficient time, Charles Darwins 1859 book The Origin of Species explained how species could arise by natural selection. Genes can sometimes be exchanged between species by horizontal transfer, and species may become extinct for a variety of reasons. In his biology, Aristotle used the term γένος to mean a kind, such as a bird or fish, a kind was distinguished by its attributes, for instance, a bird has feathers, a beak, wings, a hard-shelled egg, and warm blood. A form was distinguished by being shared by all its members, Aristotle believed all kinds and forms to be distinct and unchanging. His approach remained influential until the Renaissance, when observers in the Early Modern period began to develop systems of organization for living things, they placed each kind of animal or plant into a context. Many of these early delineation schemes would now be considered whimsical, animals likewise that differ specifically preserve their distinct species permanently, one species never springs from the seed of another nor vice versa. In the 18th century, the Swedish scientist Carl Linnaeus classified organisms according to shared physical characteristics and he established the idea of a taxonomic hierarchy of classification based upon observable characteristics and intended to reflect natural relationships. At the time, however, it was widely believed that there was no organic connection between species, no matter how similar they appeared. However, whether or not it was supposed to be fixed, by the 19th century, naturalists understood that species could change form over time, and that the history of the planet provided enough time for major changes. Jean-Baptiste Lamarck, in his 1809 Zoological Philosophy, described the transmutation of species, proposing that a species could change over time, in 1859, Charles Darwin and Alfred Russel Wallace provided a compelling account of evolution and the formation of new species. Darwin argued that it was populations that evolved, not individuals and this required a new definition of species. Darwin concluded that species are what appear to be, ideas
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Genus
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A genus is a taxonomic rank used in the biological classification of living and fossil organisms in biology. In the hierarchy of classification, genus comes above species. In binomial nomenclature, the name forms the first part of the binomial species name for each species within the genus. Felis catus and Felis silvestris are two species within the genus Felis, Felis is a genus within the family Felidae. The composition of a genus is determined by a taxonomist, the standards for genus classification are not strictly codified, so different authorities often produce different classifications for genera. Moreover, genera should be composed of units of the same kind as other genera. The term comes from the Latin genus, a noun form cognate with gignere, linnaeus popularized its use in his 1753 Species Plantarum, but the French botanist Joseph Pitton de Tournefort is considered the founder of the modern concept of genera. The scientific name of a genus may be called the name or generic epithet. It plays a role in binomial nomenclature, the system of naming organisms. The rules for the names of organisms are laid down in the Nomenclature Codes. The standard way of scientifically describing species and other lower-ranked taxa is by binomial nomenclature, the generic name forms its first half. For example, the gray wolfs binomial name is Canis lupus, with Canis being the name shared by the wolfs close relatives. The specific name is written in lower-case and may be followed by names in zoology or a variety of infraspecific names in botany. Especially with these names, when the generic name is known from context. Because animals are typically only grouped within subspecies, it is written as a trinomen with a third name. Dog breeds, meanwhile, are not scientifically distinguished, there are several divisions of plant species and therefore their infraspecific names generally include contractions explaining the relation. For example, the genus Hibiscus includes hundreds of other species apart from the Rose of Sharon or common garden hibiscus, Rose of Sharon doesnt have subspecies but has cultivars that carry desired traits, such as the bright white H. syriaca Diana. Hawaiian hibiscus, meanwhile, includes several separate species, since not all botanists agree on the divisions or names between species, it is common to specify the source of the name using author abbreviations
13.
Family (biology)
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In biological classification, family is one of the eight major taxonomic ranks, it is classified between order and genus. A family may be divided into subfamilies, which are intermediate ranks above the rank of genus. In vernacular usage, a family may be named one of its common members, for example, walnuts and hickory trees belong to the family Juglandaceae. What does or does not belong to a family—or whether a family should be recognized at all—are proposed and determined by practicing taxonomists. There are no rules for describing or recognizing a family. Taxonomists often take different positions about descriptions of taxa, and there may be no broad consensus across the community for some time. Some described taxa are accepted broadly and quickly, but others only rarely, if at all, the naming of families is codified by various international codes. In zoological nomenclature, the names of animals end with the suffix -idae. The concept of rank at time was not yet settled, and in the preface to the Prodromus Magnol spoke of uniting his families into larger genera. Carolus Linnaeus used the word familia in his Philosophia botanica to denote groups of plants, trees, herbs, ferns, palms. He used this term only in the section of the book. In zoology, the family as an intermediate between order and genus was introduced by Pierre André Latreille in his Précis des caractères génériques des insectes. He used families in some but not in all his orders of insects, families can be used for evolutionary, palaeontological and generic studies because they are more stable than lower taxonomic levels such as genera and species
14.
Evergreen
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In botany, an evergreen is a plant that has leaves throughout the year, always green. This contrasts with deciduous plants, which lose their foliage during the winter or dry season. There are many different kinds of plants, both trees and shrubs. Deciduous trees shed their leaves, usually as an adaptation to a cold or dry/wet season, evergreen trees do lose leaves, but each tree loses its leaves gradually and not all at once. Most warm temperate climate plants are also evergreen, in cool temperate climates, fewer plants are evergreen, with a predominance of conifers, as few evergreen broadleaf plants can tolerate severe cold below about −30 °C. In areas where there is a reason for being deciduous, being evergreen is usually an adaptation to low nutrient levels, deciduous trees lose nutrients whenever they lose their leaves. In warmer areas, species such as pines and cypresses grow on poor soils. In Rhododendron, a genus with many broadleaf evergreens, several species grow in forests but are usually found on highly acidic soil where the nutrients are less available to plants. In taiga or boreal forests, it is too cold for the matter in the soil to decay rapidly, so the nutrients in the soil are less easily available to plants. These conditions favour the growth of more evergreens and make it difficult for deciduous plants to persist. In addition, the shelter provided by existing evergreen plants can make it easier for younger evergreen plants to survive cold and/or drought
15.
Deciduous
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In a more general sense, deciduous means the dropping of a part that is no longer needed or falling away after its purpose is finished. In plants it is the result of natural processes, in botany and horticulture, deciduous plants, including trees, shrubs and herbaceous perennials, are those that lose all of their leaves for part of the year. In some cases leaf loss coincides with winter—namely in temperate or polar climates, in other parts of the world, including tropical, subtropical, and arid regions, plants lose their leaves during the dry season or other seasons, depending on variations in rainfall. The converse of deciduous is coniferous, where foliage is shed on a different schedule from deciduous trees, plants that are intermediate may be called semi-deciduous, they lose old foliage as new growth begins. Other plants are semi-evergreen and lose their leaves before the growing season. Many deciduous plants flower during the period when they are leafless, the absence of leaves improves wind transmission of pollen for wind-pollinated plants and increases the visibility of the flowers to insects in insect-pollinated plants. This strategy is not without risks, as the flowers can be damaged by frost or, in dry season regions, leaf drop or abscission involves complex physiological signals and changes within plants. The process of photosynthesis steadily degrades the supply of chlorophylls in foliage, the brightest leaf colors are produced when days grow short and nights are cool, but remain above freezing. These other pigments include carotenoids that are yellow, brown, anthocyanin pigments produce red and purple colors, though they are not always present in the leaves. Rather, they are produced in the foliage in late summer, parts of the world that have showy displays of bright autumn colors are limited to locations where days become short and nights are cool. In other parts of the world, the leaves of deciduous trees simply fall off without turning the bright colors produced from the accumulation of anthocyanin pigments, the beginnings of leaf drop starts when an abscission layer is formed between the leaf petiole and the stem. This layer is formed in the spring during active new growth of the leaf, the cells are sensitive to a plant hormone called auxin that is produced by the leaf and other parts of the plant. The elongation of cells break the connection between the different cell layers, allowing the leaf to break away from the plant. It also forms a layer that seals the break, so the plant does not lose sap, in the spring, these proteins are used as a nitrogen source during the growth of new leaves or flowers. Plants with deciduous foliage have advantages and disadvantages compared to plants with evergreen foliage, evergreens suffer greater water loss during the winter and they also can experience greater predation pressure, especially when small. Losing leaves in winter may reduce damage from insects, repairing leaves, removing leaves also reduces cavitation which can damage xylem vessels in plants. This then allows deciduous plants to have xylem vessels with larger diameters, the deciduous characteristic has developed repeatedly among woody plants. Trees include maple, many oaks and nothofagus, elm, aspen, Deciduous shrubs include honeysuckle, viburnum, and many others
16.
Temperate climate
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In geography, temperate or tepid latitudes of Earth lie between the tropics and the polar regions. The temperatures in these regions are relatively moderate, rather than extremely hot or cold. The north temperate zone extends from the Tropic of Cancer to the Arctic Circle, the south temperate zone extends from the Tropic of Capricorn to the Antarctic Circle. In some climate classifications, the zone is often divided into several smaller climate zones. These include Humid subtropical climate, Mediterranean climate, oceanic, subtropical climates are located between 23. 5° and 35. 0° north or south latitude on the eastern or leeward sides of landmasses. This climate has long, generally hot, summers and short, mild winters and these climates may occur in southern Asia, the southeastern United States, parts of eastern Australia, and in eastern coastal South America. Mediterranean climates, occur between 30° and 42° north and south latitude, on the sides of landmasses. This climate has hot summers and short mild winters, however, seasonal rainfall is the opposite of that of the subtropical humid type. These climates occur near the rimlands of the Mediterranean Sea, in western Australia, in California, the oceanic climates occur in the higher middle latitudes, between 45° and 60° north and south latitude. They are created by the flow from the cool high latitude oceans to their west. This causes the climate to have cool summers and cool winters, annual rainfall is spread throughout the entire year. Regions with this climate include Western Europe, northwestern North America, the Continental climates occur in middle latitudes, between 35° or 40° to 55°. These climates are normally inland or on sides of landmasses. They feature warm to hot summers and cold winters, with a large temperature variation. Regions with this climate include northern temperate Asia, the northern United States, southern Canada, the vast majority of the worlds human population resides in temperate zones, especially in the northern hemisphere, due to its greater mass of land. The richest temperate flora in the world is found in southern Africa, geographical zone Habitat Köppen climate classification Middle latitudes Polar circle Subtropics
17.
Subtropics
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The subtropics are geographic and climate zones located roughly between the tropic circle of latitude at 23.5 latitude and temperate latitudes. Subtropical climates are characterized by warm to hot summers and cool to mild winters with infrequent frost. Subtropical climates can occur at elevations within the tropics, such as in the southern end of the Mexican Plateau and in Vietnam. Six climate classifications use the term to define the various temperature. A great portion of the deserts are located within the subtropics. Within savanna regimes in the subtropics, a wet season is seen annually during the summer, within Mediterranean climate regimes, the wet season occurs during the winter. Areas bordering warm oceans are prone to heavy rainfall from tropical cyclones. Plants such as palms, citrus, mango, lychee, the tropics have been historically defined as lying between the Tropic of Cancer and Tropic of Capricorn, located at 23. 45° north and south latitude respectively. The poleward fringe of the subtropics is located at approximately 40° north and south latitude respectively, northern fringes of the type can go further north due to moderating effects of ocean streams, like in parts of Southern Europe due to heat transported by the Gulf Stream. Several methods have used to define the subtropical climate. In the Trewartha climate classification, a subtropical region should have at least eight months with a temperature greater than 10 °C. According to the Troll-Paffen climate classification, there exists one large subtropical zone named the warm-temperate subtropical zone. According to the E. Neef climate classification, the zone is divided into two parts, Rainy winters of the west sides and Eastern subtropical climate. According to the Wilhelm Lauer & Peter Frankenberg climate classification, the zone is divided into three parts, high-continental, continental, and maritime. According to the Siegmund/Frankenberg climate classification, subtropical is one of six zones in the world. Heating of the earth near the equator leads to large amounts of upward motion and convection along the trough or Intertropical convergence zone. The upper-level divergence over the near-equatorial trough leads to air rising and moving away from the equator aloft, as the air moves towards the Mid-Latitudes, it cools and sinks, which leads to subsidence near the 30th parallel of both hemispheres. This circulation is known as the Hadley cell and leads to the formation of the subtropical ridge, many of the worlds deserts are caused by these climatological high-pressure areas, located within the subtropics
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Laurel forest
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Laurel forest, also called laurisilva or laurissilva, is a type of subtropical forest found in areas with high humidity and relatively stable, mild temperatures. The forest is characterized by tree species with evergreen, glossy and elongated leaves. Plants from the family may or may not be present. Laurel forests are specific to wet forests from sea level to the highest mountains and they need an ecosystem of high humidity, such as cloud forests, with abundant rainfall throughout the year. Laurel forests typically occur on the slopes of tropical or subtropical mountains, some evergreen tree species will survive short frosts, but most species will not survive hard freezes and prolonged cool weather. They need a climate with annual temperature oscillation moderated by the proximity of the ocean. These conditions of temperature and moisture occur in four different geographical regions, along the western continental coasts between 35° and 50° latitude. On islands between 25° and 35° or 40° latitude, in humid montane regions of the tropics. Some laurel forests are a type of cloud forest, the moisture in the air condenses as rain or fog, creating a habitat characterized by cool, moist conditions in the air and soil. The resulting climate is wet and mild, with the oscillation of the temperature moderated by the proximity of the ocean. Laurel forests are characterized by evergreen and hardwood trees, reaching up to 40 metres in height, Laurel forest, laurisilva, and laurissilva all refer to plant communities that resemble the laurel bay. Some species belong to the laurel family or Lauraceae. As in any other rainforest, plants of the forests must adapt to high rainfall. The trees adapted in response to these drivers by developing analogous structures, leaves that repel water. The scientific names laurina, laurifolia, laurophylla, lauriformis, the term Lucidophyll, referring to the shiny surface of the leaves, was proposed in 1969 by Tatuo Kira. The term lauroid is also applied to climbing plants such as ivies whose leaves resemble those of the Lauraceae, mature laurel forests typically have a dense tree canopy and low light levels at the forest floor. Some forests are characterized by an overstory of emergent trees, Laurel forests are typically multi-species, and diverse in both the number of species and the genera and families represented. This species diversity contrasts with other temperate forest types, which typically have a canopy dominated by one or a few species, species diversity generally increases towards the tropics
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Tree
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In botany, a tree is a perennial plant with an elongated stem, or trunk, supporting branches and leaves in most species. In some usages, the definition of a tree may be narrower, including only woody plants with secondary growth, plants that are usable as lumber or plants above a specified height. Trees are not a group but include a variety of plant species that have independently evolved a woody trunk. In looser senses, the palms, the tree ferns, bananas. Trees tend to be long-lived, some reaching several thousand years old, the tallest known tree, a coast redwood named Hyperion, stands 115.6 m high. Trees have been in existence for 370 million years and it is estimated that there are just over 3 trillion mature trees in the world. A tree typically has many secondary branches supported clear of the ground by the trunk and this trunk typically contains woody tissue for strength, and vascular tissue to carry materials from one part of the tree to another. For most trees it is surrounded by a layer of bark which serves as a protective barrier, below the ground, the roots branch and spread out widely, they serve to anchor the tree and extract moisture and nutrients from the soil. Above ground, the divide into smaller branches and shoots. The shoots typically bear leaves, which light energy and convert it into sugars by photosynthesis, providing the food for the trees growth. Flowers and fruit may also be present, but some trees, such as conifers, instead have pollen cones and seed cones, others, such as tree ferns, trees play a significant role in reducing erosion and moderating the climate. They remove carbon dioxide from the atmosphere and store large quantities of carbon in their tissues, trees and forests provide a habitat for many species of animals and plants. Tropical rainforests are one of the most biodiverse habitats in the world, trees provide shade and shelter, timber for construction, fuel for cooking and heating, and fruit for food as well as having many other uses. In parts of the world, forests are shrinking as trees are cleared to increase the amount of available for agriculture. Because of their longevity and usefulness, trees have always revered, with sacred groves in various cultures. Although tree is a term of common parlance, there is no universally recognised precise definition of what a tree is, either botanically or in common language. In its broadest sense, a tree is any plant with the form of an elongated stem, or trunk. Trees are also defined by height, with smaller plants from 0.5 to 10 m being called shrubs
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Type species
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A similar concept is used for suprageneric groups called a type genus. In botanical nomenclature, these terms have no standing under the code of nomenclature. In botany, the type of a name is a specimen which is also the type of a species name. The species name that has that type can also be referred to as the type of the genus name, names of genus and family ranks, the various subdivisions of those ranks, and some higher-rank names based on genus names, have such types. In bacteriology, a species is assigned for each genus. Every named genus or subgenus in zoology, whether or not currently recognized as valid, is associated with a type species. In practice, however, there is a backlog of untypified names defined in older publications when it was not required to specify a type, a type species is both a concept and a practical system that is used in the classification and nomenclature of animals. The type species represents the species and thus definition for a particular genus name. In the Glossary, type species is defined as The nominal species that is the type of a nominal genus or subgenus. The type species permanently attaches a formal name to a genus by providing just one species within that genus to which the name is permanently linked. The species name in turn is fixed, in theory, to a type specimen, for example, the type species for the land snail genus Monacha is Monacha cartusiana. That genus is placed within the family Hygromiidae. The type genus for that family is the genus Hygromia, the concept of the type species in zoology was introduced by Pierre André Latreille. The International Code of Zoological Nomenclature states that the name of the type species should always be cited. It gives an example in Article 67.1, Astacus marinus Fabricius,1775 was later designated as the type species of the genus Homarus, thus giving it the name Homarus marinus. However, the species of Homarus should always be cited using its original name. 3, the type of the genus name Elodes is quoted as the type of the species name Hypericum aegypticum, Glossary of scientific naming Genetypes – genetic sequence data from type specimens. Holotype Paratype Principle of Typification Type Type genus
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Dioecy
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Dioecy is a characteristic of a species, meaning that it has distinct male and female individual organisms or colonies, meaning that a colony contains only either male or female individuals. Dioecy is one method that excludes self-fertilization and promotes allogamy, flowering plants have several other methods of excluding self-fertilization, called Self-incompatibility. In zoology, dioecious species may be opposed to hermaphroditic species, meaning that an individual is of one sex. Dioecy may also describe colonies, such as the colonies of Siphonophorae, Dioecious colonies contain members of only one sex, whereas monoecious colonies contain members of both sexes. Dioecious species have the male and female structures on separate plants. The meaning of male and female in the context of plants is different from that used in animal groups and this issue is discussed below in the Alternation of generations section. About 6 percent of species are entirely dioecious and about 7% of angiosperm genera contain some dioecious species. Examples of dioecious plant species include ginkgos, willows, cannabis, more examples of dioecious plants can be found in the Wikipedia category, Dioecious plants. Dioecy occurs in a variety of plant families. However it is common in woody plants and heterotrophic species. There are several alternatives to dioecy for sexual reproductive structure organization in plants including bisexual flowers and these are described at Plant reproductive morphology. Land plants have alternation of generations, for which the sporophyte produces spores rather than gametes. Strictly speaking, the sporophytes of plants do not have either male or female reproductive organs. The gametophytes of flowering plants are of a sex, the male gametophytes are contained within the pollen. The sporophyte generation is called dioecious when each sporophyte has only one kind of spore-producing organ whose spores ultimately give rise to all male gametes or all female gametes. Slightly different terms, dioicous and monoicous, may be used for the gametophyte generation, the definition avoids reference to male and female reproductive structures, which are rare in fungi. An individual of a fungal species not only requires a partner for mating. A monoecious fungal species can perform roles, but may not be self-compatible
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Flower
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A flower, sometimes known as a bloom or blossom, is the reproductive structure found in plants that are floral. The biological function of a flower is to effect reproduction, usually by providing a mechanism for the union of sperm with eggs, Flowers may facilitate outcrossing or allow selfing. Some flowers produce diaspores without fertilization, Flowers contain sporangia and are the site where gametophytes develop. Many flowers have evolved to be attractive to animals, so as to them to be vectors for the transfer of pollen. After fertilization, the ovary of the flower develops into fruit containing seeds, the essential parts of a flower can be considered in two parts, the vegetative part, consisting of petals and associated structures in the perianth, and the reproductive or sexual parts. A stereotypical flower consists of four kinds of structures attached to the tip of a short stalk, each of these kinds of parts is arranged in a whorl on the receptacle. The four main whorls are as follows, Collectively the calyx, corolla, the next whorl toward the apex, composed of units called petals, which are typically thin, soft and colored to attract animals that help the process of pollination. Androecium, the whorl, consisting of units called stamens. Stamens consist of two parts, a called a filament, topped by an anther where pollen is produced by meiosis. Gynoecium, the innermost whorl of a flower, consisting of one or more units called carpels, the carpel or multiple fused carpels form a hollow structure called an ovary, which produces ovules internally. Ovules are megasporangia and they in turn produce megaspores by meiosis which develop into female gametophytes and these give rise to egg cells. The gynoecium of a flower is described using an alternative terminology wherein the structure one sees in the innermost whorl is called a pistil. A pistil may consist of a carpel or a number of carpels fused together. The sticky tip of the pistil, the stigma, is the receptor of pollen, the supportive stalk, the style, becomes the pathway for pollen tubes to grow from pollen grains adhering to the stigma. The relationship to the gynoecium on the receptacle is described as hypogynous, perigynous, although the arrangement described above is considered typical, plant species show a wide variation in floral structure. These modifications have significance in the evolution of flowering plants and are used extensively by botanists to establish relationships among plant species, the four main parts of a flower are generally defined by their positions on the receptacle and not by their function. Many flowers lack some parts or parts may be modified into other functions and/or look like what is typically another part, in some families, like Ranunculaceae, the petals are greatly reduced and in many species the sepals are colorful and petal-like. Other flowers have modified stamens that are petal-like, the flowers of Peonies and Roses are mostly petaloid stamens
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Fruit
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In botany, a fruit is the seed-bearing structure in flowering plants formed from the ovary after flowering. Fruits are the means by which angiosperms disseminate seeds, accordingly, fruits account for a substantial fraction of the worlds agricultural output, and some have acquired extensive cultural and symbolic meanings. On the other hand, in usage, fruit includes many structures that are not commonly called fruits, such as bean pods, corn kernels, tomatoes. The section of a fungus that produces spores is called a fruiting body. Many common terms for seeds and fruit do not correspond to the botanical classifications, however, in botany, a fruit is the ripened ovary or carpel that contains seeds, a nut is a type of fruit and not a seed, and a seed is a ripened ovule. Examples of culinary vegetables and nuts that are botanically fruit include corn, cucurbits, eggplant, legumes, sweet pepper, in addition, some spices, such as allspice and chili pepper, are fruits, botanically speaking. g. Botanically, a grain, such as corn, rice, or wheat, is also a kind of fruit. However, the wall is very thin and is fused to the seed coat. The outer, often edible layer, is the pericarp, formed from the ovary and surrounding the seeds, the pericarp may be described in three layers from outer to inner, the epicarp, mesocarp and endocarp. Fruit that bears a prominent pointed terminal projection is said to be beaked, a fruit results from maturation of one or more flowers, and the gynoecium of the flower forms all or part of the fruit. Inside the ovary/ovaries are one or more ovules where the megagametophyte contains the egg cell, after double fertilization, these ovules will become seeds. The ovules are fertilized in a process starts with pollination. After pollination, a tube grows from the pollen through the stigma into the ovary to the ovule, later the zygote will give rise to the embryo of the seed, and the endosperm mother cell will give rise to endosperm, a nutritive tissue used by the embryo. As the ovules develop into seeds, the ovary begins to ripen and the ovary wall, in some multiseeded fruits, the extent to which the flesh develops is proportional to the number of fertilized ovules. The pericarp is often differentiated into two or three distinct layers called the exocarp, mesocarp, and endocarp, in some fruits, especially simple fruits derived from an inferior ovary, other parts of the flower, fuse with the ovary and ripen with it. In other cases, the sepals, petals and/or stamens and style of the fall off. When such other floral parts are a significant part of the fruit, it is called an accessory fruit, since other parts of the flower may contribute to the structure of the fruit, it is important to study flower structure to understand how a particular fruit forms. There are three modes of fruit development, Apocarpous fruits develop from a single flower having one or more separate carpels
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Berry
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In everyday language, a berry is a small, pulpy, and often edible fruit. Berries are usually juicy, rounded, brightly colored, sweet or sour, common examples are strawberries, raspberries, blueberries, and red- and blackcurrants. In Britain, soft fruit is a term for such fruits. The scientific usage of the term differs from common usage. In scientific terminology, a berry is a fruit produced from the ovary of a flower in which the outer layer of the ovary wall develops into an edible fleshy portion. The definition includes many fruits that are not commonly known as berries, such as grapes, tomatoes, cucumbers, eggplants, fruits excluded by the botanical definition include strawberries, raspberries, and blackberries, which are aggregate fruits. A plant bearing berries is said to be bacciferous or baccate, while many berries are edible, some are poisonous to humans, such as deadly nightshade and pokeweed. Others, such as the white mulberry, red mulberry, and elderberry, are poisonous when unripe, berries are eaten worldwide and often used in jams, preserves, cakes or pies. The berry industry varies from country to country as do types of cultivated or growing in the wild. Berries have been valuable as a source for humans since before the start of agriculture. They were a staple for early hunter-gatherers for thousands of years. In time, humans learned to store berries so that they could be used in the winter and they may be made into fruit preserves, and among Native Americans, mixed with meat and fats as pemmican. Berries also began to be cultivated in Europe and other countries, in Japan, between the 10th and 18th centuries, the term ichibigo referred to many berry crops. The most widely cultivated berry of modern times, however, is the strawberry, the strawberry was mentioned by ancient Romans, who thought it had medicinal properties, but it was then not a staple of agriculture. Woodland strawberries began to be grown in French gardens in the 14th century, the musky-flavored strawberry began to be grown in European gardens in the late 16th century. Later, the Virginia strawberry was grown in Europe and the United States, the most commonly consumed strawberry, the garden strawberry, is an accidental hybrid of the Virginia strawberry and a Chilean variety Fragaria chiloensis. Soon after, Antoine Nicolas Duchesne began to study the breeding of strawberries and made several discoveries crucial to the science of plant breeding, such as the sexual reproduction of strawberry. A form of the berry is found in all the Germanic languages, for example, Old English berie compares with Old Saxon and Old High German beri
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Drupe
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In botany, a drupe is an indehiscent fruit in which an outer fleshy part surrounds a shell of hardened endocarp with a seed inside. These fruits usually develop from a carpel, and mostly from flowers with superior ovaries. Other fleshy fruits may have an enclosure that comes from the seed coat surrounding the seed. The term drupaceous is applied to a fruit which has the structure and texture of a drupe, the boundary between a drupe and a berry is not always clear. Thus some sources describe the fruit of species of the genus Persea, one definition of berry requires the endocarp to be less than 2 mm thick, other fruits with a stony endocarp being drupes. In marginal cases, terms such as drupaceous or drupe-like may be used, the term stone fruit can be a synonym for drupe or, more typically, it can mean just the fruit of the genus Prunus. Freestone refers to a drupe having a stone which can be removed from the flesh with ease, the flesh is not attached to the stone and does not need to be cut to free the stone. Freestone varieties of fruits are preferred for uses that require removal of the stone. Freestone plums are preferred for making homegrown prunes, and freestone sour cherries are preferred for making pies, clingstone refers to a drupe having a stone which cannot easily be removed from the flesh. The flesh is attached strongly to the stone and must be cut to free the stone, clingstone varieties of fruits in the genus Prunus are preferred as table fruit and for jams, because the flesh of clingstone fruits tends to be more tender and juicy throughout. Tryma is a term for such nut-like drupes that are difficult to categorize. Hickory nuts and walnuts in the Juglandaceae family grow within a husk, these fruits are technically drupes or drupaceous nuts. Many drupes, with their sweet, fleshy outer layer, attract the attention of animals as a food, and the plant population benefits from the resulting dispersal of its seeds. The endocarp is sometimes dropped after the part is eaten, but is often swallowed, passing through the digestive tract. This passage through the tract can reduce the thickness of the endocarp. The process is known as scarification, typical drupes include apricots, olives, peaches, plums, cherries, mangoes and amlas. The coconut is also a drupe, but the mesocarp is fibrous or dry, so this type of fruit is classified as a simple dry, unlike other drupes, the coconut seed is unlikely to be dispersed by being swallowed by fauna, due to its large size. It can, however, float extremely long distances across oceans, bramble fruits are aggregates of drupelets
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Seed
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A seed is an embryonic plant enclosed in a protective outer covering. The formation of the seed is part of the process of reproduction in seed plants, Seeds are the product of the ripened ovule, after fertilization by pollen and some growth within the mother plant. The embryo is developed from the zygote and the coat from the integuments of the ovule. Seed plants now dominate biological niches on land, from forests to both in hot and cold climates. The term seed also has a meaning that antedates the above—anything that can be sown, e. g. seed potatoes. In the case of sunflower and corn seeds, what is sown is the seed enclosed in a shell or husk, many structures commonly referred to as seeds are actually dry fruits. Plants producing berries are called baccate, sunflower seeds are sometimes sold commercially while still enclosed within the hard wall of the fruit, which must be split open to reach the seed. Different groups of plants have other modifications, the stone fruits have a hardened fruit layer fused to. Nuts are the one-seeded, hard-shelled fruit of plants with an indehiscent seed. Seeds are produced in several related groups of plants, and their manner of production distinguishes the angiosperms from the gymnosperms, angiosperm seeds are produced in a hard or fleshy structure called a fruit that encloses the seeds, hence the name. Some fruits have layers of hard and fleshy material. In gymnosperms, no special structure develops to enclose the seeds, however, the seeds do become covered by the cone scales as they develop in some species of conifer. Seed production in natural plant populations varies widely from year-to-year in response to weather variables, insects and diseases, over a 20-year period, for example, forests composed of loblolly pine and shortleaf pine produced from 0 to nearly 5 million sound pine seeds per hectare. Over this period, there were six bumper, five poor, and nine good seed crops, right after fertilization, the zygote is mostly inactive, but the primary endosperm divides rapidly to form the endosperm tissue. This tissue becomes the food the young plant will consume until the roots have developed after germination, after fertilization the ovules develop into the seeds. The ovule consists of a number of components, The funicle or seed stalk which attaches the ovule to the placenta and hence ovary or fruit wall, the nucellus, the remnant of the megasporangium and main region of the ovule where the megagametophyte develops. The micropyle, a pore or opening in the apex of the integument of the ovule where the pollen tube usually enters during the process of fertilization. The chalaza, the base of the ovule opposite the micropyle, the shape of the ovules as they develop often affects the final shape of the seeds
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Parthenogenesis
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Parthenogenesis is a natural form of asexual reproduction in which growth and development of embryos occur without fertilization. In animals, parthenogenesis means development of an embryo from an egg cell. In plants parthenogenesis is a component process of apomixis, gynogenesis and pseudogamy are closely related phenomena in which a sperm or pollen triggers the development of the egg cell into an embryo but makes no genetic contribution to the embryo. The rest of the cytology and genetics of these phenomena are identical to that of parthenogenesis. However, these species still use fertilization, parthenogenesis occurs naturally in many plants, some invertebrate animal species and a few vertebrates. This type of reproduction has been induced artificially in a few species including fish, normal egg cells form after meiosis and are haploid, with half as many chromosomes as their mothers body cells. Haploid individuals, however, are usually non-viable, and parthenogenetic offspring usually have the diploid chromosome number, depending on the mechanism involved in restoring the diploid number of chromosomes, parthenogenetic offspring may have anywhere between all and half of the mothers alleles. The offspring having all of the genetic material are called full clones. Full clones are usually formed without meiosis, if meiosis occurs, the offspring will get only a fraction of the mothers alleles. Parthenogenetic offspring in species that use either the XY or the X0 sex-determination system have two X chromosomes and are female. In species that use the ZW sex-determination system, they have either two Z chromosomes or two W chromosomes, or they could have one Z and one W chromosome, some species reproduce exclusively by parthenogenesis, while others can switch between sexual reproduction and parthenogenesis. The switch between sexuality and parthenogenesis in such species may be triggered by the season, or by a lack of males or by conditions that favour rapid population growth, in these species asexual reproduction occurs either in summer or as long as conditions are favourable. This is because in asexual reproduction a successful genotype can spread quickly without being modified by sex or wasting resources on male offspring who wont give birth, in times of stress, offspring produced by sexual reproduction may be fitter as they have new, possibly beneficial gene combinations. Many taxa with heterogony have within them species that have lost the sexual phase and are now completely asexual, many other cases of obligate parthenogenesis are found among polyploids and hybrids where the chromosomes cannot pair for meiosis. The production of offspring by parthenogenesis is referred to as thelytoky while the production of males by parthenogenesis is referred to as arrhenotoky. When unfertilized eggs develop into males and females, the phenomenon is called deuterotoky. Parthenogenesis can occur without meiosis through mitotic oogenesis, mature egg cells are produced by mitotic divisions, and these cells directly develop into embryos. In flowering plants, cells of the gametophyte can undergo this process, the offspring produced by apomictic parthenogenesis are full clones of their mother
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Poison
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In biology, poisons are substances that cause disturbances in organisms, usually by chemical reaction or other activity on the molecular scale, when an organism absorbs a sufficient quantity. The fields of medicine and zoology often distinguish a poison from a toxin, toxins are poisons produced by organisms in nature, and venoms are toxins injected by a bite or sting. The difference between venom and other poisons is the delivery method, industry, agriculture, and other sectors use poisons for reasons other than their toxicity. Pesticides are one group of substances whose toxicity to various insects, in 2013,3.3 million cases of unintentional poisonings occurred. This resulted in 98,000 deaths worldwide, down from 120,000 deaths in 1990, the use of poison as an adjective dates from the 1520s. Using the word poison with plant names dates from the 18th century, the term poison ivy, for example, was first used in 1784 and the term poison oak was first used in 1743. The term poison gas was first used in 1915, paracelsus, the father of toxicology, once wrote, Everything is poison, there is poison in everything. Only the dose makes a thing not a poison, the term poison is also used in a figurative sense, His brothers presence poisoned the atmosphere at the party. The law defines poison more strictly, substances not legally required to carry the label poison can also cause a medical condition of poisoning. Some poisons are also toxins, which is any poison produced by animals, vegetables or bacterium, such as the proteins that cause tetanus. A distinction between the two terms is not always observed, even among scientists, the derivative forms toxic and poisonous are synonymous. Animal poisons delivered subcutaneously are also called venom, in normal usage, a poisonous organism is one that is harmful to consume, but a venomous organism uses venom to kill its prey or defend itself while still alive. A single organism can be poisonous and venomous, but that is rare. Human antimicrobial peptides which are toxic to viruses, fungi, bacteria, in nuclear physics, a poison is a substance that obstructs or inhibits a nuclear reaction. For an example, see nuclear poison, environmentally hazardous substances are not necessarily poisons, and vice versa. Biologically speaking, any substance, if given in large amounts, is poisonous. For instance, several kilograms worth of water would constitute a lethal dose, many substances used as medications—such as fentanyl—have an LD50 only one order of magnitude greater than the ED50. An alternative classification distinguishes between lethal substances that provide a value and those that do not
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Quercus ilex
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Quercus ilex, the evergreen oak, holly oak or holm oak, is a large evergreen oak native to the Mediterranean region. It takes its name from holm, an ancient name for holly and it is a member of the white oak section of the genus, with acorns that mature in a single summer. The first trees to be grown from acorns in England are still to be found within the grounds of Mamhead Park. From Britton & Brayley The Beauties of England and Wales, The resemblance of the foliage to that of the common European holly Ilex aquifolium, has led to its common and botanic names. The name ilex was originally the classical Latin name for the holm oak, the trunk is sometimes over 6 m in girth. The young shoots are clothed with a close grey felt, fruits are produced one to three together on a short downy stalk, ripening the first season, the acorns usually 12–18 mm long in the UK, the cups with appressed, downy scales. There are two subspecies, Quercus ilex subsp, native in the north and east of the species range, from northern Iberia and France east to Greece. Leaves narrow, acorns 2 cm long, bitter tasting, native in the southwest of the species range, in central and southern Iberia and northwest Africa. Leaves broader, acorns 2.5 cm long, sweet tasting, Holm oak grows in pure stands or mixed forest in the Mediterranean and often at low or moderate elevations, such as coastal California. One of the plant associations in which oak is found is the holm oak/Atlas cedar forests of the Atlas Mountains. In Morocco, some of mixed forests are habitat to the endangered primate, Barbary macaque. Holm oak is prevalent from Portugal to Italy along the northern Mediterranean coastal belt, Holm oak is damaging biodiversity in the United Kingdom, and is listed as an alien invader. Normally the tree is unable to withstand severe frost, which would prevent it from spreading north, the wood is hard and tough, and has been used since ancient times for general construction purposes as pillars, tools, wagons, vessels, and wine casks. It is also used as firewood, or in charcoal manufacture, the holm oak is one of the top three trees used in the establishment of truffle orchards, or truffières. Truffles grow in an association with the trees roots. The acorns, like those of the oak, are edible. Boiled in water, the acorns can also be used as a treatment for wound disinfections. Q. ilex can be clipped to form a tall hedge and it forms a picturesque rounded head, with pendulous low-hanging branches
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Old English
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Old English or Anglo-Saxon is the earliest historical form of the English language, spoken in England and southern and eastern Scotland in the early Middle Ages. It was brought to Great Britain by Anglo-Saxon settlers probably in the mid 5th century, Old English developed from a set of Anglo-Frisian or North Sea Germanic dialects originally spoken by Germanic tribes traditionally known as the Angles, Saxons, and Jutes. As the Anglo-Saxons became dominant in England, their language replaced the languages of Roman Britain, Common Brittonic, a Celtic language, Old English had four main dialects, associated with particular Anglo-Saxon kingdoms, Mercian, Northumbrian, Kentish and West Saxon. It was West Saxon that formed the basis for the standard of the later Old English period, although the dominant forms of Middle. The speech of eastern and northern parts of England was subject to strong Old Norse influence due to Scandinavian rule, Old English is one of the West Germanic languages, and its closest relatives are Old Frisian and Old Saxon. Like other old Germanic languages, it is different from Modern English. Old English grammar is similar to that of modern German, nouns, adjectives, pronouns, and verbs have many inflectional endings and forms. The oldest Old English inscriptions were using a runic system. Old English was not static, and its usage covered a period of 700 years, from the Anglo-Saxon settlement of Britain in the 5th century to the late 11th century, some time after the Norman invasion. While indicating that the establishment of dates is a process, Albert Baugh dates Old English from 450 to 1150, a period of full inflections. Perhaps around 85 per cent of Old English words are no longer in use, Old English is a West Germanic language, developing out of Ingvaeonic dialects from the 5th century. It came to be spoken over most of the territory of the Anglo-Saxon kingdoms which became the Kingdom of England and this included most of present-day England, as well as part of what is now southeastern Scotland, which for several centuries belonged to the Anglo-Saxon kingdom of Northumbria. Other parts of the island – Wales and most of Scotland – continued to use Celtic languages, Norse was also widely spoken in the parts of England which fell under Danish law. Anglo-Saxon literacy developed after Christianisation in the late 7th century, the oldest surviving text of Old English literature is Cædmons Hymn, composed between 658 and 680. There is a corpus of runic inscriptions from the 5th to 7th centuries. The Old English Latin alphabet was introduced around the 9th century, with the unification of the Anglo-Saxon kingdoms by Alfred the Great in the later 9th century, the language of government and literature became standardised around the West Saxon dialect. In Old English, typical of the development of literature, poetry arose before prose, a later literary standard, dating from the later 10th century, arose under the influence of Bishop Æthelwold of Winchester, and was followed by such writers as the prolific Ælfric of Eynsham. This form of the language is known as the Winchester standard and it is considered to represent the classical form of Old English
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French language
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French is a Romance language of the Indo-European family. It descended from the Vulgar Latin of the Roman Empire, as did all Romance languages, French has evolved from Gallo-Romance, the spoken Latin in Gaul, and more specifically in Northern Gaul. Its closest relatives are the other langues doïl—languages historically spoken in northern France and in southern Belgium, French was also influenced by native Celtic languages of Northern Roman Gaul like Gallia Belgica and by the Frankish language of the post-Roman Frankish invaders. Today, owing to Frances past overseas expansion, there are numerous French-based creole languages, a French-speaking person or nation may be referred to as Francophone in both English and French. French is a language in 29 countries, most of which are members of la francophonie. As of 2015, 40% of the population is in Europe, 35% in sub-Saharan Africa, 15% in North Africa and the Middle East, 8% in the Americas. French is the fourth-most widely spoken mother tongue in the European Union, 1/5 of Europeans who do not have French as a mother tongue speak French as a second language. As a result of French and Belgian colonialism from the 17th and 18th century onward, French was introduced to new territories in the Americas, Africa, most second-language speakers reside in Francophone Africa, in particular Gabon, Algeria, Mauritius, Senegal and Ivory Coast. In 2015, French was estimated to have 77 to 110 million native speakers, approximately 274 million people are able to speak the language. The Organisation internationale de la Francophonie estimates 700 million by 2050, in 2011, Bloomberg Businessweek ranked French the third most useful language for business, after English and Standard Mandarin Chinese. Under the Constitution of France, French has been the language of the Republic since 1992. France mandates the use of French in official government publications, public education except in specific cases, French is one of the four official languages of Switzerland and is spoken in the western part of Switzerland called Romandie, of which Geneva is the largest city. French is the language of about 23% of the Swiss population. French is also a language of Luxembourg, Monaco, and Aosta Valley, while French dialects remain spoken by minorities on the Channel Islands. A plurality of the worlds French-speaking population lives in Africa and this number does not include the people living in non-Francophone African countries who have learned French as a foreign language. Due to the rise of French in Africa, the total French-speaking population worldwide is expected to reach 700 million people in 2050, French is the fastest growing language on the continent. French is mostly a language in Africa, but it has become a first language in some urban areas, such as the region of Abidjan, Ivory Coast and in Libreville. There is not a single African French, but multiple forms that diverged through contact with various indigenous African languages, sub-Saharan Africa is the region where the French language is most likely to expand, because of the expansion of education and rapid population growth
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Old Dutch
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In linguistics, Old Dutch or Old Low Franconian is the set of Franconian dialects spoken in the Low Countries during the Early Middle Ages, from around the 5th to the 12th century. Old Dutch is mostly recorded on fragmentary relics, and words have been reconstructed from Middle Dutch and it evolved into Middle Dutch around the 12th century. Before the advent of Old Dutch or any of the Germanic languages, the North Sea Germanic dialects were spoken in the whole of the coastal parts of the Netherlands and Belgium. It was largely replaced by Weser-Rhine Germanic dialects, spoken by the Salian Franks and it spread from northern Belgium and the southern Netherlands to the coast and evolved into Old Low Franconian or Old Dutch. It has, however, a North sea Germanic substrate, which is why some philologists put the language in that branch, linguists typically date this transition to around the 5th century. Old Dutch is divided in Old West Low Franconian and Old East Low Franconian, however, the divergence being that the latter shares more traits with neighboring historical forms of Central Franconian dialects such as Ripuarian and Moselle Franconian. Old English, Old Frisian, Old Saxon and Old Dutch share the subsumption of the Ingvaeonic nasal spirant law, much of the grammatical variation between Old Dutch and Old Saxon is similar to that between Old Dutch and Old High German. Old Dutch naturally evolved into Middle Dutch with some distinctions that approximate those found in most medieval West Germanic languages. The year 1150 CE is often cited as the time of the discontinuity, the most notable difference between Old and Middle Dutch is in a feature of speech known as vowel reduction. Round vowels in word-final syllables are rather frequent in Old Dutch, in Modern Dutch, recasting is necessary to form a coherent sentence. Old Dutch texts are rare and much more limited when compared to related languages like Old English. Most of the earliest texts written in the Netherlands were written in Latin rather than Old Dutch, some of these Latin texts, however, contained Old Dutch words interspersed with the Latin text. Also, it is hard to determine whether a text actually was written in Old Dutch, wad Several words that are known to have developed in the Netherlands before Old Dutch was spoken have been found, and are sometimes called Oudnederlands in a geographic sense. The oldest known example, wad, was mentioned in 108 AD by Tacitus. The word exclusively referred to the region and ground type that is now known as the Wadden Sea, however, since this word existed long before Old Dutch did, it cannot be considered part of its vocabulary, but rather of Proto-Germanic. Haþuþuwas ann kusjam loguns This sentence has been interpreted as of Haþuþewaz, I bestow upon the choosers of the swords. It was discovered on a sword sheath, excavated in 1996 in the Dutch village of Bergakker and is better described as Frankish than Old Dutch. The text however, does show the beginning of Old Dutch morphology, the word ann, found in the partially translated inscription is coined as the oldest Dutch by linguists Nicoline van der Sijs and Tanneke Schoonheim from Genootschap Onze Taal
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Old High German
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Old High German is the earliest stage of the German language, conventionally covering the period from around 700 to 1050. Coherent written texts do not appear until the half of the 8th century. There are, however, a number of Elder Futhark inscriptions dating to the 6th century, as well as single words, during the migration period, the Elbe Germanic tribes settled in what became Alamannia, the Duchy of Bavaria and the Kingdom of Lombardy. Old High German comprises the dialects of these groups which underwent the Second Sound Shift during the 6th Century, namely all of Elbe Germanic, in the south, the Langobards, who had settled in Northern Italy, maintained their dialect until their conquest by Charlemagne in 774. This area did not become German-speaking again until the German eastward expansion of the early 12th century, though there was some attempt at conquest, Old High German literacy is a product of the monasteries, notably at St. Gallen, Reichenau and Fulda. Its origins lie in the establishment of the German church by Boniface in the mid 8th century, einhard tells how Charlemagne himself ordered that the epic lays should be collected for posterity. It was the neglect or religious zeal of later generations that led to the loss of these records, thus, it was Charlemagnes weak successor, Louis the Pious, who destroyed his fathers collection of epic poetry on account of its pagan content. Hrabanus Maurus, a student of Alcuins and abbot at Fulda from 822, was an important advocate of the cultivation of German literacy, among his students were Walafrid Strabo and Otfrid of Weissenburg. Notker Labeo towards the end of the Old High German period was among the greatest stylists in the language, the main difference between Old High German and the West Germanic dialects from which it developed is that it underwent the High German consonant shift. This is generally dated approximately to the late 5th and early 6th centuries—hence dating its start to around 500, the result of this sound change is that the consonantal system of German remains different from all other West Germanic languages, including English and Low German. Grammatically, however, Old High German remained very similar to Old English, Old Dutch, by the mid 11th century the many different vowels found in unstressed syllables had all been reduced to /ə/. Since these vowels were part of the endings in the nouns and verbs. For these reasons,1050 is seen as the start of the Middle High German period, for this reason the dialects may be termed monastery dialects. It declined after the conquest of the Lombard Kingdom by the Franks in 774 and it is classified as Upper German on the basis of evidence of the Second Sound Shift. The continued existence of a West Frankish dialect in the Western, claims that this might have been the language of the Carolingian court or that it is attested in the Ludwigslied, whose presence in a French manuscript suggests bilingualism, are controversial. The charts show the vowel and consonant systems of the East Franconian dialect in the 9th century and this is the dialect of the monastery of Fulda, and specifically of the Old High German Tatian. Old High German had five long vowels and six phonemic short vowels. Both occurred in stressed and unstressed syllables, notes, All back vowels likely had front-vowel allophones as a result of Umlaut
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Low German
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Low German or Low Saxon is a West Germanic language spoken mainly in northern Germany and the eastern part of the Netherlands. It is descended from Old Saxon in its earliest form, as an Ingvaeonic language, Low German is quite distinct from the Irminonic languages like Standard German. It is closely related to Anglo-Frisian group of languages and more distantly to Dutch and this difference resulted from the High German consonant shift, with the Uerdingen and Benrath lines being two notable linguistic borders. Dialects of Low German are widely spoken in the area of the Netherlands and are written there with an orthography based on Standard Dutch orthography. Small portions of northern Hesse and northern Thuringia are traditionally Low Saxon-speaking too, historically, Low German was also spoken in formerly German parts of Poland as well as in East Prussia and the Baltic States of Estonia and Latvia. Under the name Low Saxon, there are speakers in the Dutch north-eastern provinces of Groningen, Drenthe, Stellingwerf, Overijssel, German speakers in this area fled the Red Army or were forcibly expelled after the border changes at the end of World War II. Today, there are still speakers outside Germany and the Netherlands to be found in the areas of present-day Poland. In some of these countries, the language is part of the Mennonite religion, there are Mennonite communities in Ontario, Saskatchewan, Alberta, British Columbia, Manitoba, and Minnesota which use Low German in their religious services and communities. The type of Low German spoken in communities and in the Midwest region of the United States has diverged since emigration. The survival of the language is tenuous in many places and has died out in places where assimilation has occurred. Mennonite colonies in Paraguay, Belize, and Chihuahua, Mexico have made Low German a co-official language of the community, in Germany, native speakers of Low German call it Platt, Plattdüütsch or Nedderdüütsch. In the Netherlands, native speakers refer to their language as dialect, plat, nedersaksies, or the name of their village, officially, Low German is called Niederdeutsch by the German authorities and Nedersaksisch by the Dutch authorities. Plattdeutsch/Niederdeutsch and Platduits/Nedersaksisch are seen in texts from the German. In Danish it is called Plattysk, Nedertysk or, rarely, Mennonite Low German is called Plautdietsch. Etymologically Platt meant clear in the sense of a language the people could understand. In Dutch, the word Plat can also mean improper, or rude, the ISO 639-2 language code for Low German has been nds since May 2000. The question of whether Low German should be considered a separate language, linguistics offers no simple, generally accepted criterion to decide this question. Scholarly arguments have been put forward in favour of classifying Low German as a German dialect, as said, these arguments are not linguistic but rather socio-political and build mainly around the fact that Low German has no official standard form or use in sophisticated media
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Low Franconian
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The Frankish language, also Old Frankish, was the language of the Franks. It is a West Germanic language and was spoken in Merovingian times, the Franks first established themselves in the Netherlands and Flanders before they started to fight their way down south and east. The language had a significant impact on Old French and it evolved into Old Dutch in the north and it was replaced step by step by the langues doïl in the south. Old Frankish is not directly attested except in glosses and small phrases and it has been reconstructed using the comparative method from loanwords in Old French and from Old Dutch. One known phrase in Old Frankish is found in the Salic law of the sixth century. It was a language of the Old Frankish language. Old Low Franconian is sometimes divided in two groups, Old Dutch and Old East Low Franconian, Dutch, like other Germanic languages, is conventionally divided into three phases. During the 19th and 20th centuries these dialects have partly and gradually replaced by todays Standard German. Sometimes, Low Franconian is grouped together with Low German, the contemporary continental Low Franconian language area is decreasing in size. French Flanders has become more and more francophone during the last century, brussels Capital Region is officially bilingual, but largely francophone. In Germany, Low Franconian only exists as Meuse-Rhenish regiolects and dialects, at the Rhine near Duisburg, it adjoins a smaller strip of other Low Franconian varieties called Bergisch. Together these distinct varieties, now combined with the Kleve dialects as Meuse-Rhenish. Limburgish straddles the borderline between Low Franconian and Middle Franconian varieties and they are more-or-less mutually intelligible with the Ripuarian dialects, but show fewer High German shifts. Afrikaans originated from the Dutch language, the dialect became known as Cape Dutch. Later, Afrikaans was sometimes referred to as African Dutch or Kitchen Dutch. Dutch remained an official language until the new 1961 constitution finally stipulated the two languages in South Africa to be Afrikaans and English. It is the only Indo-European language of significance that underwent distinct development on the African continent, the South African Constitution of post 1994 considers Afrikaans as one of 11 official languages. The majority of Afrikaans speakers in South Africa are coloureds of mixed European/Black African/Khoisan/Indonesian descent, Afrikaans Dutch dialects Dutch language Franconian languages Istvaeones History of Dutch Low Dietsch Meuse-Rhenish Middle Dutch Euler, Wolfram
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Germanic languages
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It is the third most spoken Indo-European subdivision, behind Italic and Indo-Iranian, and ahead of Balto-Slavic languages. Limburgish varieties have roughly 1.3 million speakers along the Dutch–Belgian–German border, the main North Germanic languages are Norwegian, Danish, Swedish, Icelandic, and Faroese, which have a combined total of about 20 million speakers. The East Germanic branch included Gothic, Burgundian, and Vandalic, the last to die off was Crimean Gothic, spoken in the late 18th century in some isolated areas of Crimea. The total number of Germanic languages throughout history is unknown, as some of them—especially East Germanic languages—disappeared during or after the Migration Period. Proto-Germanic, along all of its descendants, is characterized by a number of unique linguistic features. Early varieties of Germanic enter history with the Germanic tribes moving south from Scandinavia in the 2nd century BC, to settle in the area of todays northern Germany, furthermore, it is the de facto language of the United Kingdom, the United States and Australia. It is also a language in Nicaragua and Malaysia. German is a language of Austria, Belgium, Germany, Liechtenstein, Luxembourg and Switzerland and has regional status in Italy, Poland, Namibia. German also continues to be spoken as a minority language by immigrant communities in North America, South America, Central America, Mexico, a German dialect, Pennsylvania Dutch, is still present amongst Anabaptist populations in Pennsylvania in the United States. Dutch is a language of Aruba, Belgium, Curaçao. The Netherlands also colonised Indonesia, but Dutch was scrapped as a language after Indonesian independence. Dutch was until 1925 an official language in South Africa, but evolved in and was replaced by Afrikaans, Afrikaans is one of the 11 official languages in South Africa and is a lingua franca of Namibia. It is used in other Southern African nations as well, low German is a collection of sometimes very diverse dialects spoken in the northeast of the Netherlands and northern Germany. Scots is spoken in Lowland Scotland and parts of Ulster, frisian is spoken among half a million people who live on the southern fringes of the North Sea in the Netherlands, Germany, and Denmark. Luxembourgish is mainly spoken in the Grand Duchy of Luxembourg, though it extends into small parts of Belgium, France. Limburgish varieties are spoken in the Limburg and Rhineland regions, along the Dutch–Belgian–German border, Swedish is also one of the two official languages in Finland, along with Finnish, and the only official language in the Åland Islands. Danish is also spoken natively by the Danish minority in the German state of Schleswig-Holstein, Norwegian is the official language of Norway. Icelandic is the language of Iceland, and is spoken by a significant minority in the Faroe Islands
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Celtic languages
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The Celtic languages are descended from Proto-Celtic, or Common Celtic, a branch of the greater Indo-European language family. Modern Celtic languages are spoken on the north-western edge of Europe, notably in Ireland, Scotland, Wales, Brittany, Cornwall. There are also a number of Welsh speakers in the Patagonia area of Argentina. Some people speak Celtic languages in the other Celtic diaspora areas of the United States, Canada, Australia, in all these areas, the Celtic languages are now only spoken by minorities though there are continuing efforts at revitalisation. Welsh is the only Celtic language not classified as endangered by UNESCO, the spread to Cape Breton and Patagonia occurred in modern times. SIL Ethnologue lists six living Celtic languages, of four have retained a substantial number of native speakers. These are the Goidelic languages and the Brittonic languages, the other two, Cornish and Manx, died in modern times with their presumed last native speakers in 1777 and 1974 respectively. For both these languages, however, revitalisation movements have led to the adoption of these languages by adults and children, taken together, there were roughly one million native speakers of Celtic languages as of the 2000s. In 2010, there were more than 1.4 million speakers of Celtic languages, shelta, based largely on Irish with influence from an undocumented source. Some forms of Welsh-Romani or Kååle also combined Romany itself with Welsh language, beurla-reagaird, Highland travellers language Celtic divided into various branches, Lepontic, the oldest attested Celtic language. Anciently spoken in Switzerland and in Northern-Central Italy, from the Alps to Umbria, coins with Lepontic inscriptions have been found in Noricum and Gallia Narbonensis. Celtiberian, anciently spoken in the Iberian peninsula, in parts of modern Galicia, Asturias, La Rioja, Aragón, Cantabria, Old Castile, the relationship of Celtiberian with Gallaecian, in the northwest of the peninsula, is uncertain. Gallaecian, anciently spoken in the former Gallaecia, northwest of the peninsula, Gaulish languages, including Galatian and possibly Noric. These languages were spoken in a wide arc from Belgium to Turkey. Brittonic, including the living languages Breton, Cornish, and Welsh, before the arrival of Scotti on the Isle of Man in the 9th century, there may have been a Brittonic language in the Isle of Man. Goidelic, including the living languages Irish, Manx, and Scottish Gaelic, scholarly handling of the Celtic languages has been rather argumentative owing to scarceness of primary source data. Other scholars distinguish between P-Celtic and Q-Celtic, putting most of the Gaulish and Brittonic languages in the former group, the P-Celtic languages are sometimes seen as a central innovating area as opposed to the more conservative peripheral Q-Celtic languages. In the P/Q classification schema, the first language to split off from Proto-Celtic was Gaelic and it has characteristics that some scholars see as archaic, but others see as also being in the Brittonic languages
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Welsh language
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Welsh is a member of the Brittonic branch of the Celtic languages. It is spoken natively in Wales, by some in England, historically it has also been known in English as the British tongue, Cambrian, Cambric and Cymric. The United Kingdom Census 2011 counted 3.1 million residents of Wales, 27% of whom had been born outside Wales, and 73% of whom reported having no Welsh language skills. Of residents of Wales aged three and over, 19% reported being able to speak Welsh, and 77% of these were able to speak, read and this can be compared with the 2001 Census, in which 20. 8% of the population reported being able to speak Welsh. 787,854 of residents in Wales aged three and over had one or more skills in Welsh, in surveys carried out between 2004 and 2006, 57% of Welsh speakers described themselves as fluent in the written language. An estimated 110,000 to 150,000 people speak Welsh in England, Welsh emerged in the 6th century from Common Brittonic, the common ancestor of Welsh, Breton, Cornish and the extinct language known as Cumbric. The Middle Welsh period is considered to have lasted from then until the 14th century, when the Modern Welsh period began, the name Welsh originated as an exonym given to its speakers by the Anglo-Saxons, meaning foreign speech. The native term for the language is Cymraeg, and for the name of the country of Wales it is Cymru, Welsh evolved from Common Brittonic, the Celtic language spoken by the ancient Celtic Britons. Classified as Insular Celtic, the British language probably arrived in Britain during the Bronze Age or Iron Age and was spoken throughout the island south of the Firth of Forth. During the Early Middle Ages the British language began to fragment due to increased dialect differentiation, evolving into Welsh and it is not clear when Welsh became distinct. Kenneth H. Jackson suggested that the evolution in syllabic structure and sound pattern was complete by around 550, Jackson, however, believed that the two varieties were already distinct by that time. The earliest Welsh poetry – that attributed to the Cynfeirdd or Early Poets – is generally considered to date to the Primitive Welsh period. However, much of this poetry was composed in the Hen Ogledd, raising further questions about the dating of the material. An 8th century inscription in Tywyn shows the language already dropping inflections in the declension of nouns, the next main period, somewhat better attested, is Old Welsh, poetry from both Wales and Scotland has been preserved in this form of the language. Both the works of Aneirin and the Book of Taliesin were in this era, Middle Welsh is the label attached to the Welsh of the 12th to 14th centuries, of which much more remains than for any earlier period. This is the language of all surviving early manuscripts of the Mabinogion. It is also the language of the existing Welsh law manuscripts, Middle Welsh is reasonably intelligible, albeit with some work, to a modern-day Welsh speaker. The famous cleric Gerald of Wales tells, in his Descriptio Cambriae, during one of the Kings many raids in the 12th century, Henry asked an old man of Pencader, Carmarthenshire whether the Welsh people could resist his army
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Breton language
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Breton /ˈbrɛtən/ is a Southwestern Brittonic Celtic language spoken in Brittany. Breton is most closely related to Cornish, both being Southwestern Brittonic languages, Welsh and the extinct Cumbric are the more distantly related Brittonic languages. The other regional language of Brittany, Gallo, is a langue doïl, Gallo is consequently close to French, although not mutually intelligible, and a Romance language descended from Latin. However, the number of children attending bilingual classes has risen 33% between 2006 and 2012 to 14,709, Breton is spoken in West Brittany, roughly to the west of a line linking Plouha and La Roche-Bernard. It comes from a Brittonic language community that extended from Great Britain to Armorica and had even established a toehold in Galicia. Old Breton is attested from the 9th century and it was the language of the upper classes until the 12th century, after which it became the language of commoners in Lower Brittany. The nobility, followed by the bourgeoisie, adopted French, the written language of the Duchy of Brittany was Latin, switching to French in the 15th century. There exists a tradition of Breton literature. Some Old Breton vocabulary remains in the present day as philosophical, during the French Revolution, the government introduced policies favouring French over the regional languages, which it pejoratively referred to as patois. The revolutionaries assumed that reactionary and monarchist forces preferred regional languages to try to keep the peasant masses underinformed, in 1794, Bertrand Barère submitted his report on the patois to the Committee of Public Safety in which he said that federalism and superstition speak Breton. Teachers humiliated students for using their regional languages, and such practices prevailed until the late 1960s, the majority of todays speakers are more than 60 years old, and Breton is now classified as an endangered language. At the beginning of the 20th century, half of the population of Lower Brittany knew only Breton, by 1950, there were only 100,000 monolingual Bretons, and this rapid decline has continued, with likely no monolingual speakers left today. A statistical survey in 1997 found around 300,000 speakers in Lower Brittany, few 15- to 19-year-olds spoke Breton. In 1925, Professor Roparz Hemon founded the Breton-language review Gwalarn, during its 19-year run, Gwalarn tried to raise the language to the level of a great international language. Its publication encouraged the creation of literature in all genres. In 1946, Al Liamm replaced Gwalarn, other Breton-language periodicals have been published, which established a fairly large body of literature for a minority language. In 1977, Diwan schools were founded to teach Breton by immersion and they taught a few thousand young people from elementary school to high school. See the education section for more information, the Asterix comic series has been translated into Breton
40.
Irish language
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Irish, also referred to as Gaelic or Irish Gaelic, is a Goidelic language of the Indo-European language family originating in Ireland and historically spoken by the Irish people. Irish is spoken as a first language by a minority of Irish people. Irish enjoys constitutional status as the national and first official language of the Republic of Ireland and it is also among the official languages of the European Union. The public body Foras na Gaeilge is responsible for the promotion of the language throughout the island of Ireland and it has the oldest vernacular literature in Western Europe. The fate of the language was influenced by the power of the English state in Ireland. Elizabethan officials viewed the use of Irish unfavourably, as being a threat to all things English in Ireland and its decline began under English rule in the 17th century. In the latter part of the 19th century, there was a decrease in the number of speakers. Irish-speaking areas were hit especially hard, by the end of British rule, the language was spoken by less than 15% of the national population. Since then, Irish speakers have been in the minority, efforts have been made by the state, individuals and organisations to preserve, promote and revive the language, but with mixed results. Around the turn of the 21st century, estimates of native speakers ranged from 20,000 to 80,000 people. In the 2011 Census, these numbers had increased to 94,000 and 1.3 million, there are several thousand Irish speakers in Northern Ireland. It has been estimated that the active Irish-language scene probably comprises 5 to 10 per cent of Irelands population, there has been a significant increase in the number of urban Irish speakers, particularly in Dublin. In Gaeltacht areas, however, there has been a decline of the use of Irish. Údarás na Gaeltachta predicted that, by 2025, Irish will no longer be the language in any of the designated Gaeltacht areas. Survey data suggest that most Irish people think highly of Irish as a marker of identity. It has also argued that newer urban groups of Irish speakers are a disruptive force in this respect. In An Caighdeán Oifigiúil the name of the language is Gaeilge, before the spelling reform of 1948, this form was spelled Gaedhilge, originally this was the genitive of Gaedhealg, the form used in Classical Irish. Older spellings of this include Gaoidhealg in Classical Irish and Goídelc in Old Irish, the modern spelling results from the deletion of the silent dh in the middle of Gaedhilge, whereas Goidelic languages, used to refer to the language family including Irish, comes from Old Irish
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Romance languages
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Today, around 800 million people are native speakers worldwide, mainly in Europe, Africa and the Americas, but also elsewhere. Additionally, the major Romance languages have many speakers and are in widespread use as lingua francas. This is especially the case for French, which is in use throughout Central and West Africa, Madagascar, Mauritius. The five most widely spoken Romance languages by number of speakers are Spanish, Portuguese, French, Italian. Because of the difficulty of imposing boundaries on a continuum, various counts of the modern Romance languages are given, for example, Dalby lists 23 based on mutual intelligibility. Between 350 BC and 150 AD, the expansion of the Empire, together with its administrative and educational policies, Latin also exerted a strong influence in southeastern Britain, the Roman province of Africa, western Germany and the Balkans north of the Jireček Line. Despite other influences, the phonology, morphology, and lexicon of all Romance languages consist mainly of evolved forms of Vulgar Latin, however, some notable differences occur between todays Romance languages and their Roman ancestor. With only one or two exceptions, Romance languages have lost the system of Latin and, as a result, have SVO sentence structure. From this adverb the noun romance originated, which applied initially to anything written romanice, the word romance with the modern sense of romance novel or love affair has the same origin. In the medieval literature of Western Europe, serious writing was usually in Latin, while popular tales, often focusing on love, were composed in the vernacular, for example, the Portuguese word fresta is descended from Latin fenestra window, but now means skylight and slit. Cognates may exist but have become rare, such as finiestra in Spanish, the Spanish and Portuguese terms defenestrar meaning to throw through a window and fenestrado meaning replete with windows also have the same root, but are later borrowings from Latin. Galician has both fiestra and the frequently used ventá and xanela. As an alternative to lei, Italian has the pronoun ella, a cognate of the words for she. Sardinian balcone comes from Old Italian and is similar to other Romance languages such as French balcon, Portuguese balcão, Romanian balcon, Spanish balcón, Catalan balcó and Corsican balconi. Documentary evidence is limited about Vulgar Latin for the purposes of research. Many of its speakers were soldiers, slaves, displaced peoples, other scholars argue that the distinctions are more rightly viewed as indicative of sociolinguistic and register differences normally found within any language. Both were mutually intelligible as one and the language, which was true until the second half of the 7th century. Central Europe and the Balkans were occupied by the Germanic and Slavic tribes, as well as by the Huns, over the course of the fourth to eighth centuries, Vulgar Latin, by this time highly dialectalized, broke up into discrete languages that were no longer mutually intelligible
42.
Occitan
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Occitan, also known as lenga dòc by its native speakers, is a Romance language. It is spoken in southern France, Italys Occitan Valleys, Monaco, and Spains Val dAran, collectively, Occitan is also spoken in the linguistic enclave of Guardia Piemontese. However, there is controversy about the unity of the language, others include Catalan in this family, as the distance between this language and some Occitan dialects is similar to the distance among different Occitan dialects. In fact, Catalan was considered an Occitan dialect until the end of the 19th century, today, Occitan is an official language in Catalonia, where a subdialect of Gascon known as Aranese is spoken in the Val dAran. Since September 2010, the Parliament of Catalonia has considered Aranese Occitan to be the preferred language for use in the Val dAran. Unlike other Romance languages such as French or Spanish, there is no written standard language called Occitan. Instead, there are competing norms for writing Occitan, some of which attempt to be pan-dialectal, There are also significant lexical differences, where some dialects have words cognate with French, and others have Catalan and Spanish cognates. Nonetheless, there is a significant amount of mutual intelligibility, the long-term survival of Occitan is in grave doubt. According to the UNESCO Red Book of Endangered Languages, four of the six dialects of Occitan are considered severely endangered. The name Occitan comes from lenga dòc, òc being the Occitan word for yes and this was not, of course, the only defining characteristic of each group. The word òc came from Vulgar Latin hoc, while oïl originated from Latin hoc illud, Old Catalan, and now the Catalan of Northern Catalonia also have hoc. Other Romance languages derive their word for yes from the Latin sic, thus, etc. such as Spanish sí, Eastern Lombard sé, Sicilian and Italian sì, or Portuguese sim. French uses si to answer yes in response to questions that are asked in the negative sense, the name Occitan is sometimes considered a neologism, however, it was attested around 1300 as occitanus, a crossing of oc and aquitanus. For many centuries, the Occitan dialects were referred to as Limousin or Provençal, after Frédéric Mistrals Félibrige movement in the 19th century, Provençal achieved the greatest literary recognition and so became the most popular term for Occitan. The term first came into fashion in Italy, currently, linguists use the terms Provençal and Limousin strictly to refer to specific varieties within Occitania, keeping the name Occitan for the language as a whole. Many non-specialists, however, continue to refer to the language as Provençal, NO·L LI TOLRÀ NO·L LI DEVEDARÀ NI NO LEN DECEBRÀ. Nec societatem non AURÀ, si per castellum recuperare NON O FA, et si recuperare potuerit in potestate Froterio et Raimundo LO TORNARÀ, carolingian litanies, both written and sung in Latin, were answered to in Old Occitan by the audience. Occitan was the vehicle for the poetry of the medieval troubadours and trobairises, At that time
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Phylogeography
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Phylogeography is the study of the historical processes that may be responsible for the contemporary geographic distributions of individuals. This is accomplished by considering the geographic distribution of individuals in light of genetics and this term was introduced to describe geographically structured genetic signals within and among species. An explicit focus on a species biogeography/biogeographical past sets phylogeography apart from population genetics and phylogenetics. Past events that can be inferred include population expansion, population bottlenecks, vicariance, the term phylogeography was first used by John Avise in his 1987 work Intraspecific Phylogeography, The Mitochondrial DNA Bridge Between Population Genetics and Systematics. Historical biogeography addresses how historical geological, climatic and ecological conditions influenced the current distribution of species, as part of historical biogeography, researchers had been evaluating the geographical and evolutionary relationships of organisms years before. The resulting school of thought was vicariance biogeography, which explained the origin of new lineages through geological events like the drifting apart of continents or the formation of rivers, when a continuous population is divided by a new river or a new mountain range, two populations are created. Paleogeography, geology and paleoecology are all important fields that supply information that is integrated into phylogeographic analyses, Phylogeography takes a population genetics and phylogenetic perspective on biogeography. In the mid-1970s, population genetic analyses turned to mitochondrial markers, the advent of the polymerase chain reaction, the process where millions of copies of a DNA segment can be replicated, was crucial in the development of phylogeography. Thanks to this breakthrough, the contained in mitochondrial DNA sequences was much more accessible. Advances in both laboratory methods that allowed easier sequencing DNA and computational methods that make use of the data have helped improve phylogeographic inference. Early phylogeographic work has recently criticized for its narrative nature. Recent approaches have taken a stronger approach to phylogeography than was done initially. Example Climate change, such as the cycles of the past 2.4 million years, has periodically restricted some species into disjunct refugia. These restricted ranges may result in population bottlenecks that reduce genetic variation, once a reversal in climate change allows for rapid migration out of refugial areas, these species spread rapidly into newly available habitat. A number of studies find genetic signatures of both animal and plant species that support this scenario of refugia and postglacial expansion. This has occurred both in the tropics as well as regions that were directly influenced by glaciers. Phylogeography can help in the prioritization of areas of high value for conservation, using phylogeographical approaches, the authors found that hidden within what was thought to be a single, widely distributed species, an ancient and previously undetected species was also present. Conservation decisions can now be made to ensure that both lineages received protection, results like this are not an uncommon outcome from phylogeographic studies
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Speciation
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Speciation is the evolutionary process by which biological populations evolve to become distinct species. The biologist Orator F. Cook coined the term speciation in 1906 for the splitting of lineages or cladogenesis, Charles Darwin was the first to describe the role of natural selection in speciation in his 1859 book The Origin of Species. He also identified sexual selection as a mechanism, but found it problematic. There are four modes of speciation in nature, based on the extent to which speciating populations are isolated from one another, allopatric, peripatric, parapatric. Speciation may also be induced artificially, through animal husbandry, agriculture, whether genetic drift is a minor or major contributor to speciation is the subject matter of much ongoing discussion. All forms of speciation have taken place over the course of evolution, however. During allopatric speciation, a population splits into two isolated populations. When the populations come back into contact, they have evolved such that they are isolated and are no longer capable of exchanging genes. Island genetics is the associated with the tendency of small. Examples include insular dwarfism and the changes among certain famous island chains. The Galápagos Islands are particularly famous for their influence on Charles Darwin, though the finches were less important for Darwin, more recent research has shown the birds now known as Darwins finches to be a classic case of adaptive evolutionary radiation. In peripatric speciation, a subform of allopatric speciation, new species are formed in isolated and it is related to the concept of a founder effect, since small populations often undergo bottlenecks. Genetic drift is often proposed to play a significant role in peripatric speciation, parapatric speciation may be associated with differential landscape-dependent selection. Even if there is a gene flow between two populations, strong differential selection may impede assimilation and different species may eventually develop, habitat differences may be more important in the development of reproductive isolation than the isolation time. Ecologists refer to parapatric and peripatric speciation in terms of ecological niches, a niche must be available in order for a new species to be successful. Ring species such as Larus gulls have been claimed to illustrate speciation in progress, the grass Anthoxanthum odoratum may be starting parapatric speciation in areas of mine contamination. Sympatric speciation refers to the formation of two or more descendant species from an ancestral species all occupying the same geographic location. Often-cited examples of sympatric speciation are found in insects that become dependent on different host plants in the same area, however, the existence of sympatric speciation as a mechanism of speciation remains highly debated
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