1.
Megaannum
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A year is the orbital period of the Earth moving in its orbit around the Sun. Due to the Earths axial tilt, the course of a year sees the passing of the seasons, marked by changes in weather, the hours of daylight, and, consequently, vegetation and soil fertility. In temperate and subpolar regions around the globe, four seasons are recognized, spring, summer, autumn. In tropical and subtropical regions several geographical sectors do not present defined seasons, but in the seasonal tropics, a calendar year is an approximation of the number of days of the Earths orbital period as counted in a given calendar. The Gregorian, or modern, calendar, presents its calendar year to be either a common year of 365 days or a year of 366 days, as do the Julian calendars. For the Gregorian calendar the average length of the year across the complete leap cycle of 400 years is 365.2425 days. The ISO standard ISO 80000-3, Annex C, supports the symbol a to represent a year of either 365 or 366 days, in English, the abbreviations y and yr are commonly used. In astronomy, the Julian year is a unit of time, it is defined as 365.25 days of exactly 86400 seconds, totalling exactly 31557600 seconds in the Julian astronomical year. The word year is used for periods loosely associated with, but not identical to, the calendar or astronomical year, such as the seasonal year, the fiscal year. Similarly, year can mean the period of any planet, for example. The term can also be used in reference to any long period or cycle, west Saxon ġēar, Anglian ġēr continues Proto-Germanic *jǣran. Cognates are German Jahr, Old High German jār, Old Norse ár and Gothic jer, all the descendants of the Proto-Indo-European noun *yeh₁rom year, season. Cognates also descended from the same Proto-Indo-European noun are Avestan yārǝ year, Greek ὥρα year, season, period of time, Old Church Slavonic jarŭ, Latin annus is from a PIE noun *h₂et-no-, which also yielded Gothic aþn year. Both *yeh₁-ro- and *h₂et-no- are based on verbal roots expressing movement, *h₁ey- and *h₂et- respectively, the Greek word for year, ἔτος, is cognate with Latin vetus old, from the PIE word *wetos- year, also preserved in this meaning in Sanskrit vat-sa- yearling and vat-sa-ras year. Derived from Latin annus are a number of English words, such as annual, annuity, anniversary, etc. per annum means each year, anno Domini means in the year of the Lord. No astronomical year has an number of days or lunar months. Financial and scientific calculations often use a 365-day calendar to simplify daily rates, in the Julian calendar, the average length of a year is 365.25 days. In a non-leap year, there are 365 days, in a year there are 366 days
2.
Precambrian
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The Precambrian is the earliest period of Earths history, set before the current Phanerozoic Eon. The Precambrian is a supereon that is subdivided into three eons of the time scale. It spans from the formation of Earth about 4.6 billion years ago to the beginning of the Cambrian Period, about 541 million years ago, the Precambrian accounts for 89% of geologic time. Relatively little is known about the Precambrian, despite it making up roughly seven-eighths of the Earths history, the Precambrian fossil record is poorer than that of the succeeding Phanerozoic, and fossils from that time are of limited biostratigraphic use. This is because many Precambrian rocks have been metamorphosed, obscuring their origins, while others have been destroyed by erosion. A stable crust was apparently in place by 4,412 Ma, the term Precambrian is recognized by the International Commission on Stratigraphy as a general term including the Archean and Proterozoic eons. It is still used by geologists and paleontologists for general discussions not requiring the more specific eon names and it was briefly called the Cryptozoic eon. A specific date for the origin of life has not been determined, carbon found in 3.8 billion year old rocks from islands off western Greenland may be of organic origin. Well-preserved microscopic fossils of bacteria older than 3.46 billion years have found in Western Australia. Probable fossils 100 million years older have been found in the same area, there is a fairly solid record of bacterial life throughout the remainder of the Precambrian. The oldest fossil evidence from that era of such complex life comes from the Lantian formation of the Ediacarian period, a very diverse collection of soft-bodied forms is found in a variety of locations worldwide and date to between 635 and 542 Ma. These are referred to as Ediacaran or Vendian biota, hard-shelled creatures appeared toward the end of that time span, marking the beginning of the Phanerozoic era. By the middle of the following Cambrian period, a diverse fauna is recorded in the Burgess Shale. The explosion in diversity of lifeforms during the early Cambrian is called the Cambrian explosion of life, while land seems to have been devoid of plants and animals, cyanobacteria and other microbes formed prokaryotic mats that covered terrestrial areas. Evidence of the details of plate motions and other activity in the Precambrian has been poorly preserved. It is generally believed that small proto-continents existed prior to 4280 Ma, the supercontinent, known as Rodinia, broke up around 750 Ma. A number of glacial periods have been identified going as far back as the Huronian epoch, one of the best studied is the Sturtian-Varangian glaciation, around 850–635 Ma, which may have brought glacial conditions all the way to the equator, resulting in a Snowball Earth. The atmosphere of the early Earth is not well understood, most geologists believe it was composed primarily of nitrogen, carbon dioxide, and other relatively inert gases, and was lacking in free oxygen
3.
Cambrian
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The Cambrian Period was the first geological period of the Paleozoic Era, of the Phanerozoic Eon. The Cambrian lasted 55.6 million years from the end of the preceding Ediacaran Period 541 million years ago to the beginning of the Ordovician Period 485.4 mya and its subdivisions, and its base, are somewhat in flux. The period was established by Adam Sedgwick, who named it after Cambria, the Latinised form of Cymru, the Welsh name for Wales, as a result, our understanding of the Cambrian biology surpasses that of some later periods. The rapid diversification of lifeforms in the Cambrian, known as the Cambrian explosion, most of the continents were probably dry and rocky due to a lack of vegetation. Shallow seas flanked the margins of several continents created during the breakup of the supercontinent Pannotia, the seas were relatively warm, and polar ice was absent for much of the period. The United States Federal Geographic Data Committee uses a barred capital C ⟨Є⟩ character similar to the capital letter Ukrainian Ye ⟨Є⟩ to represent the Cambrian Period, the proper Unicode character is U+A792 Ꞓ LATIN CAPITAL LETTER C WITH BAR. Despite the long recognition of its distinction from younger Ordovician Period rocks and older Supereon Precambrian rocks, the base of the Cambrian lies atop a complex assemblage of trace fossils known as the Treptichnus pedum assemblage. Pedum in Namibia, Spain and Newfoundland, and possibly, in the western USA, the stratigraphic range of T. pedum overlaps the range of the Ediacaran fossils in Namibia, and probably in Spain. The Cambrian Period followed the Ediacaran Period and was followed by the Ordovician Period, the Cambrian is divided into four epochs and ten ages. Currently only two series and five stages are named and have a GSSP, because the international stratigraphic subdivision is not yet complete, many local subdivisions are still widely used. In some of these subdivisions the Cambrian is divided into three epochs with locally differing names – the Early Cambrian, Middle Cambrian and Furongian, rocks of these epochs are referred to as belonging to the Lower, Middle, or Upper Cambrian. Trilobite zones allow biostratigraphic correlation in the Cambrian, each of the local epochs is divided into several stages. The International Commission on Stratigraphy list the Cambrian period as beginning at 541 million years ago, the lower boundary of the Cambrian was originally held to represent the first appearance of complex life, represented by trilobites. The recognition of small shelly fossils before the first trilobites, and Ediacara biota substantially earlier and this formal designation allowed radiometric dates to be obtained from samples across the globe that corresponded to the base of the Cambrian. Early dates of 570 million years ago quickly gained favour, though the used to obtain this number are now considered to be unsuitable. A more precise date using modern radiometric dating yield a date of 541 ±0.3 million years ago, most continental land was clustered in the Southern Hemisphere at this time, but was drifting north. Large, high-velocity rotational movement of Gondwana appears to have occurred in the Early Cambrian, the sea levels fluctuated somewhat, suggesting there were ice ages, associated with pulses of expansion and contraction of a south polar ice cap. In Baltoscandia a Lower Cambrian transgression transformed large swathes of the Sub-Cambrian peneplain into a epicontinental sea, the Earth was generally cold during the early Cambrian, probably due to the ancient continent of Gondwana covering the South Pole and cutting off polar ocean currents
4.
Ordovician
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The Ordovician is a geologic period and system, the second of six periods of the Paleozoic Era. The Ordovician spans 41.2 million years from the end of the Cambrian Period 485.4 million years ago to the start of the Silurian Period 443.8 Mya. Lapworth recognized that the fauna in the disputed strata were different from those of either the Cambrian or the Silurian periods. It received international sanction in 1960, when it was adopted as a period of the Paleozoic Era by the International Geological Congress. Life continued to flourish during the Ordovician as it did in the earlier Cambrian period, invertebrates, namely molluscs and arthropods, dominated the oceans. The Great Ordovician Biodiversification Event considerably increased the diversity of life, fish, the worlds first true vertebrates, continued to evolve, and those with jaws may have first appeared late in the period. Life had yet to diversify on land, about 100 times as many meteorites struck the Earth during the Ordovician compared with today. The Ordovician Period began with a major extinction called the Cambrian–Ordovician extinction event and it lasted for about 42 million years and ended with the Ordovician–Silurian extinction event, about 443.8 Mya which wiped out 60% of marine genera. The dates given are recent radiometric dates and vary slightly from those found in other sources and this second period of the Paleozoic era created abundant fossils that became major petroleum and gas reservoirs. The boundary chosen for the beginning of both the Ordovician Period and the Tremadocian stage is highly significant and it correlates well with the occurrence of widespread graptolite, conodont, and trilobite species. The base of the Tremadocian allows scientists to relate these species not only to each other and this makes it easier to place many more species in time relative to the beginning of the Ordovician Period. A number of terms have been used to subdivide the Ordovician Period. In 2008, the ICS erected an international system of subdivisions. There exist Baltoscandic, British, Siberian, North American, Australian, the Ordovician Period in Britain was traditionally broken into Early, Middle and Late epochs. The corresponding rocks of the Ordovician System are referred to as coming from the Lower, Middle, the Floian corresponds to the lower Arenig, the Arenig continues until the early Darriwilian, subsuming the Dapingian. The Llanvirn occupies the rest of the Darriwilian, and terminates with it at the base of the Late Ordovician. The Sandbian represents the first half of the Caradoc, the Caradoc ends in the mid-Katian, during the Ordovician, the southern continents were collected into Gondwana. Gondwana started the period in equatorial latitudes and, as the period progressed, drifted toward the South Pole, the small continent Avalonia separated from Gondwana and began to move north towards Baltica and Laurentia, opening the Rheic Ocean between Gondwana and Avalonia
5.
Silurian
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The Silurian is a geologic period and system spanning 24.6 million years from the end of the Ordovician Period, at 443.8 million years ago, to the beginning of the Devonian Period,419.2 Mya. As with other periods, the rock beds that define the periods start and end are well identified. The base of the Silurian is set at a major Ordovician-Silurian extinction event when 60% of marine species were wiped out, a significant evolutionary milestone during the Silurian was the diversification of jawed and bony fish. However, terrestrial life would not greatly diversify and affect the landscape until the Devonian, the Silurian system was first identified by British geologist Sir Roderick Impey Murchison, who was examining fossil-bearing sedimentary rock strata in south Wales in the early 1830s. He named the sequences for a Celtic tribe of Wales, the Silures, inspired by his friend Adam Sedgwick and this naming does not indicate any correlation between the occurrence of the Silurian rocks and the land inhabited by the Silures. As it was first identified, the Silurian series when traced farther afield quickly came to overlap Sedgwicks Cambrian sequence, however, charles Lapworth resolved the conflict by defining a new Ordovician system including the contested beds. An early alternative name for the Silurian was Gotlandian after the strata of the Baltic island of Gotland, the French geologist Joachim Barrande, building on Murchisons work, used the term Silurian in a more comprehensive sense than was justified by subsequent knowledge. He divided the Silurian rocks of Bohemia into eight stages and his interpretation was questioned in 1854 by Edward Forbes, and the later stages of Barrande, F, G and H, have since been shown to be Devonian. Despite these modifications in the groupings of the strata, it is recognized that Barrande established Bohemia as a classic ground for the study of the earliest fossils. The epoch is named for the town of Llandovery in Carmarthenshire, the Wenlock, which lasted from 433.4 ±1.5 to 427.4 ±2.8 mya, is subdivided into the Sheinwoodian and Homerian ages. It is named after Wenlock Edge in Shropshire, England, during the Wenlock, the oldest known tracheophytes of the genus Cooksonia, appear. The first terrestrial animals also appear in the Wenlock, represented by air-breathing millipedes from Scotland. The Ludlow, lasting from 427.4 ±1.5 to 423 ±2.8 mya, comprises the Gorstian stage, lasting until 425.6 million years ago, and it is named for the town of Ludlow in Shropshire, England. The Pridoli, lasting from 423 ±1.5 to 419.2 ±2.8 mya, is the final and it is named after one locality at the Homolka a Přídolí nature reserve near the Prague suburb Slivenec in the Czech Republic. Přídolí is the old name of a field area. The high sea levels of the Silurian and the flat land resulted in a number of island chains. The southern continents remained united during this period, the melting of icecaps and glaciers contributed to a rise in sea level, recognizable from the fact that Silurian sediments overlie eroded Ordovician sediments, forming an unconformity. The continents of Avalonia, Baltica, and Laurentia drifted together near the equator and this event is the Caledonian orogeny, a spate of mountain building that stretched from New York State through conjoined Europe and Greenland to Norway
6.
Devonian
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The Devonian is a geologic period and system of the Paleozoic, spanning 60 million years from the end of the Silurian,419.2 million years ago, to the beginning of the Carboniferous,358.9 Mya. It is named after Devon, England, where rocks from this period were first studied, the first significant adaptive radiation of life on dry land occurred during the Devonian. Free-sporing vascular plants began to spread across dry land, forming extensive forests which covered the continents, by the middle of the Devonian, several groups of plants had evolved leaves and true roots, and by the end of the period the first seed-bearing plants appeared. Various terrestrial arthropods also became well-established, Fish reached substantial diversity during this time, leading the Devonian to often be dubbed the Age of Fish. The first ray-finned and lobe-finned bony fish appeared, while the placodermi began dominating almost every aquatic environment. The ancestors of all four-limbed vertebrates began adapting to walking on land, as their strong pectoral, in the oceans, primitive sharks became more numerous than in the Silurian and Late Ordovician. The first ammonites, species of molluscs, appeared, trilobites, the mollusk-like brachiopods and the great coral reefs, were still common. The Late Devonian extinction which started about 375 million years ago severely affected marine life, killing off all placodermi, and all trilobites, save for a few species of the order Proetida. The palaeogeography was dominated by the supercontinent of Gondwana to the south, the continent of Siberia to the north, while the rock beds that define the start and end of the Devonian period are well identified, the exact dates are uncertain. According to the International Commission on Stratigraphy, the Devonian extends from the end of the Silurian 419.2 Mya, another common term is Age of the Fishes, referring to the evolution of several major groups of fish that took place during the period. Older literature on the Anglo-Welsh basin divides it into the Downtonian, Dittonian, Breconian and Farlovian stages, in the Late Devonian, by contrast, arid conditions were less prevalent across the world and temperate climates were more common. The Devonian Period is formally broken into Early, Middle and Late subdivisions, the rocks corresponding to those epochs are referred to as belonging to the Lower, Middle and Upper parts of the Devonian System. Early Devonian The Early Devonian lasted from 419.2 ±2.8 to 393.3 ±2.5 and began with the Lochkovian stage, which lasted until the Pragian. It spanned from 410.8 ±2.8 to 407.6 ±2.5, and was followed by the Emsian, which lasted until the Middle Devonian began,393. 3±2.7 million years ago. Middle Devonian The Middle Devonian comprised two subdivisions, first the Eifelian, which gave way to the Givetian 387. 7±2.7 million years ago. Late Devonian Finally, the Late Devonian started with the Frasnian,382.7 ±2.8 to 372.2 ±2.5, during which the first forests took shape on land. The first tetrapods appeared in the record in the ensuing Famennian subdivision. This lasted until the end of the Devonian,358. 9±2.5 million years ago, the Devonian was a relatively warm period, and probably lacked any glaciers
7.
Carboniferous
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The Carboniferous is a geologic period and system that spans 60 million years from the end of the Devonian Period 358.9 million years ago, to the beginning of the Permian Period,298.9 Mya. The name Carboniferous means coal-bearing and derives from the Latin words carbō and ferō, and was coined by geologists William Conybeare and William Phillips in 1822. Based on a study of the British rock succession, it was the first of the system names to be employed. The Carboniferous is often treated in North America as two periods, the earlier Mississippian and the later Pennsylvanian. Terrestrial life was established by the Carboniferous period. Amphibians were the dominant land vertebrates, of one branch would eventually evolve into amniotes. Arthropods were also common, and many were much larger than those of today. Vast swaths of forest covered the land, which would eventually be laid down, the atmospheric content of oxygen also reached their highest levels in geological history during the period, 35% compared with 21% today, allowing terrestrial invertebrates to evolve to great size. A major marine and terrestrial extinction event, the Carboniferous rainforest collapse, occurred in the middle of the period, the later half of the period experienced glaciations, low sea level, and mountain building as the continents collided to form Pangaea. In the United States the Carboniferous is usually broken into Mississippian and Pennsylvanian subperiods, the Silesian is roughly contemporaneous with the late Mississippian Serpukhovian plus the Pennsylvanian. In Britain the Dinantian is traditionally known as the Carboniferous Limestone, the Namurian as the Millstone Grit, and the Westphalian as the Coal Measures and Pennant Sandstone. There was also a drop in south polar temperatures, southern Gondwanaland was glaciated throughout the period and these conditions apparently had little effect in the deep tropics, where lush swamps, later to become coal, flourished to within 30 degrees of the northernmost glaciers. Mid-Carboniferous, a drop in sea level precipitated a major extinction, one that hit crinoids. This sea level drop and the unconformity in North America separate the Mississippian subperiod from the Pennsylvanian subperiod. This happened about 323 million years ago, at the onset of the Permo-Carboniferous Glaciation, the Carboniferous was a time of active mountain-building, as the supercontinent Pangaea came together. The southern continents remained tied together in the supercontinent Gondwana, which collided with North America–Europe along the present line of eastern North America, in the same time frame, much of present eastern Eurasian plate welded itself to Europe along the line of the Ural mountains. Most of the Mesozoic supercontinent of Pangea was now assembled, although North China, the Late Carboniferous Pangaea was shaped like an O. There were two major oceans in the Carboniferous—Panthalassa and Paleo-Tethys, which was inside the O in the Carboniferous Pangaea, other minor oceans were shrinking and eventually closed - Rheic Ocean, the small, shallow Ural Ocean and Proto-Tethys Ocean
8.
Permian
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The Permian is a geologic period and system which spans 46.7 million years from the end of the Carboniferous Period 298.9 million years ago, to the beginning of the Triassic Period 252.2 Mya. It is the last period of the Paleozoic Era, the following Triassic Period belongs to the Mesozoic Era, the concept of the Permian was introduced in 1841 by geologist Sir Roderick Murchison, who named it after the city of Perm. The Permian witnessed the diversification of the early amniotes into the groups of the mammals, turtles, lepidosaurs. The world at the time was dominated by two known as Pangaea and Siberia, surrounded by a global ocean called Panthalassa. The Carboniferous rainforest collapse left behind vast regions of desert within the continental interior, amniotes, who could better cope with these drier conditions, rose to dominance in place of their amphibian ancestors. The Permian ended with the Permian–Triassic extinction event, the largest mass extinction in Earths history, in which nearly 90% of marine species and it would take well into the Triassic for life to recover from this catastrophe. Recovery from the Permian-Triassic extinction event was protracted, on land, the term Permian was introduced into geology in 1841 by Sir R. I. Murchison, president of the Geological Society of London, who identified typical strata in extensive Russian explorations undertaken with Edouard de Verneuil, the region now lies in the Perm Krai of Russia. This could have in part caused the extinctions of marine species at the end of the period by severely reducing shallow coastal areas preferred by many marine organisms. During the Permian, all the Earths major landmasses were collected into a supercontinent known as Pangaea. The Cimmeria continent rifted away from Gondwana and drifted north to Laurasia, a new ocean was growing on its southern end, the Tethys Ocean, an ocean that would dominate much of the Mesozoic Era. Large continental landmass interiors experience climates with extreme variations of heat and cold, deserts seem to have been widespread on Pangaea. Such dry conditions favored gymnosperms, plants with seeds enclosed in a cover, over plants such as ferns that disperse spores in a wetter environment. The first modern trees appeared in the Permian, the climate in the Permian was quite varied. At the start of the Permian, the Earth was still in an Ice Age, glaciers receded around the mid-Permian period as the climate gradually warmed, drying the continents interiors. In the late Permian period, the drying continued although the temperature cycled between warm and cool cycles, Permian marine deposits are rich in fossil mollusks, echinoderms, and brachiopods. By the close of the Permian, trilobites and a host of other groups became extinct. Terrestrial life in the Permian included diverse plants, fungi, arthropods, the period saw a massive desert covering the interior of Pangaea
9.
Triassic
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The Triassic is a geologic period and system which spans 50.9 million years from the end of the Permian Period 252.17 million years ago, to the beginning of the Jurassic Period 201.3 Mya. The Triassic is the first period of the Mesozoic Era, both the start and end of the period are marked by major extinction events. Therapsids and archosaurs were the terrestrial vertebrates during this time. A specialized subgroup of archosaurs, called dinosaurs, first appeared in the Late Triassic, the vast supercontinent of Pangaea existed until the mid-Triassic, after which it began to gradually rift into two separate landmasses, Laurasia to the north and Gondwana to the south. The global climate during the Triassic was mostly hot and dry, however, the climate shifted and became more humid as Pangaea began to drift apart. The end of the period was marked by yet another mass extinction, the Triassic-Jurassic extinction event. The Triassic is usually separated into Early, Middle, and Late Triassic Epochs, from the east, along the equator, the Tethys sea penetrated Pangaea, causing the Paleo-Tethys Ocean to be closed. Later in the mid-Triassic a similar sea penetrated along the equator from the west, the remaining shores were surrounded by the world-ocean known as Panthalassa. All the deep-ocean sediments laid down during the Triassic have disappeared through subduction of oceanic plates, thus, the supercontinent Pangaea was rifting during the Triassic—especially late in that period—but had not yet separated. In North America, for example, marine deposits are limited to a few exposures in the west, thus Triassic stratigraphy is mostly based on organisms that lived in lagoons and hypersaline environments, such as Estheria crustaceans. At the beginning of the Mesozoic Era, Africa was joined with Earths other continents in Pangaea, Africa shared the supercontinents relatively uniform fauna which was dominated by theropods, prosauropods and primitive ornithischians by the close of the Triassic period. Late Triassic fossils are found throughout Africa, but are common in the south than north. The time boundary separating the Permian and Triassic marks the advent of an event with global impact. At Paleorrota geopark, located in Rio Grande do Sul, Brazil, in these formations, one of the earliest dinosaurs, Staurikosaurus, as well as the mammal ancestors Brasilitherium and Brasilodon have been discovered. The Triassic continental interior climate was hot and dry, so that typical deposits are red bed sandstones and evaporites. Pangaeas large size limited the effect of the global ocean, its continental climate was highly seasonal, with very hot summers. The strong contrast between the Pangea supercontinent and the global ocean triggered intense cross-equatorial monsoons, the best studied of such episodes of humid climate, and probably the most intense and widespread, was the Carnian Pluvial Event. On land, the vascular plants included the lycophytes, the dominant cycadophytes, ginkgophyta, ferns, horsetails
10.
Jurassic
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The Jurassic is a geologic period and system that spans 56.3 million years from the end of the Triassic Period 201.3 million years ago to the beginning of the Cretaceous Period 145 Mya. The Jurassic constitutes the middle period of the Mesozoic Era, also known as the Age of Reptiles, the start of the period is marked by the major Triassic–Jurassic extinction event. The Jurassic is named after the Jura Mountains within the European Alps, by the beginning of the Jurassic, the supercontinent Pangaea had begun rifting into two landmasses, Laurasia to the north and Gondwana to the south. This created more coastlines and shifted the continental climate from dry to humid, on land, the fauna transitioned from the Triassic fauna, dominated by both dinosauromorph and crocodylomorph archosaurs, to one dominated by dinosaurs alone. The first birds also appeared during the Jurassic, having evolved from a branch of theropod dinosaurs, other major events include the appearance of the earliest lizards, and the evolution of therian mammals, including primitive placentals. Crocodilians made the transition from a terrestrial to a mode of life. The oceans were inhabited by marine reptiles such as ichthyosaurs and plesiosaurs, the chronostratigraphic term Jurassic is directly linked to the Jura Mountains. The name Jura is derived from the Celtic root jor, which was Latinised into juria, the Jurassic period is divided into the Early Jurassic, Middle, and Late Jurassic epochs. The Jurassic System, in stratigraphy, is divided into the Lower Jurassic, Middle, the separation of the term Jurassic into three sections goes back to Leopold von Buch. The Jurassic North Atlantic Ocean was relatively narrow, while the South Atlantic did not open until the following Cretaceous period, the Tethys Sea closed, and the Neotethys basin appeared. Climates were warm, with no evidence of glaciation, as in the Triassic, there was apparently no land over either pole, and no extensive ice caps existed. In contrast, the North American Jurassic record is the poorest of the Mesozoic, the Jurassic was a time of calcite sea geochemistry in which low-magnesium calcite was the primary inorganic marine precipitate of calcium carbonate. Carbonate hardgrounds were thus very common, along with calcitic ooids, calcitic cements, the first of several massive batholiths were emplaced in the northern American cordillera beginning in the mid-Jurassic, marking the Nevadan orogeny. Important Jurassic exposures are found in Russia, India, South America, Japan, Australasia. As the Jurassic proceeded, larger and more groups of dinosaurs like sauropods and ornithopods proliferated in Africa. Middle Jurassic strata are well represented nor well studied in Africa. Late Jurassic strata are also poorly represented apart from the spectacular Tendaguru fauna in Tanzania, the Late Jurassic life of Tendaguru is very similar to that found in western North Americas Morrison Formation. During the Jurassic period, the primary living in the sea were fish
11.
Cretaceous
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The Cretaceous is a geologic period and system that spans 79 million years from the end of the Jurassic Period 145 million years ago to the beginning of the Paleogene Period 66 Mya. It is the last period of the Mesozoic Era, the Cretaceous Period is usually abbreviated K, for its German translation Kreide. The Cretaceous was a period with a warm climate, resulting in high eustatic sea levels that created numerous shallow inland seas. These oceans and seas were populated with now-extinct marine reptiles, ammonites and rudists, during this time, new groups of mammals and birds, as well as flowering plants, appeared. The Cretaceous ended with a mass extinction, the Cretaceous–Paleogene extinction event, in which many groups, including non-avian dinosaurs, pterosaurs. The end of the Cretaceous is defined by the abrupt Cretaceous–Paleogene boundary, the name Cretaceous was derived from Latin creta, meaning chalk. The Cretaceous is divided into Early and Late Cretaceous epochs, or Lower and Upper Cretaceous series, in older literature the Cretaceous is sometimes divided into three series, Neocomian, Gallic and Senonian. A subdivision in eleven stages, all originating from European stratigraphy, is now used worldwide, in many parts of the world, alternative local subdivisions are still in use. As with other geologic periods, the rock beds of the Cretaceous are well identified. No great extinction or burst of diversity separates the Cretaceous from the Jurassic and this layer has been dated at 66.043 Ma. A140 Ma age for the Jurassic-Cretaceous boundary instead of the usually accepted 145 Ma was proposed in 2014 based on a study of Vaca Muerta Formation in Neuquén Basin. Víctor Ramos, one of the authors of the study proposing the 140 Ma boundary age sees the study as a first step toward formally changing the age in the International Union of Geological Sciences, due to the high sea level there was extensive space for such sedimentation. Because of the young age and great thickness of the system. Chalk is a type characteristic for the Cretaceous. It consists of coccoliths, microscopically small calcite skeletons of coccolithophores, the group is found in England, northern France, the low countries, northern Germany, Denmark and in the subsurface of the southern part of the North Sea. Chalk is not easily consolidated and the Chalk Group still consists of sediments in many places. The group also has other limestones and arenites, among the fossils it contains are sea urchins, belemnites, ammonites and sea reptiles such as Mosasaurus. In southern Europe, the Cretaceous is usually a marine system consisting of competent limestone beds or incompetent marls
12.
Paleogene
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The Paleogene is a geologic period and system that spans 43 million years from the end of the Cretaceous Period 66 million years ago to the beginning of the Neogene Period 23.03 Mya. It is the beginning of the Cenozoic Era of the present Phanerozoic Eon and this period consists of the Paleocene, Eocene and Oligocene epochs. The terms Paleogene System and lower Tertiary System are applied to the rocks deposited during the Paleogene Period. By dividing the Tertiary Period into two periods instead of directly into five epochs, the periods are more comparable to the duration of periods of the preceding Mesozoic and Paleozoic Eras. The trend was caused by the formation of the Antarctic Circumpolar Current. During the Paleogene, the continued to drift closer to their current positions. India was in the process of colliding with Asia, subsequently forming the Himalayas, the Atlantic Ocean continued to widen by a few centimeters each year. Africa was moving north to meet with Europe and form the Mediterranean, inland seas retreated from North America early in the period. Australia had also separated from Antarctica and was drifting towards Southeast Asia, mammals began a rapid diversification during this period. Some of these mammals would evolve into forms that would dominate the land, while others would become capable of living in marine, specialized terrestrial. Those that took to the oceans became modern cetaceans, while those that took to the trees became primates, the group to which humans belong. Birds, which were well established by the end of the Cretaceous. In comparison to birds and mammals, most other branches of life remained unchanged during this period. As the Earth cooled, tropical plants became less numerous and were now restricted to equatorial regions, deciduous plants, which could survive through the seasonal climates the world was now experiencing, became more common. The Paleogene is notable in the context of offshore oil drilling, and especially in Gulf of Mexico oil exploration and these rock formations represent the current cutting edge of deep-water oil discovery. Lower Tertiary explorations to date include, Kaskida Oil Field Tiber Oil Field Jack 2 Paleogene Microfossils, 180+ images of Foraminifera
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Neogene
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The Neogene is a geologic period and system that spans 20.45 million years from the end of the Paleogene Period 23.03 million years ago to the beginning of the present Quaternary Period 2.58 Mya. The Neogene is sub-divided into two epochs, the earlier Miocene and the later Pliocene, some geologists assert that the Neogene cannot be clearly delineated from the modern geological period, the Quaternary. During this period, mammals and birds continued to evolve into modern forms. Early hominids, the ancestors of humans, appeared in Africa near the end of the period, some continental movement took place, the most significant event being the connection of North and South America at the Isthmus of Panama, late in the Pliocene. This cut off the ocean currents from the Pacific to the Atlantic ocean. The global climate cooled considerably over the course of the Neogene, the terms Neogene System and upper Tertiary System describe the rocks deposited during the Neogene Period. The continents in the Neogene were very close to their current positions, the Isthmus of Panama formed, connecting North and South America. The Indian subcontinent continued to collide with Asia, forming the Himalayas, sea levels fell, creating land bridges between Africa and Eurasia and between Eurasia and North America. The global climate became seasonal and continued an overall drying and cooling trend which began at the start of the Paleogene. The ice caps on both poles began to grow and thicken, and by the end of the period the first of a series of glaciations of the current Ice Age began, marine and continental flora and fauna have a modern appearance. The reptile group Choristodera became extinct in the part of the period. Mammals and birds continued to be the dominant terrestrial vertebrates, the first hominids, the ancestors of humans, appeared in Africa and spread into Eurasia. In response to the cooler, seasonal climate, tropical plant species gave way to deciduous ones, grasses therefore greatly diversified, and herbivorous mammals evolved alongside it, creating the many grazing animals of today such as horses, antelope, and bison. The Neogene traditionally ended at the end of the Pliocene Epoch, just before the definition of the beginning of the Quaternary Period. However, there was a movement amongst geologists to also include ongoing geological time in the Neogene, by dividing the Cenozoic Era into three periods instead of seven epochs, the periods are more closely comparable to the duration of periods in the Mesozoic and Paleozoic eras. The International Commission on Stratigraphy once proposed that the Quaternary be considered a sub-era of the Neogene, with a date of 2.58 Ma. In the 2004 proposal of the ICS, the Neogene would have consisted of the Miocene and Pliocene epochs, thus the Neogene Period ends bounding the succeeding Quaternary Period at 2.58 Mya. Digital Atlas of Neogene Life for the Southeastern United States — by San Jose State University via Web Archive
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Taxonomy (biology)
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Taxonomy is the science of defining groups of biological organisms on the basis of shared characteristics and giving names to those groups. The exact definition of taxonomy varies from source to source, but the core of the remains, the conception, naming. There is some disagreement as to whether biological nomenclature is considered a part of taxonomy, the broadest meaning of taxonomy is used here. The word taxonomy was introduced in 1813 by Candolle, in his Théorie élémentaire de la botanique, the term alpha taxonomy is primarily used today to refer to the discipline of finding, describing, and naming taxa, particularly species. In earlier literature, the term had a different meaning, referring to morphological taxonomy, ideals can, it may be said, never be completely realized. They have, however, a value of acting as permanent stimulants. Some of us please ourselves by thinking we are now groping in a beta taxonomy, turrill thus explicitly excludes from alpha taxonomy various areas of study that he includes within taxonomy as a whole, such as ecology, physiology, genetics, and cytology. He further excludes phylogenetic reconstruction from alpha taxonomy, thus, Ernst Mayr in 1968 defined beta taxonomy as the classification of ranks higher than species. This activity is what the term denotes, it is also referred to as beta taxonomy. How species should be defined in a group of organisms gives rise to practical and theoretical problems that are referred to as the species problem. The scientific work of deciding how to define species has been called microtaxonomy, by extension, macrotaxonomy is the study of groups at higher taxonomic ranks, from subgenus and above only, than species. While some descriptions of taxonomic history attempt to date taxonomy to ancient civilizations, earlier works were primarily descriptive, and focused on plants that were useful in agriculture or medicine. There are a number of stages in scientific thinking. Early taxonomy was based on criteria, the so-called artificial systems. Later came systems based on a complete consideration of the characteristics of taxa, referred to as natural systems, such as those of de Jussieu, de Candolle and Bentham. The publication of Charles Darwins Origin of Species led to new ways of thinking about classification based on evolutionary relationships and this was the concept of phyletic systems, from 1883 onwards. This approach was typified by those of Eichler and Engler, the advent of molecular genetics and statistical methodology allowed the creation of the modern era of phylogenetic systems based on cladistics, rather than morphology alone. Taxonomy has been called the worlds oldest profession, and naming and classifying our surroundings has likely been taking place as long as mankind has been able to communicate
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Animal
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Animals are multicellular, eukaryotic organisms of the kingdom Animalia. The animal kingdom emerged as a clade within Apoikozoa as the group to the choanoflagellates. Animals are motile, meaning they can move spontaneously and independently at some point in their lives and their body plan eventually becomes fixed as they develop, although some undergo a process of metamorphosis later in their lives. All animals are heterotrophs, they must ingest other organisms or their products for sustenance, most known animal phyla appeared in the fossil record as marine species during the Cambrian explosion, about 542 million years ago. Animals can be divided broadly into vertebrates and invertebrates, vertebrates have a backbone or spine, and amount to less than five percent of all described animal species. They include fish, amphibians, reptiles, birds and mammals, the remaining animals are the invertebrates, which lack a backbone. These include molluscs, arthropods, annelids, nematodes, flatworms, cnidarians, ctenophores, the study of animals is called zoology. The word animal comes from the Latin animalis, meaning having breath, the biological definition of the word refers to all members of the kingdom Animalia, encompassing creatures as diverse as sponges, jellyfish, insects, and humans. Aristotle divided the world between animals and plants, and this was followed by Carl Linnaeus, in the first hierarchical classification. In Linnaeuss original scheme, the animals were one of three kingdoms, divided into the classes of Vermes, Insecta, Pisces, Amphibia, Aves, and Mammalia. Since then the last four have all been subsumed into a single phylum, in 1874, Ernst Haeckel divided the animal kingdom into two subkingdoms, Metazoa and Protozoa. The protozoa were later moved to the kingdom Protista, leaving only the metazoa, thus Metazoa is now considered a synonym of Animalia. Animals have several characteristics that set apart from other living things. Animals are eukaryotic and multicellular, which separates them from bacteria and they are heterotrophic, generally digesting food in an internal chamber, which separates them from plants and algae. They are also distinguished from plants, algae, and fungi by lacking cell walls. All animals are motile, if only at life stages. In most animals, embryos pass through a stage, which is a characteristic exclusive to animals. With a few exceptions, most notably the sponges and Placozoa and these include muscles, which are able to contract and control locomotion, and nerve tissues, which send and process signals
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Chordate
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Chordates are deuterostomes, as during the embryo development stage the anus forms before the mouth. They are also bilaterally symmetric coelomates, in the case of vertebrate chordates, the notochord is usually replaced by a vertebral column during development, and they may have body plans organized via segmentation. There are also additional extinct taxa, the Vertebrata are sometimes considered as a subgroup of the clade Craniata, consisting of chordates with a skull, the Craniata and Tunicata compose the clade Olfactores. Of the more than 65,000 living species of chordates, the worlds largest and fastest animals, the blue whale and peregrine falcon respectively, are chordates, as are humans. Fossil chordates are known from at least as early as the Cambrian explosion, Hemichordata, which includes the acorn worms, has been presented as a fourth chordate subphylum, but it now is usually treated as a separate phylum. The Hemichordata, along with the Echinodermata, form the Ambulacraria, the Chordata and Ambulacraria form the superphylum Deuterostomia, composed of the deuterostomes. Attempts to work out the relationships of the chordates have produced several hypotheses. All of the earliest chordate fossils have found in the Early Cambrian Chengjiang fauna. Because the fossil record of early chordates is poor, only molecular phylogenetics offers a prospect of dating their emergence. However, the use of molecular phylogenetics for dating evolutionary transitions is controversial and it has also proved difficult to produce a detailed classification within the living chordates. Attempts to produce family trees shows that many of the traditional classes are paraphyletic. While this has been known since the 19th century, an insistence on only monophyletic taxa has resulted in vertebrate classification being in a state of flux. Although the name Chordata is attributed to William Bateson, it was already in prevalent use by 1880, ernst Haeckel described a taxon comprising tunicates, cephalochordates, and vertebrates in 1866. Though he used the German vernacular form, it is allowed under the ICZN code because of its subsequent latinization, among the vertebrate sub-group of chordates the notochord develops into the spine, and in wholly aquatic species this helps the animal to swim by flexing its tail. In fish and other vertebrates, this develops into the spinal cord, the pharynx is the part of the throat immediately behind the mouth. In fish, the slits are modified to form gills, a muscular tail that extends backwards behind the anus. This is a groove in the wall of the pharynx. In filter-feeding species it produces mucus to gather food particles, which helps in transporting food to the esophagus and it also stores iodine, and may be a precursor of the vertebrate thyroid gland
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Mammal
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Mammals are any vertebrates within the class Mammalia, a clade of endothermic amniotes distinguished from reptiles by the possession of a neocortex, hair, three middle ear bones and mammary glands. All female mammals nurse their young with milk, secreted from the mammary glands, Mammals include the largest animals on the planet, the great whales. The basic body type is a quadruped, but some mammals are adapted for life at sea, in the air, in trees. The largest group of mammals, the placentals, have a placenta, Mammals range in size from the 30–40 mm bumblebee bat to the 30-meter blue whale. With the exception of the five species of monotreme, all modern mammals give birth to live young, most mammals, including the six most species-rich orders, belong to the placental group. The largest orders are the rodents, bats and Soricomorpha, the next three biggest orders, depending on the biological classification scheme used, are the Primates, the Cetartiodactyla, and the Carnivora. Living mammals are divided into the Yinotheria and Theriiformes There are around 5450 species of mammal, in some classifications, extant mammals are divided into two subclasses, the Prototheria, that is, the order Monotremata, and the Theria, or the infraclasses Metatheria and Eutheria. The marsupials constitute the group of the Metatheria, and include all living metatherians as well as many extinct ones. Much of the changes reflect the advances of cladistic analysis and molecular genetics, findings from molecular genetics, for example, have prompted adopting new groups, such as the Afrotheria, and abandoning traditional groups, such as the Insectivora. The mammals represent the only living Synapsida, which together with the Sauropsida form the Amniota clade, the early synapsid mammalian ancestors were sphenacodont pelycosaurs, a group that produced the non-mammalian Dimetrodon. At the end of the Carboniferous period, this group diverged from the line that led to todays reptiles. Some mammals are intelligent, with some possessing large brains, self-awareness, Mammals can communicate and vocalize in several different ways, including the production of ultrasound, scent-marking, alarm signals, singing, and echolocation. Mammals can organize themselves into fission-fusion societies, harems, and hierarchies, most mammals are polygynous, but some can be monogamous or polyandrous. They provided, and continue to provide, power for transport and agriculture, as well as commodities such as meat, dairy products, wool. Mammals are hunted or raced for sport, and are used as model organisms in science, Mammals have been depicted in art since Palaeolithic times, and appear in literature, film, mythology, and religion. Defaunation of mammals is primarily driven by anthropogenic factors, such as poaching and habitat destruction, Mammal classification has been through several iterations since Carl Linnaeus initially defined the class. No classification system is accepted, McKenna & Bell and Wilson & Reader provide useful recent compendiums. Though field work gradually made Simpsons classification outdated, it remains the closest thing to a classification of mammals
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Primate
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A primate is a mammal of the order Primates. In taxonomy, primates include two distinct lineages, strepsirrhines and haplorhines, Primates arose from ancestors that lived in the trees of tropical forests, many primate characteristics represent adaptations to life in this challenging three-dimensional environment. Most primate species remain at least partly arboreal, with the exception of humans, who inhabit every continent except for Antarctica, most primates live in tropical or subtropical regions of the Americas, Africa and Asia. Based on fossil evidence, the earliest known true primates, represented by the genus Teilhardina, an early close primate relative known from abundant remains is the Late Paleocene Plesiadapis, c. Molecular clock studies suggest that the branch may be even older. The order Primates was traditionally divided into two groupings, prosimians and anthropoids. Prosimians have characteristics more like those of the earliest primates, and include the lemurs of Madagascar, lorisoids, simians include monkeys, apes and hominins. Simians are divided into two groups, catarrhine monkeys and apes of Africa and Southeast Asia and platyrrhine or New World monkeys of South, catarrhines consist of Old World monkeys, gibbons and great apes, New World monkeys include the capuchin, howler and squirrel monkeys. Humans are the only extant catarrhines to have spread successfully outside of Africa, South Asia, New primate species are still being discovered. More than 25 species were described in the decade of the 2000s. Considered generalist mammals, primates exhibit a range of characteristics. Some primates are primarily terrestrial rather than arboreal, but all species possess adaptations for climbing trees, locomotion techniques used include leaping from tree to tree, walking on two or four limbs, knuckle-walking, and swinging between branches of trees. Primates are characterized by large brains relative to other mammals, as well as a reliance on stereoscopic vision at the expense of smell. These features are developed in monkeys and apes and noticeably less so in lorises. Three-color vision has developed in some primates, most also have opposable thumbs and some have prehensile tails. Many species are dimorphic, differences include body mass, canine tooth size. Primates have slower rates of development than other similarly sized mammals and reach maturity later, depending on the species, adults may live in solitude, in mated pairs, or in groups of up to hundreds of members. The relationships among the different groups of primates were not clearly understood until relatively recently, for example, ape has been used either as an alternative for monkey or for any tailless, relatively human-like primate
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Haplorhini
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Haplorrhini is a clade containing the tarsiers and the simians. The name is also spelt Haplorrhini, the simians include catarrhines, and the platyrrhines. The extinct omomyids, which are considered to be the most basal haplorhines, are believed to be closely related to the tarsiers than to other haplorhines. Haplorhines share a number of derived features that distinguish them from the strepsirrhine wet-nosed primates, the haplorhine upper lip, which has replaced the ancestral rhinarium found in strepsirrhines, is not directly connected to their nose or gum, allowing a large range of facial expressions. Their brain-to-body mass ratio is greater than the strepsirrhines. Haplorhines have a plate, unlike the postorbital bar found in strepsirrhines. All anthropoids have a uterus, tarsiers have a bicornate uterus like the strepsirrhines. Most species typically have single births, although twins and triplets are common for marmosets, despite similar gestation periods, haplorhine newborns are relatively much larger than strepsirrhine newborns, but have a longer dependence period on their mother. This difference in size and dependence is credited to the complexity of their behavior. The taxonomic name Haplorhini derives from the Ancient Greek haploûs and rhinos and it refers to the lack of a rhinarium or wet nose, which is found in many mammals, including strepsirrhine primates. Haplorhini and its sister clade, Strepsirrhini, diverged about 65 million years ago, the fossil Archicebus may be similar to the most recent common ancestor at this time. The other major clade within Haplorhini, the simians, is divided into two parvorders, Platyrrhini and Catarrhini, the New World monkeys split from catarrhines about 40 mya, while the apes diverged from Old World monkeys about 25 mya. The available fossil evidence indicates both the hominoid and cercopithecoid clades originated in Africa. The following is the listing of the living families, and their placement in the Order Primates, ORDER PRIMATES Suborder Strepsirrhini, lemurs, lorises
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Simian
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The simians are the monkeys, cladistically including the apes, the New World monkeys or platyrrhines, and the catarrhine clade consisting of the Old World monkeys and apes. The simian line and the line diverged about 60 million years ago. Forty million years ago, simians from Africa colonized South America, the remaining simians split 25 million years ago into apes and Old World monkeys. Under modern classification, the tarsiers and simians are grouped under the suborder Haplorhini while the strepsirrhines are placed in suborder Strepsirrhini, in anthropoidea, evidences indicate that the Old and the New World primates went through parallel evolution. Primatology, paleoanthropology, and other related fields are split on their usage of the synonymous infraorder names, Simiiformes and Anthropoidea. According to Robert Hoffstetter, the term Simiiformes has priority over Anthropoidea because of the taxonomic term Simii by van der Hoeven, from which it is constructed, dates to 1833. In contrast, Anthropoidea by Mivart dates to 1864, while Simiiformes by Haeckel dates to 1866, Hoffstetter also argued that Simiiformes is also constructed like a proper infraorder name, whereas Anthropoidea ends in -oidea, which is reserved for superfamilies. He also noted that Anthropoidea is too easily confused with anthropoïdes, the simians are split into three distinct groups. The New World monkeys in parvorder Platyrrhini split from the rest of the line about 40 mya. This group split about 25 mya between the Old World monkeys and the apes
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Hominidae
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Several revisions in classifying the great apes have caused the use of the term hominid to vary over time. Its original meaning referred only to humans and their closest non-extant relatives and that restrictive meaning has now been largely assumed by the term hominin, which comprises all members of the human clade after the split from the chimpanzees. The current, 21st-century meaning of hominid includes all the great apes including humans, the most recent common ancestor of all Hominidae lived roughly 14 million years ago, when the ancestors of the orangutans speciated from the ancestral line of the other three genera. Those ancestors of the family Hominidae had already speciated from the family Hylobatidae, in the early Miocene, about 22 million years ago, there were many species of arboreally adapted primitive catarrhines from East Africa, the variety suggests a long history of prior diversification. Fossils at 20 million years ago include fragments attributed to Victoriapithecus, the most recent of these far-flung Miocene apes is Oreopithecus, from the fossil-rich coal beds in northern Italy and dated to 9 million years ago. Species close to the last common ancestor of gorillas, chimpanzees and humans may be represented by Nakalipithecus fossils found in Kenya and Ouranopithecus found in Greece. Molecular evidence suggests that between 8 and 4 million years ago, first the gorillas, and then the split off from the line leading to the humans. Human DNA is approximately 98. 4% identical to that of chimpanzees when comparing single nucleotide polymorphisms, the earliest fossils argued by some to belong to the human lineage are Sahelanthropus tchadensis and Orrorin tugenensis, followed by Ardipithecus, with species Ar. kadabba and Ar. ramidus. The classification of the apes has been revised several times in the last few decades. The original meaning of the referred to only humans and their closest relatives—what is now the modern meaning of the term hominin. And the meaning of the taxon Hominidae changed gradually, leading to a different usage of hominid that today all the great apes including humans. A hominid is a member of the family Hominidae, the apes, orangutans, gorillas, chimpanzees. A hominine is a member of the subfamily Homininae, gorillas, chimpanzees, a hominin is a member of the tribe Hominini, Chimpanzees and humans. A homininan is a member of the subtribe Hominina of the tribe Hominini, a human is a member of the genus Homo, of which Homo sapiens is the only extant species, and within that Homo sapiens sapiens is the only surviving subspecies. For each clade it is indicated approximately when newer extant clades emerged, some texts will refer to Homonini as the Hominina branch. Many scientists, including paleoanthropologists, continue to use the term hominid to mean humans, as mentioned, Hominidae was originally the name given to the family of humans and their close relatives, with the other great apes all being placed in a separate family, the Pongidae. However, that eventually made Pongidae paraphyletic because at least one great ape species proved to be more closely related to humans than to other great apes. Most taxonomists today encourage monophyletic groups—this would require, in this case, thus, many biologists now assign Pongo to the family Hominidae
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Homininae
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Homininae is a subfamily of Hominidae. It includes two tribes, with their extant as well as species, the Hominini tribe. It comprises all hominids that arose after orangutans split from the line of great apes, the Homininae cladogram has three main branches, which lead to gorillas, and to humans and chimpanzees via the tribe Hominini and subtribes Hominina and Panina. There are two living species of Panina and two living species of gorillas, but only one extant human species, traces of hypothetical Homo species, including Homo floresiensis and Homo denisova, have been found with dates as recent as 12,000 years ago. Organisms in this class are described as hominine or hominines, until 1980, the family Hominidae meant humans only, the non-human great apes were assigned to the family Pongidae. The subfamily Homininae can be subdivided into three branches, the tribe Gorillini, and the tribe Hominini with subtribes Panina and Hominina. Their existence suggests that the Homininae tribes diverged not earlier than about 8 million years ago, today, chimpanzees and gorillas live in tropical forests with acid soils that rarely preserve fossils. This shows that some chimpanzees lived close to Homo at the time, australopithecus and early Paranthropus may have been bipedal. Very early hominins such as Ardipithecus ramidus may have possessed a type of bipedalism. It is proposed that they evolved from H. erectus as a case of insular dwarfism, in spite of their smaller brain, there is evidence that H. floresiensis used fire and made stone tools at least as sophisticated as those of their proposed ancestors H. erectus. In this case, it seems that for intelligence, the structure of the brain is more important than its size, sexuality is related to family structure and partly shapes it. The involvement of fathers in education is unique to humans. Concealed ovulation and menopause in women both also occur in a few other primates however, but are uncommon in other species, testis and penis size seems to be related to family structure, monogamy or promiscuity, or harem, in humans, chimpanzees or gorillas, respectively. The levels of sexual dimorphism are seen as a marker of sexual selection. Concealed ovulation means that the phase of fertility is not detectable in women, women can be partly aware of their ovulation along the menstrual phases, but men are essentially unable to detect ovulation in women. Menopause also occurs in rhesus monkeys, and possibly in chimpanzees, chimpanzee-human last common ancestor List of human evolution fossils Human Timeline – Smithsonian, National Museum of Natural History
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Hominini
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Members of the human clade, that is, the Hominina, including Homo and those species of the australopithecines that arose after the split from the chimpanzees, are called homininans. Not all homininans are directly related to the emergence of early Homo and this is a modern cladogram, For each clade, the cladogram above shows approximately when newer extant clades emerged. Some texts refer to Homonini as the Hominina branch, the subtribe Hominina is the human branch, that is, it contains only the genus Homo. Researchers proposed the taxon Hominini on the basis that the least similar species of a trichotomy should be separated from the other two. The common chimpanzee and the bonobo of the genus Pan are the closest living relatives to humans. All the extinct genera listed to the right are ancestral to, or offshoots of, however, both Orrorin and Sahelanthropus existed around the time of the split, and so may be ancestral to both Pan and Homo. In the proposal of Mann and Weiss, the tribe Hominini includes Pan as well as Homo, Homo and all bipedal apes are referred to the subtribe Hominina, while Pan is assigned to the subtribe Panina. Wood discusses the different views of this taxonomy, the assumption of late hybridization was in particular based on the similarity of the X chromosome in humans and chimpanzees, suggesting a divergence as late as some 4 million years ago. Sahelanthropus tchadensis is an extinct species that lived seven million years ago. Human Timeline – Smithsonian, National Museum of Natural History
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Homo
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The genus is between 2 and 3 million years old, taken to emerge with the appearance of Homo habilis. It is derived from the genus Australopithecus, which itself had previously split from the lineage of Pan, taxonomically, Homo is the only genus assigned to the subtribe Hominina which, with the subtribes Australopithecina and Panina, comprise the tribe Hominini. All species of the genus Homo plus those species of the australopithecines that arose after the split from Pan are called hominins, Homo erectus appeared about two million years ago in East Africa and, in several early migrations, it spread throughout Africa and Eurasia. It was likely the first hominin to live in a hunter-gatherer society, the subspecies Homo sapiens sapiens or anatomically modern humans emerged about 200,000 years ago in East Africa. DNA analysis provides evidence of interbreeding between archaic and modern humans. See Hominidae for an overview of taxonomy, the Latin noun homō means human being or man in the generic sense of human being, mankind. The binomial name Homo sapiens was coined by Carl Linnaeus, names for other species of the genus were introduced beginning in the second half of the 19th century. Even today, the genus Homo has not been properly defined, since the early human fossil record began to slowly emerge from the earth, the boundaries and definitions of the genus Homo have been poorly defined and constantly in flux. Because there was no reason to think it would ever have any additional members, the discovery of Neanderthal brought the first addition. The genus Homo was given its name to suggest that its member species can be classified as human. Many such names are now dubbed as synonyms with Homo, including Pithecanthropus, Protanthropus, Sinanthropus, Cyphanthropus, Africanthropus, Telanthropus, Atlanthropus, classifying the genus Homo into species and subspecies is subject to incomplete information and remains poorly done. Some recently extinct species in the genus Homo are only recently discovered, john Edward Gray was an early advocate of classifying taxa by designating tribes and families. Several species, including Australopithecus garhi, Australopithecus sediba, Australopithecus africanus and these species have morphological features that align them with Homo, but there is no consensus as to which gave rise to Homo. The advent of Homo was traditionally taken to coincide with the first use of stone tools, the emergence of Homo also coincides roughly with the onset of Quaternary glaciation, the beginning of the current ice age. A fossil mandible fragment dated to 2.8 million years ago which may represent a stage between Australopithecus and Homo was discovered in 2015 in Afar, Ethiopia. Some authors would push the development of Homo past 3 Mya, the most salient physiological development between the earlier australopithecine species and Homo is the increase in cranial capacity, from about 450 cm3 in A. garhi to 600 cm3 in H. habilis. Within the genus Homo, cranial capacity again doubled from H. habilis through Homo ergaster or H. erectus to Homo heidelbergensis by 0.6 million years ago, the cranial capacity of H. heidelbergensis overlaps with the range found in modern humans. Homo erectus has often been assumed to have developed anagenetically from Homo habilis from about 2 million years ago
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Binomial nomenclature
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Such a name is called a binomial name, a binomen, binominal name or a scientific name, more informally it is also called a Latin name. The first part of the name identifies the genus to which the species belongs, for example, humans belong to the genus Homo and within this genus to the species Homo sapiens. The formal introduction of system of naming species is credited to Carl Linnaeus. But Gaspard Bauhin, in as early as 1623, had introduced in his book Pinax theatri botanici many names of genera that were adopted by Linnaeus. Although the general principles underlying binomial nomenclature are common to these two codes, there are differences, both in the terminology they use and in their precise rules. Similarly, both parts are italicized when a binomial name occurs in normal text, thus the binomial name of the annual phlox is now written as Phlox drummondii. In scientific works, the authority for a name is usually given, at least when it is first mentioned. In zoology Patella vulgata Linnaeus,1758, the original name given by Linnaeus was Fringilla domestica, the parentheses indicate that the species is now considered to belong in a different genus. The ICZN does not require that the name of the person who changed the genus be given, nor the date on which the change was made, in botany Amaranthus retroflexus L. – L. is the standard abbreviation used in botany for Linnaeus. – Linnaeus first named this bluebell species Scilla italica, Rothmaler transferred it to the genus Hyacinthoides, the ICN does not require that the dates of either publication be specified. Prior to the adoption of the binomial system of naming species. Together they formed a system of polynomial nomenclature and these names had two separate functions. First, to designate or label the species, and second, to be a diagnosis or description, such polynomial names may sometimes look like binomials, but are significantly different. For example, Gerards herbal describes various kinds of spiderwort, The first is called Phalangium ramosum, Branched Spiderwort, is aptly termed Phalangium Ephemerum Virginianum, Soon-Fading Spiderwort of Virginia. The Latin phrases are short descriptions, rather than identifying labels, the Bauhins, in particular Caspar Bauhin, took some important steps towards the binomial system, by pruning the Latin descriptions, in many cases to two words. The adoption by biologists of a system of binomial nomenclature is due to Swedish botanist and physician Carl von Linné. It was in his 1753 Species Plantarum that he first began using a one-word trivial name together with a generic name in a system of binomial nomenclature. This trivial name is what is now known as an epithet or specific name
26.
Archaic humans
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A number of varieties of Homo are grouped into the broad category of archaic humans in the period beginning 500,000 years ago. It typically includes Homo neanderthalensis, Homo rhodesiensis, Homo heidelbergensis and this category is contrasted with anatomically modern humans, which include the subspecies Homo sapiens sapiens and Homo sapiens idaltu. Modern humans are theorized to have evolved from humans, who in turn evolved from Homo erectus. Varieties of archaic humans are sometimes included under the binomial name Homo sapiens because their size is very similar to that of modern humans. Archaic humans had a brain size averaging 1200 to 1400 cubic centimeters, archaics are distinguished from anatomically modern humans by having a thick skull, prominent supraorbital ridges and the lack of a prominent chin. Anatomically modern humans appear from about 200,000 years ago and after 70,000 years ago, non-modern varieties of Homo are certain to have survived until after 30,000 years ago, and perhaps until as recent as 10,000 years ago. Which of these, if any, are included under the archaic human is a matter of definition. Nonetheless, according to recent genetic studies, modern humans may have bred with at least two groups of ancient humans, Neanderthals and Denisovans. New evidence suggests another group may also have been extant as recently as 11,500 years ago, chris Stringer of the Natural History Museum in London has suggested that these people could be a result of mating between Denisovans and modern humans. Other scientists remain skeptical, suggesting that the features are within the variations expected for human populations. The category archaic human lacks a single, agreed upon definition, according to one definition, Homo sapiens is a single species comprising several subspecies that include the archaics and modern humans. Under this definition, modern humans are referred to as Homo sapiens sapiens, for example, the Neanderthals are Homo sapiens neanderthalensis, and Homo heidelbergensis is Homo sapiens heidelbergensis. Other taxonomists prefer not to consider archaics and modern humans as a single species, in this case the standard taxonomy is used, i. e. Homo rhodesiensis, or Homo neanderthalensis. The evolutionary dividing lines that separate humans from archaic Homo sapiens. However, these modern humans do possess a number of archaic traits, such as moderate. The emergence of humans is sometimes used as an example of punctuated equilibrium. This occurs when a species undergoes significant biological evolution within a short period. Subsequently, the species undergoes very little change for long periods until the next punctuation, the brain size of archaic humans expanded significantly from 900 cm2 in erectus to 1,300 cm2
27.
Lower Paleolithic
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The Lower Paleolithic is the earliest subdivision of the Paleolithic or Old Stone Age. It spans the time from around 3, the Middle Paleolithic followed the Lower Paleolithic and recorded the appearance of the more advanced prepared-core tool-making technologies such as the Mousterian. Whether the earliest control of fire by hominins dates to the Lower or to the Middle Paleolithic remains an open question. The Lower Paleolithic began with the appearance of the oldest stone tools in the world, roughly 3.3 million ago in eastern Africa, the Gelasian, some 2.5 million years ago, saw the appearance of the Homo genus, possibly developing from australopithecine forebears. Homo habilis is assumed to have lived primarily on scavenging, using tools to cleave meat off carrion or to break bones to extract the marrow, decreasing oceanic evaporation produced a drier climate and the expansion of the savannah at the expense of forests. Reduced availability of fruits stimulated some proto-australopithecines to search out new food sources found in the drier savannah ecology, Homo erectus appeared by about 1.8 million years ago, via the transitional variety Homo ergaster. Homo erectus moved from scavenging to hunting, developing the hunting-gathering lifestyle that would remain dominant throughout the Paleolithic into the Mesolithic, Homo erectus migrated out of Africa and dispersed throughout Eurasia. Stone tools in Malaysia have been dated to be 1.83 million years old, the Peking Man fossil, discovered in 1929, is roughly 700,000 years old. In Europe, the Olduwan tradition split into two parallel traditions, the Clactonian, a tradition, and the Acheulean, a hand-axe tradition. The Levallois technique for knapping flint developed during this time, the carrier species from Africa to Europe was undoubtedly Homo erectus. This type of human is more linked to the flake tradition. Many Mousterian finds in the Middle Paleolithic have been knapped using a Levallois technique, monte Poggiolo, near Forlì, Italy, is the location of an Acheulian littoral handaxe industry dating from 1.8 to 1.1 million years ago. The appearance of Homo heidelbergensis about 600,000 years ago heralds a number of new varieties, such as Homo rhodesiensis. Homo heidelbergensis is a candidate for first developing a form of symbolic language. Whether control of fire and earliest burials date to this period or only appear during the Middle Paleolithic is an open question, the hand-axe tradition originates in the same period. The intermediate may have been Homo heidelbergensis, held responsible for the manufacture of improved Mode 2 Acheulean tool types, in Africa, flakes and axes coexisted in Europe, sometimes at the same site. The axe tradition, however, spread to a different range in the east and it appears in Arabia and India, but more importantly, it does not appear in southeast Asia. Homo sapiens first appear about 200,000 years ago, control of fire by early humans Lomekwi, site of the oldest tools discovered The First People and Culture at Indiana University Bloomington
28.
Western Europe
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Western Europe, or West Europe, is the region comprising the western part of Europe. Below, some different geographic and geopolitical definitions of the term are outlined, prior to the Roman conquest, a large part of Western Europe had adopted the newly developed La Tène culture. This cultural and linguistic division was reinforced by the later political east-west division of the Roman Empire. The division between these two was enhanced during Late Antiquity and the Middle Ages by a number of events, the Western Roman Empire collapsed, starting the Early Middle Ages. By contrast, the Eastern Roman Empire, mostly known as the Greek or Byzantine Empire, survived, in East Asia, Western Europe was historically known as taixi in China and taisei in Japan, which literally translates as the Far West. The term Far West became synonymous with Western Europe in China during the Ming dynasty, the Italian Jesuit priest Matteo Ricci was one of the first writers in China to use the Far West as an Asian counterpart to the European concept of the Far East. In his writings, Ricci referred to himself as Matteo of the Far West, the term was still in use in the late 19th and early 20th centuries. Post-war Europe would be divided into two spheres, the West, influenced by the United States, and the Eastern Bloc. With the onset of the Cold War, Europe was divided by the Iron Curtain, behind that line lie all the capitals of the ancient states of Central and Eastern Europe. Although some countries were neutral, they were classified according to the nature of their political. This division largely defined the popular perception and understanding of Western Europe, the world changed dramatically with the fall of the Iron Curtain in 1989. The Federal Republic of Germany peacefully absorbed the German Democratic Republic, COMECON and the Warsaw Pact were dissolved, and in 1991, the Soviet Union ceased to exist. Several countries which had part of the Soviet Union regained full independence. Although the term Western Europe was more prominent during the Cold War, it remains much in use, in 1948 the Treaty of Brussels was signed between Belgium, France, Luxembourg, the Netherlands and the United Kingdom. It was further revisited in 1954 at the Paris Conference, when the Western European Union was established and it was declared defunct in 2011, after the Treaty of Lisbon, and the Treaty of Brussels was terminated. When the Western European Union was dissolved, it had 10 member countries, six member countries, five observer countries. The CIA divides Western Europe into two smaller subregions, regional voting blocs were formed in 1961 to encourage voting to various UN bodies from different regional groups. The European Union is an economic and political union of 28 member states that are located primarily in Europe, some Western and Northern European countries of Iceland, Norway, Switzerland and Liechtenstein are members of EFTA, though cooperating to varying degree with the European Union
29.
Eudald Carbonell
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Eudald Carbonell i Roura is a Spanish archaeologist, anthropologist and paleontologist. Educated in Girona, Barcelona and Paris, he holds a PhD in Geology of the Quaternary from Pierre and Marie Curie University and his findings include the discovery of Homo antecessor and well-preserved remains of Homo heidelbergensis. Carbonell, E. M. Mosquera, A. Ollé, X. P. Rodríguez Álvarez, M. Sahnouni, R. Sala, structure morphotechnique de lindustrie lithique du Pléistocène inférieur et moyen dAtapuerca. LAnthropologie 105, 259-280 Aguirre, E. and E. Carbonell, early human expansions on Eurasia, The Atapuerca evidence. Bermúdez de Castro, J. M. Carbonell, E & Arsuaga, J. L. Gran Dolina Site, J. of Human Evolution,37, 309-700 Arsuaga, J. L. Lorenzo, C. Carretero, J. M. Gracia, A. Martínez, garcía, N. Bermúdez de Castro, J. M. & Carbonell, E. A complete human pelvis from the Middle Pleistocene of Spain, Carbonell, E. M. Mosquera, et al. Out of Africa, The Dispersal of the Earliest Technical Systems Reconsidered, Carbonell, E. & Vaquero, M. Behavioral Complexity and Biocultural Change in Europe Around Forty Thousand Years Ago. Bermúdez de Castro, J. M. Arsuaga, J. L. Carbonell, E. Rosas, A. Martínez, a Hominid from the Lower Pleistocene of Atapuerca, Spain, Possible Ancestor to Neandertals and Modern Humans. Carbonell, E. & Vaquero, M. eds, the Last Neandertals - The First Anatomically Modern Humans. Cultural Change and Human Evolution, The Crisis at 40 Ka BP, Ed. Carbonell, E. Bermúdez de Castro, J. M. Arsuaga, J. L. Díez, J. C. Rosas, A. Cuenca-Bescos, G. Sala, R. Mosquera, M. & Rodriguez, Lower Pleistocene Hominids and Artifacts from Atapuerca-TD6. Castro-Curel, Z. & Carbonell, E. Wood Pseudomorphs From Level I at Abric Romaní, Barcelona, Carbonell, E. Giralt, S. & Vaquero, M. Abric Romani, Une importante séquence anthropisée du Pléistocene Supérieur. Early Middle Pleistocene deposits and artefacts in the Gran Dolina site of the Sierra de Atapuerca, fundación Atapuerca Institut Català de Paleoecologia Humana i Evolució Social
30.
Juan Luis Arsuaga
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As a child he already showed a great interest in prehistory after reading The Quest for Fire and visiting a dig in nearby Bilbao. The finds at Atapuerca have shed new light on the first humans in Europe and this contrasts with the secretive atmosphere surrounding the digs to near Orce, in southern Spain, which has yielded tools indicating human presence that predate the finds at Atapuerca. In 2013, Arsuaga co-authored a paper which reported the finding of the oldest human DNA ever, the mitochondrial DNA that stemmed from a fossil found in a cave in Sima de los Huesos had similarities to mitochondrial genomes previously found in the extinct Denisovans in Siberia. He has been a lecturer at the universities of London, Cambridge, Zurich, Rome, Arizona, Philadelphia, Berkeley, New York, Tel Aviv, among others. He authored and/or published several publications in Nature, Science, Journal of Archaeological Science, American Journal of Physical Anthropology
31.
Homo heidelbergensis
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Homo heidelbergensis – also Homo rhodesiensis – is an extinct species of the genus Homo that lived in Africa, Europe and western Asia between 600,000 and 200,000 years ago. The skulls of this species share features with both Homo erectus and anatomically modern Homo sapiens, its brain was nearly as large as that of Homo sapiens. The Sima de los Huesos cave at Atapuerca in northern Spain holds particularly rich layers of deposits that represent an exceptional reserve of data where excavations are still in progress. Neanderthals, Denisovans, and modern humans are all considered to have descended from Homo heidelbergensis that appeared around 700,000 years ago in Africa, fossils have been recovered in Ethiopia, Namibia and South Africa. Between 300,000 and 400,000 years ago a group of Homo heidelbergensis migrated into Europe and West Asia via yet unknown routes, archaeological sites exist in Spain, Italy, France, England, Germany, Hungary and Greece. Another Homo heidelbergensis group ventured eastwards into continental Asia, eventually developing into Denisovans, some researchers suggest that the finds associated to Homo heidelbergensis are mere variants of Homo erectus. Both Homo antecessor and Homo heidelbergensis are likely to have descended from the very similar Homo ergaster from Africa. The anatomy is more primitive than that of Neanderthal, but the harmoniously rounded dental arch. Heidelbergensis from other known human species, the species name heidelbergensis only experienced a renaissance with the many discoveries of the past 30 years and appears now to be recognized by an increasing number of researchers. Any form of segregation is considered arbitrary, which is why these researchers forgo the name H. heidelbergensis altogether. Paleoanthropologists often refer to the surrounding the specimens, their dating and morphology, as “the muddle in the middle. ”The fact that there seem to be no clear transitions makes it difficult to draw up a list of unique characteristics of H. heidelbergensis that distinguishes it from H. erectus. In general, the show a continuation of evolutionary trends that are emerging from around the Lower into Middle Pleistocene. Along with changes in the robustness of cranial and dental features, male H. heidelbergensis averaged about 1.75 m tall and 62 kg. Females averaged 1.57 m and 51 kg, the available space would allow for a far greater flexibility of development and It shows a combination of features, which has been previously found neither on a recent nor a fossil human mandible. The actual proof that we are dealing with human parts here only lies within the nature of the dentition, the completely preserved teeth bear the stamp human as evidence, The canines show no trace of a stronger expression in relation to the other groups of teeth. They suggest a moderate and harmonious co-evolution, as it is the case in recent humans, recent findings in a pit in Atapuerca of 28 human skeletons suggest that H. heidelbergensis might have been the first species of the Homo genus to bury its dead. Steven Mithen believes that H. heidelbergensis, like its descendant H. neanderthalensis, acquired a pre-linguistic system of communication. No forms of art have been uncovered, although red ochre, the morphology of the outer and middle ear suggests they had an auditory sensitivity similar to modern humans and very different from chimpanzees
32.
Homo erectus
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Homo erectus is an extinct species of hominid that lived throughout most of the Pleistocene geological epoch. Its earliest fossil evidence dates to 1.9 million years ago and extends to 70,000 years ago, or, possibly, as recently as 35,000 years ago. It is generally thought that H. erectus originated in Africa and spread from there, migrating throughout Eurasia as far as Georgia, India, Sri Lanka, China, a new debate appeared in 2013, with the documentation of the Dmanisi skulls.58 million years ago. From there it migrated, in part, by 2, the fossil record shows that its development from about 1.8 mya to one mya was widely distributed, in Africa, the Transcaucasus, Indonesia, and in Vietnam, China, and India. The second hypothesis is that H. erectus evolved in Eurasia and they occupied the Dmanisi site from 1.85 million to 1.77 million years ago, which was about the same time or slightly before their earliest evidence in Africa. There are several proposed explanations of the dispersal of H. erectus georgicus—including whether or not Africa is the source), the Dutch anatomist Eugène Dubois was fascinated by Darwins theory of evolution especially as it applied to humankind. In 1886, he set out for Asia—which then was the region accepted as the cradle of evolution despite Darwins theory of African origin. In 1891, his team discovered a human fossil on the island of Java, the Java fossil from Indonesia aroused much public interest. It was dubbed by the press as Java Man, but few scientists accepted Dubois argument that his fossil was the transitional form—the so-called missing link—between apes. Java Man is now classified as Homo erectus, most of the spectacular discoveries of H. erectus next took place at the Zhoukoudian Project, now known as the Peking Man Site, in Zhoukoudian, China. This site was first discovered by Johan Gunnar Andersson in 1921 and was first excavated in 1921, canadian anatomist Davidson Blacks initial description of a lower molar as belonging to a previously unknown species prompted widely publicized interest. Extensive excavations followed, which altogether uncovered 200 human fossils from more than 40 individuals including five nearly complete skullcaps, german anatomist Franz Weidenreich provided much of the detailed description of this material in several monographs published in the journal Palaeontologica Sinica. Throughout much of the 20th century, anthropologists debated the role of H. erectus in human evolution, early in the century, due in part to the discoveries at Java and Zhoukoudian, it was widely accepted that modern humans first evolved in Asia. From the 1950s forward, numerous finds in East Africa confirmed the hypothesis of an African genesis and it is now generally accepted that H. erectus descended from either, 1) the earliest hominin genera, or 2) the earliest Homo-species. East Africa provided sympatric coexistence for H. erectus and H, in the 1950s, archaeologists John T. Robinson and Robert Broom named Telanthropus capensis, Robinson had discovered a jaw fragment in 1949 in Swartkrans, South Africa. Later, Simonetta proposed to re-designate it to Homo erectus, in 1961, Yves Coppens discovered a skull of Tchadanthropus uxoris, then the earliest fossil human discovered in north Africa. It was reported that the fossil had been so eroded by sand that it mimicked the appearance of an australopith. Although at first considered to be a specimen of H. habilis, T. uxoris is no longer considered a valid taxon, and has been subsumed into H. erectus
33.
Atapuerca Mountains
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The Atapuerca Mountains, is a karstic hill formation near Atapuerca town in Castile and Leon, northern Spain. The earliest specimen yet unearthed and reliably dated confirm an age between 1.2 Million and 600,000 years, the site was induced into the UNESCO list of World Heritage Sites under the designation Archeological Site of Atapuerca. Stretching alongside the Bureba corridor, a pass that connects the Ebro river valley with the Mediterranean Sea. This conjunction constitutes an ecotone, that is rich in species of both ecosystems, the mountain pass was part of an Roman causeway and the pilgrimage route of Saint James that is now traversed by the N-I and AP-1 highways. Situated strategically in between two major Drainage divides and near the pass is assumed to have been supportive for the successful. The Galería de la Eduarda y el Kolora is a cave that contains parietal rock paintings. Among numerous faunal and floral fossils a jaw fragment was found during the 1970s and a fragment in 1995. They date to between 600,000 and 400,000 years BP, the Gran Dolina site is a huge cavern, which is being excavated since September 1981. Its sediments were divided into eleven stratae TD-11, Mousterian tools found Level TD-10 presumed to have been a Homo heidelbergensis camp with tools, Level TD-8, accessible since 1994, it contained remarkable carnivore fossils. About 25% of the bones have manipulation marks that suggest cannibalism, taxation of these remains is still being debated, suggestions range from Homo erectus to Homo heidelbergensis and Homo antecessor. The Homo erectus-like fossils were found with retouched flake and core stone tools. Level TD-5 is assumed to have been a carnivore den, in TD-4, four lithic pieces were found during the 1991 excavation and several remnants of Ursus dolinensis, a sparsely described bear species. At the lowest levels no fossils Sima de los Huesos accounts for the greatest number of scientific discoveries. This site is located at the bottom of a 13 m deep shaft, associated finds include Ursus deningeri fossils and a hand axe called Excalibur. The idea sparked a renewal of the disputed evolutionary progress and the stages of human cognitive, ninety percent of the known Homo heidelbergensis fossil record have been obtained at the site. The fossil bone pit includes, The complete cranium, Skull 5, nicknamed Miguelón, a complete pelvis, humorously nicknamed Elvis Mandibles, teeth, a lot of postcranial bones Remains of a child with craniosynostosis were found and dated to 530,000 BP. The find was considered to provide evidence for food sharing in human populations. Mitochondrial DNA from a 400,000 year old femur has been sequenced, the mtDNA was found to be closer to the mtDNA of Denisova hominins than to the mtDNA of Neanderthals
34.
Homo ergaster
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It is one of the earliest hominins, which are those hominids that comprise the original members and species of the human clade after splitting from the line of the chimpanzees. Homo ergaster is thought to be ancestral to, or as sharing a common ancestor with, or as being the same species as. Some palaeoanthropologists consider H. ergaster to be a variety of H. erectus, that is, others call H. ergaster the direct ancestor of H. erectus, which then emigrated out of Africa into Eurasia and branched into a distinct species. Still others dispense with the specific epithet ergaster and make no distinctions among fossils assigned to erectus. The latest discoveries at Dmanisi, Georgia, suggest that all the groups of early Homo in Africa, including Homo ergaster, are of the same species. The binomial name was published in 1975 by Groves and Mazák, South African palaeontologist John T. Robinson discovered in 1949 a mandible of a new hominin in southern Africa, which he named Telanthropus capensis and which today is classified as Homo ergaster. That taxon was first applied to a mandible found near Lake Rudolf, Kenya, by Colin Groves and Vratislav Mazák in 1975, dubbed KNM-ER992, it became the type-specimen of the species. A near-complete skeleton of H. ergaster, KNM-WT15000, or Turkana Boy, was discovered in 1984 at Lake Turkana by Kamoya Kimeu and it is dated to 1.6 million years ago and is one of the most complete early hominin fossils found to date. Paleoanthropologists debate the defining of H. ergaster and H. erectus as separate species, thus, although Homo ergaster has gained some acceptance as a valid taxon, ergaster and erectus are often identified as separate populations of the larger species H. erectus. Sura et al declared that Homo erectus was a source of multiple events of gene flow to the Eurasian continent. The major implication of the analyses of the Dmanisi skull and the finds at Dmanisi is that all the earliest varieties of Homo are of one species. This implies that H. ergaster is subsumed under the taxon H. erectus, derived features separating it from earlier non-Homo species include reduced sexual dimorphism, a smaller, more orthognathous face, a smaller dental arcade, and a larger cranial capacity. Remains have been found in Tanzania, Ethiopia, Kenya, there are broad divisions in the scientific community re interpreting the development of the earliest species of genus Homo. H. habilis is generally accepted as the ancestor of Homo. However, habiliss status as a species within Homo is particularly contentious. It is unclear what genetic influence H. ergaster had on later hominins, recent genetic analysis has generally supported the recent-Out-of-Africa hypothesis, and that same kind of analysis may in time designate H. ergaster as the ancestor to all later hominins. H. ergaster is believed to have diverged from the lineage of H. habilis by 1.8 million years ago and these early descendants of H. habilis may have been discovered at Dmanisi, Georgia as Homo erectus georgicus. In 2013, a fragment of fossilized jawbone was discovered in the Ledi-Geraru research area in the Afar depression, Ethiopia
35.
Principle of Priority
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Priority is a fundamental principle of modern botanical nomenclature and zoological nomenclature. Essentially, it is the principle of recognising the first valid application of a name to a plant or animal. There are two aspects to this, The first formal name given to a plant or animal taxon shall be the name that is to be used, called the valid name in zoology. Once a name has been used, no subsequent publication of that name for another taxon shall be valid or validly published, there are formal provisions for making exceptions to this principle. If an archaic or obscure prior name is discovered for a taxon, the current name can be declared a nomen conservandum or conserved name. Similarly, if the current name for a taxon is found to have an archaic or obscure prior homonym, the principle of priority has not always been in place. When Carl Linnaeus laid the foundations of modern nomenclature, he offered no recognition of prior names, the botanists who followed him were just as willing to overturn Linnaeuss names. The first sign of recognition of priority came in 1813, when A. P. de Candolle laid out some principles of good nomenclatural practice and he favoured retaining prior names, but left wide scope for overturning poor prior names. During the 19th century, the principle gradually came to be accepted by almost all botanists, botanists on one side of the debate argued that priority should be universal and without exception. This would have meant a major disruption as countless names in current usage were overturned in favour of archaic prior names. In 1891, Otto Kuntze, one of the most vocal proponents of this position, did just that and he then followed with further such publications in 1893,1898 and 1903. His efforts, however, were so disruptive that they appear to have benefited his opponents, by the 1900s, the need for a mechanism for the conservation of names was widely accepted, and details of such a mechanism were under discussion. The current system of modified priority was essentially put in place at the Cambridge Congress of 1930, the Principle of Priority is one of the guiding principles of the International Code of Zoological Nomenclature, defined by Article 23. There are exceptions, another name may be given precedence by any provision of the Code or by any ruling of the Commission and it is a fundamental guiding precept that preserves the stability of biological nomenclature. It was first formulated in 1842 by an appointed by the British Association to consider the rules of zoological nomenclature. In 1855, John Edward Gray published the name Antilocapra anteflexa for a new species of pronghorn, however, it is now thought that his specimen belonged to an unusual individual of an existing species, Antilocapra americana, with a name published by George Ord in 1815. The older name, by Ord, takes priority, with Antilocapra anteflexa becoming a junior synonym, in 1856, Johann Jakob Kaup published the name Leptocephalus brevirostris for a new species of eel. However, it was realized in 1893 that the described by Kaup was in fact the juvenile form of the European eel
36.
Tomography
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Tomography is imaging by sections or sectioning, through the use of any kind of penetrating wave. The method is used in radiology, archaeology, biology, atmospheric science, geophysics, oceanography, plasma physics, materials science, astrophysics, quantum information, in most cases the production of these images is based on the mathematical procedure tomographic reconstruction. Tomography refers to imaging by sections or sectioning, through the use of any kind of penetrating wave or mechanical method. The method is used in radiology, archaeology, biology, atmospheric science, geophysics, oceanography, plasma physics, materials science, astrophysics, quantum information, and other sciences. In most cases it is based on the procedure called tomographic reconstruction. The word tomography is derived from Ancient Greek τόμος tomos, slice, section and γράφω graphō. In conventional medical X-ray tomography, clinical staff make an image through a body by moving an X-ray source. Consequently, structures in the focal plane appear sharper, while structures in other planes appear blurred, by modifying the direction and extent of the movement, operators can select different focal planes which contain the structures of interest. In a 1953 article in the medical journal Chest, B, pollak of the Fort William Sanatorium described the use of planography, another term for tomography. A device used in tomography is called a tomograph, while the image produced is a tomogram, tomography as the computed tomographic scanner was invented by Sir Godfrey Hounsfield. More modern variations of tomography involve gathering projection data from multiple directions, different types of signal acquisition can be used in similar calculation algorithms in order to create a tomographic image. The term volume imaging might describe these technologies more accurately than the term tomography, however, in the majority of cases in clinical routine, staff request output from these procedures as 2-D slice images. As more and more clinical decisions come to depend on more advanced volume visualization techniques, most algorithms fall into one of two categories, filtered back projection and iterative reconstruction. These procedures give inexact results, they represent a compromise between accuracy and computation time required, FBP demands fewer computational resources, while IR generally produces fewer artifacts at a higher computing cost. Although MRI and ultrasound are transmission methods, they typically do not require movement of the transmitter to acquire data from different directions, in MRI, both projections and higher spatial harmonics are sampled by applying spatially-varying magnetic fields, no moving parts are necessary to generate an image. On the other hand, since ultrasound uses time-of-flight to spatially encode the signal, it is not strictly a tomographic method. In this section, the principle of tomography in the case that especially uses tomography utilizing the parallel beam irradiation optical system will be explained. Tomography is a technology that uses an optical system to obtain virtual slices of specific cross section of a scanned object. There are several types of optical system including the parallel beam irradiation optical system
37.
Human
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Modern humans are the only extant members of Hominina tribe, a branch of the tribe Hominini belonging to the family of great apes. Several of these hominins used fire, occupied much of Eurasia and they began to exhibit evidence of behavioral modernity around 50,000 years ago. In several waves of migration, anatomically modern humans ventured out of Africa, the spread of humans and their large and increasing population has had a profound impact on large areas of the environment and millions of native species worldwide. Humans are uniquely adept at utilizing systems of communication for self-expression and the exchange of ideas. Humans create complex structures composed of many cooperating and competing groups, from families. Social interactions between humans have established a wide variety of values, social norms, and rituals. These human societies subsequently expanded in size, establishing various forms of government, religion, today the global human population is estimated by the United Nations to be near 7.5 billion. In common usage, the word generally refers to the only extant species of the genus Homo—anatomically and behaviorally modern Homo sapiens. In scientific terms, the meanings of hominid and hominin have changed during the recent decades with advances in the discovery, there is also a distinction between anatomically modern humans and Archaic Homo sapiens, the earliest fossil members of the species. The English adjective human is a Middle English loanword from Old French humain, ultimately from Latin hūmānus, the words use as a noun dates to the 16th century. The native English term man can refer to the species generally, the species binomial Homo sapiens was coined by Carl Linnaeus in his 18th century work Systema Naturae. The generic name Homo is a learned 18th century derivation from Latin homō man, the species-name sapiens means wise or sapient. Note that the Latin word homo refers to humans of either gender, the genus Homo evolved and diverged from other hominins in Africa, after the human clade split from the chimpanzee lineage of the hominids branch of the primates. The closest living relatives of humans are chimpanzees and gorillas, with the sequencing of both the human and chimpanzee genome, current estimates of similarity between human and chimpanzee DNA sequences range between 95% and 99%. The gibbons and orangutans were the first groups to split from the leading to the humans. The splitting date between human and chimpanzee lineages is placed around 4–8 million years ago during the late Miocene epoch, during this split, chromosome 2 was formed from two other chromosomes, leaving humans with only 23 pairs of chromosomes, compared to 24 for the other apes. There is little evidence for the divergence of the gorilla, chimpanzee. Each of these species has been argued to be an ancestor of later hominins
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Occipital bun
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An occipital bun is a prominent bulge or projection of the occipital bone at the back of the skull. It is important in descriptions of classic Neanderthal crania. While common among many of humankinds ancestors, primarily robust relatives rather than gracile, some scientists suspect occipital buns might correlate with the biomechanics of running. Another theory attributes them to enlargement of the cerebellum, a region of the brain which mediates the timing of motor actions, there are still some human populations which often exhibit occipital buns. A greater proportion of early modern Europeans had them, but prominent occipital buns in modern populations are now relatively infrequent, bone terminology Terms for anatomical location PBS. org - Neanderthals on Trial
39.
Turkana Boy
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Turkana Boy, also called Nariokotome Boy, is the common name of Homo erectus fossil KNM-WT15000, a nearly complete skeleton of a hominin youth who lived during the early Pleistocene. This specimen is the most complete human skeleton ever found. It is believed to be between 1.5 and 1.6 million years old, estimates of his age at death range from seven to 18 years old, the most recent scientific review suggests eight years. It was initially suggested that he would have grown into a 185 centimetres tall adult, the reason for this shift has been research showing that his growth maturation differed from that of modern humans in that he would have had a briefer and smaller adolescent growth spurt. The skeleton was discovered in 1984 by Kamoya Kimeu, a member of a team led by Richard Leakey, the shape of the pelvis identifies the specimen as male. Estimates of the age at death depend on whether the maturity stage of the teeth or skeleton is used, a key factor is that, while modern humans have a marked adolescent growth spurt, chimpanzees do not. While initial research assumed a modern type of growth, more recent evidence from other fossils suggests this was less present in early Homo. This affects the estimation of both his age and his stature as a fully grown adult. Anthropologists Alan Walker and Richard Leakey in 1993 estimated the boy to have been about 11–12 years old based on rates of bone maturity. Christopher Dean of University College London, in a Nova special, but Alan Walker and Richard Leakey said that dental dating often gives a younger age than a persons actual age. According to this scenario, KNM-WT15000 would have attained an adult stature ranging between 159 centimetres and 168 centimetres. Moreover, that according to our models of growth and development, growth in stature completed by 12 years of age. The specimen comprises 108 bones, making it the most complete human skeleton discovered. The skeleton is about 160 centimetres tall, in adulthood, Turkana Boy might have reached 185 centimetres tall and massed 68 kilograms. The pelvis is narrower than in Homo sapiens, which is most likely for more efficient upright walking and this further indicates a fully terrestrial bipedalism, which is unlike older hominin species that show a combined feature of bipedalism and tree climbing. The Boy was relatively tall, which increased his body surface area that would enhance heat dissipation, the overall KNM-WT15000 skeleton still had features not seen in H. sapiens. However, there are significant defining characters, such as brain size. The arms and legs are slightly longer indicating effective bipedality, the nose is projecting like those of humans rather than the open flat nose seen in apes
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Cannibalism
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In zoology, cannibalism is the act of one individual of a species consuming all or part of another individual of the same species as food. To consume the same species or show cannibalism is an ecological interaction in the animal kingdom and has been recorded for more than 1,500 species. It does not, as believed, occur only as a result of extreme food shortages or artificial conditions. Cannibalism seems to be prevalent in aquatic communities, in which up to approximately 90% of the organisms engage in cannibalism at some point of the life cycle. Cannibalism is also not restricted to species, but is commonly found in herbivores and detritivores. Sexual cannibalism is a case of cannibalism in which a female organism kills and consumes a conspecific male before, during. Sexual cannibalism has been recorded in the female spider, black widow spider, praying mantis. Size-structured cannibalism is cannibalism in which older, larger, more mature individuals consume smaller, younger conspecifics, in size-structured populations, cannibalism can be responsible for 8% to 95% of the total mortality, making it a significant and important factor for population and community dynamics. Size-structured cannibalism has commonly been observed in the wild for a variety of taxa, vertebrate examples include chimpanzees, where groups of adult males have been observed to attack and consume infants. Filial cannibalism is a type of size-structured cannibalism in which adults eat their own offspring. Though most often thought of as parents eating live young, filial cannibalism includes parental consumption of stillborn infants and miscarried fetuses as well as infertile, vertebrate examples include pigs, where savaging accounts for a sizable percentage of total piglet deaths, and cats. Filial cannibalism is common in teleost fishes, appearing in at least seventeen different families of teleosts. Within this diverse group of fish, there have been many, one of these is the energy-based hypothesis, which suggests that fish eat their offspring when they are low on energy as an investment in future reproductive success. In other words, when males of a species are low on energy, it might sometimes be beneficial for them to feed on their own offspring to survive. Another hypothesis as to the value of filial cannibalism in teleosts is that it increases density-dependent egg survivorship. In other words, filial cannibalism simply increases overall reproductive success by helping the other eggs make it to maturity by thinning out the numbers. Possible explanations as to why this is so include increasing oxygen availability to the eggs, the negative effects of accumulating embryo waste. In some species of wasps, such as P. chinensis
41.
England
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England is a country that is part of the United Kingdom. It shares land borders with Scotland to the north and Wales to the west, the Irish Sea lies northwest of England and the Celtic Sea lies to the southwest. England is separated from continental Europe by the North Sea to the east, the country covers five-eighths of the island of Great Britain in its centre and south, and includes over 100 smaller islands such as the Isles of Scilly, and the Isle of Wight. England became a state in the 10th century, and since the Age of Discovery. The Industrial Revolution began in 18th-century England, transforming its society into the worlds first industrialised nation, Englands terrain mostly comprises low hills and plains, especially in central and southern England. However, there are uplands in the north and in the southwest, the capital is London, which is the largest metropolitan area in both the United Kingdom and the European Union. In 1801, Great Britain was united with the Kingdom of Ireland through another Act of Union to become the United Kingdom of Great Britain and Ireland. In 1922 the Irish Free State seceded from the United Kingdom, leading to the latter being renamed the United Kingdom of Great Britain, the name England is derived from the Old English name Englaland, which means land of the Angles. The Angles were one of the Germanic tribes that settled in Great Britain during the Early Middle Ages, the Angles came from the Angeln peninsula in the Bay of Kiel area of the Baltic Sea. The earliest recorded use of the term, as Engla londe, is in the ninth century translation into Old English of Bedes Ecclesiastical History of the English People. According to the Oxford English Dictionary, its spelling was first used in 1538. The earliest attested reference to the Angles occurs in the 1st-century work by Tacitus, Germania, the etymology of the tribal name itself is disputed by scholars, it has been suggested that it derives from the shape of the Angeln peninsula, an angular shape. An alternative name for England is Albion, the name Albion originally referred to the entire island of Great Britain. The nominally earliest record of the name appears in the Aristotelian Corpus, specifically the 4th century BC De Mundo, in it are two very large islands called Britannia, these are Albion and Ierne. But modern scholarly consensus ascribes De Mundo not to Aristotle but to Pseudo-Aristotle, the word Albion or insula Albionum has two possible origins. Albion is now applied to England in a poetic capacity. Another romantic name for England is Loegria, related to the Welsh word for England, Lloegr, the earliest known evidence of human presence in the area now known as England was that of Homo antecessor, dating to approximately 780,000 years ago. The oldest proto-human bones discovered in England date from 500,000 years ago, Modern humans are known to have inhabited the area during the Upper Paleolithic period, though permanent settlements were only established within the last 6,000 years
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France
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France, officially the French Republic, is a country with territory in western Europe and several overseas regions and territories. The European, or metropolitan, area of France extends from the Mediterranean Sea to the English Channel and the North Sea, Overseas France include French Guiana on the South American continent and several island territories in the Atlantic, Pacific and Indian oceans. France spans 643,801 square kilometres and had a population of almost 67 million people as of January 2017. It is a unitary republic with the capital in Paris. Other major urban centres include Marseille, Lyon, Lille, Nice, Toulouse, during the Iron Age, what is now metropolitan France was inhabited by the Gauls, a Celtic people. The area was annexed in 51 BC by Rome, which held Gaul until 486, France emerged as a major European power in the Late Middle Ages, with its victory in the Hundred Years War strengthening state-building and political centralisation. During the Renaissance, French culture flourished and a colonial empire was established. The 16th century was dominated by civil wars between Catholics and Protestants. France became Europes dominant cultural, political, and military power under Louis XIV, in the 19th century Napoleon took power and established the First French Empire, whose subsequent Napoleonic Wars shaped the course of continental Europe. Following the collapse of the Empire, France endured a succession of governments culminating with the establishment of the French Third Republic in 1870. Following liberation in 1944, a Fourth Republic was established and later dissolved in the course of the Algerian War, the Fifth Republic, led by Charles de Gaulle, was formed in 1958 and remains to this day. Algeria and nearly all the colonies became independent in the 1960s with minimal controversy and typically retained close economic. France has long been a centre of art, science. It hosts Europes fourth-largest number of cultural UNESCO World Heritage Sites and receives around 83 million foreign tourists annually, France is a developed country with the worlds sixth-largest economy by nominal GDP and ninth-largest by purchasing power parity. In terms of household wealth, it ranks fourth in the world. France performs well in international rankings of education, health care, life expectancy, France remains a great power in the world, being one of the five permanent members of the United Nations Security Council with the power to veto and an official nuclear-weapon state. It is a member state of the European Union and the Eurozone. It is also a member of the Group of 7, North Atlantic Treaty Organization, Organisation for Economic Co-operation and Development, the World Trade Organization, originally applied to the whole Frankish Empire, the name France comes from the Latin Francia, or country of the Franks
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Tarragona
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Tarragona is a port city located in northeast Spain on the Costa Daurada by the Mediterranean Sea. It is the capital of the Province of Tarragona, and part of Tarragonès, geographically, it is bordered on the north by the Province of Barcelona and the Province of Lleida. The city has a population of 132,199, the real founding date of Tarragona is unknown. The city may have begun as an Iberic town called Kesse or Kosse, named for the Iberic tribe of the region, william Smith suggests that the city was probably founded by the Phoenicians, who called it Tarchon, which, according to Samuel Bochart, means a citadel. This name was derived from its situation on a high rock. It was seated on the river Sulcis or Tulcis, on a bay of the Mare Internum, the city was first named Colonia Iulia Urbs Triumphalis Tarraco and was capital of the province of Hispania Citerior. Subsequently, it became the capital of the named after it, Hispania Tarraconensis, in the Roman Empire. Tarraco lies on the road along the southeastern coast of the Iberian Peninsula. According to Mela it was the richest town on that coast and its fertile plain and sunny shores are celebrated by Martial and other poets, and its neighbourhood is described as producing good wine and flax. An inscribed stone base for a now lost statue of Tiberius Claudius Candidus was found in Tarragona during the nineteenth century and this important marble block was purchased by the British Museum in 1994. After the demise of the Western Roman Empire, it was captured first by the Vandals, the Visigothic Kingdoms rule of Tarracona was ended by the Umayyad conquest of Hispania in 714. It was an important border city of the Caliphate of Córdoba between 750 and 1013, after the demise of the Caliphate, it was part of the Taifa of Zaragoza between 1013 and 1110 and under the control of the Almoravid dynasty between 1110 and 1117. It was taken by the County of Barcelona in 1117, after the dynastic union of Aragon and Barcelona, it was part of the Kingdom of Aragon from 1164-1412. After dynastic union of Aragon and the Crown of Castile, it remained a part of Aragon until the foundation of the Spanish Empire in 1516. During the Catalan Revolt, Tarragon was captured by Catalan insurgents with French support in 1641 and it was captured by allied Portuguese, Dutch, and British troops in 1705 during the War of the Spanish Succession and remained in their hands until Treaty of Utrecht in 1713. During the war, the Catalans supported the claim of Archduke Charles, Duke of Teschen against the victorious Bourbon Duke of Anjou. He signed the Nueva Planta decrees, which abolished the Crown of Aragon and all remaining Catalan institutions, a British naval squadron commanded by Admiral Edward Codrington harassed the French besiegers with cannon fire and transported large numbers of reinforcements into the city by sea. Nevertheless, Suchets troops stormed into the defenses and killed or captured almost all the defenders and it became a subprefecture center in Bouches-de-lÈbre department of French empire
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Spain
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By population, Spain is the sixth largest in Europe and the fifth in the European Union. Spains capital and largest city is Madrid, other urban areas include Barcelona, Valencia, Seville, Bilbao. Modern humans first arrived in the Iberian Peninsula around 35,000 years ago, in the Middle Ages, the area was conquered by Germanic tribes and later by the Moors. Spain is a democracy organised in the form of a government under a constitutional monarchy. It is a power and a major developed country with the worlds fourteenth largest economy by nominal GDP. Jesús Luis Cunchillos argues that the root of the span is the Phoenician word spy. Therefore, i-spn-ya would mean the land where metals are forged, two 15th-century Spanish Jewish scholars, Don Isaac Abravanel and Solomon ibn Verga, gave an explanation now considered folkloric. Both men wrote in two different published works that the first Jews to reach Spain were brought by ship by Phiros who was confederate with the king of Babylon when he laid siege to Jerusalem. This man was a Grecian by birth, but who had given a kingdom in Spain. He became related by marriage to Espan, the nephew of king Heracles, Heracles later renounced his throne in preference for his native Greece, leaving his kingdom to his nephew, Espan, from whom the country of España took its name. Based upon their testimonies, this eponym would have already been in use in Spain by c.350 BCE, Iberia enters written records as a land populated largely by the Iberians, Basques and Celts. Early on its coastal areas were settled by Phoenicians who founded Western Europe´s most ancient cities Cadiz, Phoenician influence expanded as much of the Peninsula was eventually incorporated into the Carthaginian Empire, becoming a major theater of the Punic Wars against the expanding Roman Empire. After an arduous conquest, the peninsula came fully under Roman Rule, during the early Middle Ages it came under Germanic rule but later, much of it was conquered by Moorish invaders from North Africa. In a process took centuries, the small Christian kingdoms in the north gradually regained control of the peninsula. The last Moorish kingdom fell in the same year Columbus reached the Americas, a global empire began which saw Spain become the strongest kingdom in Europe, the leading world power for a century and a half, and the largest overseas empire for three centuries. Continued wars and other problems led to a diminished status. The Napoleonic invasions of Spain led to chaos, triggering independence movements that tore apart most of the empire, eventually democracy was peacefully restored in the form of a parliamentary constitutional monarchy. Spain joined the European Union, experiencing a renaissance and steady economic growth
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Complutense University of Madrid
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The Complutense University of Madrid is a public research university located in Madrid, and one of the oldest universities in the world. The university enrolls over 86,000 students, and consistently ranks as one of the top universities in Spain, according to the Spanish newspaper El Mundo, the university is widely regarded as the most prestigious academic institution in Spain. In the course of seven centuries, the University of Madrid has provided invaluable contributions in the sciences, fine arts. Alumni include renowned philosophers, writers, scientists, historians, military leaders, foreign leaders, in the year 1785, the University of Madrid became one of the first Universities in the world to grant a Doctorate degree to a female student. By Royal Decree of 1857, the University of Madrid was the institution in Spain authorized to grant doctorates throughout the Spanish Empire. On 13 April 1499, Cardinal Cisneros secured from Pope Alexander VI a Papal bull to expand Complutense into a full university and this Papal Bull conferred official recognition throughout Christendom to all degrees granted by the University. It also renamed the institution Universitas Complutensis, after Complutum, which was the Latin name of Alcalá de Henares, in the 1509–1510 school year, the Complutense University already operated with five major schools, Arts and Philosophy, Theology, Canon law, Philology and Medicine. During the 16th and 17th centuries, Complutense University became one of the greatest centers of Academic excellence in the world, many of the leading figures in science, arts and politics of that Age studied or taught in Complutenses classrooms. Special colleges were created for students of foreign origin, such as Flemish or Irish, in 1785, Complutense became one of the first Universities in the world to grant a Doctorate to a female student, María Isidra de Guzmán y de la Cerda. In comparison, University of Oxford did not accept female scholars until the year 1920, in 1824, Francisco Tadeo Calomarde further expanded Complutense by merging it with the University of Sigüenza. The University would be known under this name until its name of Complutense was restored in the 1970s. The University of Madrid awarded Albert Einstein a Doctor of Science degree Honoris Causa on 28 February 1923 and this was the first Doctor of Science degree Honoris Causa that Albert Einstein accepted from a European University. However, as the situation began to deteriorate throughout Europe. The University greatly expanded during the 19th century, and its accommodations in central Madrid proved to be increasingly inadequate. This situation changed in 1927, when by royal decree King Alfonso XIII officially ceded state-held lands in the proximity of the Palace of La Moncloa to establish space for the University of Madrid. A council appointed by King Alfonso XIII had decided that the new University of Madrid would require the innovative architecture and planning. Indeed, the buildings, exemplary amongst them the Schools of Medicine, Pharmacy and Odontology, are an homage to structural functionalism. The city of Alcalá de Henares decided to open a university within the campus buildings
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Burgos
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Burgos is a city in northern Spain and the historic capital of Castile. It is situated on the confluence of the Arlanzón river tributaries and it has about 180,000 inhabitants in the actual city and another 20,000 in the metropolitan area. It is the capital of the province of Burgos, in the community of Castile. A large number of churches, palaces and other buildings from the medieval age remain, the city is surrounded by the Fuentes Blancas and the Paseo de la Isla parks. The city forms the principal crossroad of northern Spain along the Camino de Santiago and it has a well-developed transportation system, forming the main communication node in northern Spain. In 2008, the international Burgos Airport started to offer commercial flights, furthermore, AVE high speed trains are planned to start service in the near future, stopping at the newly-built Rosa de Lima train station. The Museum of Human Evolution was opened in 2010, unique in its kind across the world, the museum features the first Europeans, which lived in this area 800,000 years ago. Burgos was selected as the Spanish Gastronomy Capital of 2013, there are several possible origins for the toponymy. When the city was founded, the inhabitants of the country moved into the fortified village. The city began to be called Caput Castellae, early humans occupied sites around Burgos as early as 800,000 years ago. When the Romans took possession of what is now the province of Burgos, in Roman times, it belonged to Hispania Citerior and then to Hispania Tarraconensis. The region came to be known as Castile, i. e. land of castles, the city began to be called Caput Castellae. The county of Burgos, subject to the Kings of León, continued to be governed by counts and was extended, one of these counts, Fernán González. In the 11th century, the city became the see of the Roman Catholic Diocese of Burgos, throughout the 13th and 14th centuries, Burgos was a favourite seat of the kings of León and Castile and a favoured burial site. In 1285, Sancho IV added a new body to the consejo which came to dominate it, the jurado in charge of collecting taxes and overseeing public works, the alguacil was the royal official instituted to judge disagreements. On 9 June 1345, sweeping aside the city government, Alfonso XI established direct rule of Burgos through the Regimiento of sixteen appointed men. In 1574, Pope Gregory XIII made the bishopric a Metropolitan archbishorpic, Burgos has been the scene of many wars, with the Moors, the struggles between León and Navarre, and between Castile and Aragon. In the Peninsular War against Napoleonic France, the siege of Burgos was a scene of a withdrawal for Arthur Wellesley, again in the 19th-century Carlist civil wars of the Spanish succession Burgos was the scene of a battle
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