Saurischia is one of the two basic divisions of dinosaurs. ‘Saurischia’ translates to lizard-hipped. In 1888, Harry Seeley classified dinosaurs into two orders, based on their hip structure, though today most paleontologists classify Saurischia as an unranked clade rather than an order. All carnivorous dinosaurs are traditionally classified as saurischians, as are all of the birds and one of the two primary lineages of herbivorous dinosaurs, the sauropodomorphs. At the end of the Cretaceous Period, all saurischians except the birds became extinct in the course of the Cretaceous–Paleogene extinction event. Birds, as direct descendants of one group of theropod dinosaurs, are a sub-clade of saurischian dinosaurs in phylogenetic classification. Saurischian dinosaurs are traditionally distinguished from ornithischian dinosaurs by their three-pronged pelvic structure, with the pubis pointed forward; the ornithischians' pelvis is arranged with the pubis rotated backward, parallel with the ischium also with a forward-pointing process, giving a four-pronged structure.
The saurischian hip structure led Seeley to name them "lizard-hipped" dinosaurs, because they retained the ancestral hip anatomy found in modern lizards and other reptiles. He named ornithischians "bird-hipped" dinosaurs because their hip arrangement was superficially similar to that of birds, though he did not propose any specific relationship between ornithischians and birds. However, in the view which has long been held, this "bird-hipped" arrangement evolved several times independently in dinosaurs, first in the ornithischians in the lineage of saurischians including birds, lastly in the therizinosaurians; this would be an example of convergent evolution, therizinosaurians, ornithischian dinosaurs all developed a similar hip anatomy independently of each other as an adaptation to their herbivorous or omnivorous diets. In his paper naming the two groups, Seeley reviewed previous classification schemes put forth by other paleontologists to divide up the traditional order Dinosauria, he preferred one, put forward by Othniel Charles Marsh in 1878, which divided dinosaurs into four orders: Sauropoda, Theropoda and Stegosauria.
Seeley, wanted to formulate a classification that would take into account a single primary difference between major dinosaurian groups based on a characteristic that differentiated them from other reptiles. He found this in the configuration of the hip bones, found that all four of Marsh's orders could be divided neatly into two major groups based on this feature, he placed the Stegosauria and Ornithopoda in the Ornithischia, the Theropoda and Sauropoda in the Saurischia. Furthermore, Seeley used this major difference in the hip bones, along with many other noted differences between the two groups, to argue that "dinosaurs" were not a natural grouping at all, but rather two distinct orders that had arisen independently from more primitive archosaurs; this concept that "dinosaur" was an outdated term for two distinct orders lasted many decades in the scientific and popular literature, it was not until the 1960s that scientists began to again consider the possibility that saurischians and ornithischians were more related to each other than they were to other archosaurs.
Although his concept of a polyphyletic Dinosauria is no longer accepted by most paleontologists, Seeley's basic division of the two dinosaurian groups has stood the test of time, has been supported by modern cladistic analysis of relationships among dinosaurs. One alternative hypothesis challenging Seeley's classification was proposed by Robert T. Bakker in his 1986 book The Dinosaur Heresies. Bakker's classification separated the theropods into their own group and placed the two groups of herbivorous dinosaurs together in a separate group he named the Phytodinosauria; the Phytodinosauria hypothesis was based on the supposed link between ornithischians and prosauropods, the idea that the former had evolved directly from the latter by way of an enigmatic family that seemed to possess characters of both groups, the segnosaurs. However, it was found that segnosaurs were an unusual type of herbivorous theropod saurischian related to birds, the Phytodinosauria hypothesis fell out of favor. A 2017 study by Dr Matthew Grant Baron, Dr David B. Norman and Prof. Paul M. Barrett did not find support for a monophyletic Saurischia, according to its traditional definition.
Instead, the group was found to be paraphyletic, with Theropoda removed from the group and placed as the sister group to the Ornithischia in the newly defined clade Ornithoscelida. As a result, the authors redefined Saurischia as "the most inclusive clade that contains D. carnegii, but not T. horridus", resulting in a clade containing only the Sauropodomorpha and Herrerasauridae
In the geologic timescale, the Sinemurian is an age and stage in the Early or Lower Jurassic epoch or series. It spans the time between 199.3 ± 2 Ma and 190.8 ± 1.5 Ma. The Sinemurian is followed by the Pliensbachian. In Europe the Sinemurian age, together with the Hettangian age, saw the deposition of the lower Lias, in Great Britain known as the Blue Lias; the Sinemurian stage was defined and introduced into scientific literature by French palaeontologist Alcide d'Orbigny in 1842. It takes its name from the French town near Dijon; the calcareous soil formed from the Jurassic limestone of the region is in part responsible for the character of the classic Sancerre wines. The base of the Sinemurian stage is at the first appearance of the ammonite genera Vermiceras and Metophioceras in the stratigraphic record. A global reference profile for the Sinemurian stage is located in a cliff north of the hamlet of East Quantoxhead, 6 kilometres east of Watchet, England; the top of the Sinemurian is at the first appearances of the ammonite species Bifericeras donovani and ammonite genus Apoderoceras.
The Sinemurian contains six ammonite biozones in the Tethys domain: zone of Echioceras raricostatum zone of Oxynotoceras oxynotum zone of Asteroceras obtusum zone of Caenisites turneri zone of Arnioceras semicostatum zone of Arietites bucklandi Bloos, G. & Page, K. N.. M.. G. & Smith, A. G.. D´Orbigny, A. C. V. M. D.. 1. Terrains oolitiques ou jurassiques, Paris. Komlosaurus carbonis GeoWhen Database - Sinemurian Lower Jurassic timescale, at the website of the subcommission for stratigraphic information of the ICS Stratigraphic chart of the Lower Jurassic, at the website of Norges Network of offshore records of geology and stratigraphy
The Triassic is a geologic period and system which spans 50.6 million years from the end of the Permian Period 251.9 million years ago, to the beginning of the Jurassic Period 201.3 Mya. The Triassic is the shortest period of the Mesozoic Era. Both the start and end of the period are marked by major extinction events. Triassic began in the wake of the Permian–Triassic extinction event, which left the Earth's biosphere impoverished. Therapsids and archosaurs were the chief terrestrial vertebrates during this time. A specialized subgroup of archosaurs, called dinosaurs, first appeared in the Late Triassic but did not become dominant until the succeeding Jurassic Period; the first true mammals, themselves a specialized subgroup of therapsids evolved during this period, as well as the first flying vertebrates, the pterosaurs, like the dinosaurs, were a specialized subgroup of archosaurs. The vast supercontinent of Pangaea existed until the mid-Triassic, after which it began to rift into two separate landmasses, Laurasia to the north and Gondwana to the south.
The global climate during the Triassic was hot and dry, with deserts spanning much of Pangaea's interior. However, the climate became more humid as Pangaea began to drift apart; the end of the period was marked by yet another major mass extinction, the Triassic–Jurassic extinction event, that wiped out many groups and allowed dinosaurs to assume dominance in the Jurassic. The Triassic was named in 1834 by Friedrich von Alberti, after the three distinct rock layers that are found throughout Germany and northwestern Europe—red beds, capped by marine limestone, followed by a series of terrestrial mud- and sandstones—called the "Trias"; the Triassic is separated into Early and Late Triassic Epochs, the corresponding rocks are referred to as Lower, Middle, or Upper Triassic. The faunal stages from the youngest to oldest are: During the Triassic all the Earth's land mass was concentrated into a single supercontinent centered more or less on the equator and spanning from pole to pole, called Pangaea.
From the east, along the equator, the Tethys sea penetrated Pangaea, causing the Paleo-Tethys Ocean to be closed. In the mid-Triassic a similar sea penetrated along the equator from the west; the remaining shores were surrounded by the world-ocean known as Panthalassa. All the deep-ocean sediments laid down during the Triassic have disappeared through subduction of oceanic plates; the supercontinent Pangaea was rifting during the Triassic—especially late in that period—but had not yet separated. The first nonmarine sediments in the rift that marks the initial break-up of Pangaea, which separated New Jersey from Morocco, are of Late Triassic age. S. these thick sediments comprise the Newark Group. Because a super-continental mass has less shoreline compared to one broken up, Triassic marine deposits are globally rare, despite their prominence in Western Europe, where the Triassic was first studied. In North America, for example, marine deposits are limited to a few exposures in the west, thus Triassic stratigraphy is based on organisms that lived in lagoons and hypersaline environments, such as Estheria crustaceans.
At the beginning of the Mesozoic Era, Africa was joined with Earth's other continents in Pangaea. Africa shared the supercontinent's uniform fauna, dominated by theropods and primitive ornithischians by the close of the Triassic period. Late Triassic fossils are more common in the south than north; the time boundary separating the Permian and Triassic marks the advent of an extinction event with global impact, although African strata from this time period have not been studied. During the Triassic peneplains are thought to have formed in what is now southern Sweden. Remnants of this peneplain can be traced as a tilted summit accordance in the Swedish West Coast. In northern Norway Triassic peneplains may have been buried in sediments to be re-exposed as coastal plains called strandflats. Dating of illite clay from a strandflat of Bømlo, southern Norway, have shown that landscape there became weathered in Late Triassic times with the landscape also being shaped during that time. At Paleorrota geopark, located in Rio Grande do Sul, the Santa Maria Formation and Caturrita Formations are exposed.
In these formations, one of the earliest dinosaurs, Staurikosaurus, as well as the mammal ancestors Brasilitherium and Brasilodon have been discovered. The Triassic continental interior climate was hot and dry, so that typical deposits are red bed sandstones and evaporites. There is no evidence of glaciation near either pole. Pangaea's large size limited the moderating effect of the global ocean; the strong contrast between the Pangea supercontinent and the global ocean triggered intense cross-equatorial monsoons. The Triassic may have been a dry period, but evidence exists that it was punctuated by several episodes of increased rainfall in tropical and subtropical latitudes of the Tethys Sea and its surrounding land. Sediments and fossils suggestive of a more humid climate are known from the Anisian to Ladinian of the Tethysian domain, from the Carnian and Rhaetian of a larger area that includes the Boreal domain, the North
The Precambrian is the earliest part of Earth's history, set before the current Phanerozoic Eon. The Precambrian is so named because it preceded the Cambrian, the first period of the Phanerozoic eon, named after Cambria, the Latinised name for Wales, where rocks from this age were first studied; the Precambrian accounts for 88% of the Earth's geologic time. The Precambrian is an informal unit of geologic time, subdivided into three eons of the geologic time scale, it spans from the formation of Earth about 4.6 billion years ago to the beginning of the Cambrian Period, about 541 million years ago, when hard-shelled creatures first appeared in abundance. Little is known about the Precambrian, despite it making up seven-eighths of the Earth's history, what is known has been discovered from the 1960s onwards; the Precambrian fossil record is poorer than that of the succeeding Phanerozoic, fossils from the Precambrian are of limited biostratigraphic use. This is because many Precambrian rocks have been metamorphosed, obscuring their origins, while others have been destroyed by erosion, or remain buried beneath Phanerozoic strata.
It is thought that the Earth coalesced from material in orbit around the Sun at 4,543 Ma, may have been struck by a large planetesimal shortly after it formed, splitting off material that formed the Moon. A stable crust was in place by 4,433 Ma, since zircon crystals from Western Australia have been dated at 4,404 ± 8 Ma; the term "Precambrian" is recognized by the International Commission on Stratigraphy as the only "supereon" in geologic time. "Precambrian" is still used by geologists and paleontologists for general discussions not requiring the more specific eon names. As of 2010, the United States Geological Survey considers the term informal, lacking a stratigraphic rank. A specific date for the origin of life has not been determined. Carbon found in 3.8 billion-year-old rocks from islands off western Greenland may be of organic origin. Well-preserved microscopic fossils of bacteria older than 3.46 billion years have been found in Western Australia. Probable fossils 100 million years older have been found in the same area.
However, there is evidence. There is a solid record of bacterial life throughout the remainder of the Precambrian. Excluding a few contested reports of much older forms from North America and India, the first complex multicellular life forms seem to have appeared at 1500 Ma, in the Mesoproterozoic era of the Proterozoic eon. Fossil evidence from the Ediacaran period of such complex life comes from the Lantian formation, at least 580 million years ago. A diverse collection of soft-bodied forms is found in a variety of locations worldwide and date to between 635 and 542 Ma; these are referred to as Vendian biota. Hard-shelled creatures appeared toward the end of that time span, marking the beginning of the Phanerozoic eon. By the middle of the following Cambrian period, a diverse fauna is recorded in the Burgess Shale, including some which may represent stem groups of modern taxa; the increase in diversity of lifeforms during the early Cambrian is called the Cambrian explosion of life. While land seems to have been devoid of plants and animals and other microbes formed prokaryotic mats that covered terrestrial areas.
Tracks from an animal with leg like appendages have been found in what was mud 551 million years ago. Evidence of the details of plate motions and other tectonic activity in the Precambrian has been poorly preserved, it is believed that small proto-continents existed prior to 4280 Ma, that most of the Earth's landmasses collected into a single supercontinent around 1130 Ma. The supercontinent, known as Rodinia, broke up around 750 Ma. A number of glacial periods have been identified going as far back as the Huronian epoch 2400–2100 Ma. One of the best studied is the Sturtian-Varangian glaciation, around 850–635 Ma, which may have brought glacial conditions all the way to the equator, resulting in a "Snowball Earth"; the atmosphere of the early Earth is not well understood. Most geologists believe it was composed of nitrogen, carbon dioxide, other inert gases, was lacking in free oxygen. There is, evidence that an oxygen-rich atmosphere existed since the early Archean. At present, it is still believed that molecular oxygen was not a significant fraction of Earth's atmosphere until after photosynthetic life forms evolved and began to produce it in large quantities as a byproduct of their metabolism.
This radical shift from a chemically inert to an oxidizing atmosphere caused an ecological crisis, sometimes called the oxygen catastrophe. At first, oxygen would have combined with other elements in Earth's crust iron, removing it from the atmosphere. After the supply of oxidizable surfaces ran out, oxygen would have begun to accumulate in the atmosphere, the modern high-oxygen atmosphere would have developed. Evidence for this lies in older rocks that contain massive banded iron formations that were laid down as iron oxides. A terminology has evolved covering the early years of the Earth's existence, as radiometric dating has allowed real dates to be assigned to specific formations and features; the Precambrian is divided into
Ohmdenosaurus is the name given to a genus of herbivorous dinosaur from the Early Jurassic. It was a small vulcanodontid sauropod which lived in Germany. Only a couple of fragmentary leg bones were found. In the 1970s, German palaeontologist Rupert Wild, visiting the Urwelt-Museum Hauff at Holzmaden, noticed a fossil in a display labelled as a plesiosaur which he recognised to be a dinosaur bone instead, it proved to be impossible to establish the exact provenance of the remains. In 1978 it was described by him as the type species Ohmdenosaurus liasicus; the generic name refers to Ohmden, a town in Baden-Württemberg near the quarry the remains were found in. The specific name refers to an old name for the Early Jurassic; the holotype, which lacked an inventory number, was found in the famous Posidonia Shale, marine strata dating from the middle Toarcian, as can be established from the presence of the snail Coelodiscus in the matrix rock containing the only prepared bones. It consists of an astragalus and a calcaneus.
The bones, disarticulated in the fossil, show signs of weathering, evidence that the animal died on land and that only its bones were washed into the sea. The tibia is only 405 millimetres long; the shape of the fourteen centimetres wide astragalus, like that of a sandal and not convex on top as with the derived Neosauropoda, proves that Ohmdenosaurus is a basal sauropod. In 1990 John Stanton McIntosh classified Ohmdenosaurus in the Vulcanodontidae but that concept functioned as a waste-basket taxon for all kinds of unrelated basal sauropods, it has not been confirmed by an exact analysis as a vulcanodontid in the modern sense. Wild, R.. "Ein Sauropoden-Rest aus dem Posidonienschiefer von Holzmaden". Stuttgarter Beiträge zur Naturkunde, Serie B 41: 1-15
The Carboniferous is a geologic period and system that spans 60 million years from the end of the Devonian Period 358.9 million years ago, to the beginning of the Permian Period, 298.9 Mya. The name Carboniferous means "coal-bearing" and derives from the Latin words carbō and ferō, was coined by geologists William Conybeare and William Phillips in 1822. Based on a study of the British rock succession, it was the first of the modern'system' names to be employed, reflects the fact that many coal beds were formed globally during that time; the Carboniferous is treated in North America as two geological periods, the earlier Mississippian and the Pennsylvanian. Terrestrial animal life was well established by the Carboniferous period. Amphibians were the dominant land vertebrates, of which one branch would evolve into amniotes, the first terrestrial vertebrates. Arthropods were very common, many were much larger than those of today. Vast swaths of forest covered the land, which would be laid down and become the coal beds characteristic of the Carboniferous stratigraphy evident today.
The atmospheric content of oxygen reached its highest levels in geological history during the period, 35% compared with 21% today, allowing terrestrial invertebrates to evolve to great size. The half of the period experienced glaciations, low sea level, mountain building as the continents collided to form Pangaea. A minor marine and terrestrial extinction event, the Carboniferous rainforest collapse, occurred at the end of the period, caused by climate change. In the United States the Carboniferous is broken into Mississippian and Pennsylvanian subperiods; the Mississippian is about twice as long as the Pennsylvanian, but due to the large thickness of coal-bearing deposits with Pennsylvanian ages in Europe and North America, the two subperiods were long thought to have been more or less equal in duration. In Europe the Lower Carboniferous sub-system is known as the Dinantian, comprising the Tournaisian and Visean Series, dated at 362.5-332.9 Ma, the Upper Carboniferous sub-system is known as the Silesian, comprising the Namurian and Stephanian Series, dated at 332.9-298.9 Ma.
The Silesian is contemporaneous with the late Mississippian Serpukhovian plus the Pennsylvanian. In Britain the Dinantian is traditionally known as the Carboniferous Limestone, the Namurian as the Millstone Grit, the Westphalian as the Coal Measures and Pennant Sandstone; the International Commission on Stratigraphy faunal stages from youngest to oldest, together with some of their regional subdivisions, are: A global drop in sea level at the end of the Devonian reversed early in the Carboniferous. There was a drop in south polar temperatures; these conditions had little effect in the deep tropics, where lush swamps to become coal, flourished to within 30 degrees of the northernmost glaciers. Mid-Carboniferous, a drop in sea level precipitated a major marine extinction, one that hit crinoids and ammonites hard; this sea level drop and the associated unconformity in North America separate the Mississippian subperiod from the Pennsylvanian subperiod. This happened about 323 million years ago, at the onset of the Permo-Carboniferous Glaciation.
The Carboniferous was a time of active mountain-building as the supercontinent Pangaea came together. The southern continents remained tied together in the supercontinent Gondwana, which collided with North America–Europe along the present line of eastern North America; this continental collision resulted in the Hercynian orogeny in Europe, the Alleghenian orogeny in North America. In the same time frame, much of present eastern Eurasian plate welded itself to Europe along the line of the Ural Mountains. Most of the Mesozoic supercontinent of Pangea was now assembled, although North China, South China continents were still separated from Laurasia; the Late Carboniferous Pangaea was shaped like an "O." There were two major oceans in the Carboniferous—Panthalassa and Paleo-Tethys, inside the "O" in the Carboniferous Pangaea. Other minor oceans were shrinking and closed - Rheic Ocean, the small, shallow Ural Ocean and Proto-Tethys Ocean. Average global temperatures in the Early Carboniferous Period were high: 20 °C.
However, cooling during the Middle Carboniferous reduced average global temperatures to about 12 °C. Lack of growth rings of fossilized trees suggest a lack of seasons of a tropical climate. Glaciations in Gondwana, triggered by Gondwana's southward movement, continued into the Permian and because of the lack of clear markers and breaks, the deposits of this glacial period are referred to as Permo-Carboniferous in age; the cooling and drying of the climate led to the Carboniferous Rainforest Collapse during the late Carboniferous. Tropical rainforests fragmented and were devastated by climate change. Carboniferous rocks in Europe and eastern North America consist of a repeated sequence of limestone, sandstone and coal beds. In North America, the early Carboniferous is marine
In phylogenetics, an autapomorphy is a distinctive feature, known as a derived trait, unique to a given taxon. That is, it is found only in one taxon, but not found in any others or outgroup taxa, not those most related to the focal taxon, it can therefore be considered an apomorphy in relation to a single taxon. The word autapomorphy, first introduced in 1950 by German entomologist Willi Hennig, is derived from the Greek words αὐτός, aut- = "self"; because autapomorphies are only present in a single taxon, they do not convey information about relationship. Therefore, autapomorphies are not useful to infer phylogenetic relationships. However, like synapomorphy and plesiomorphy is a relative concept depending on the taxon in question. An autapomorphy at a given level may well be a synapomorphy at a less-inclusive level. An example of an autapomorphy can be described in modern snakes. Snakes have lost the two pairs of legs that characterize all of Tetrapoda, the closest taxa to Ophidia - as well as their common ancestors - all have two pairs of legs.
Therefore, the Ophidia taxon presents an autapomorphy with respect to its absence of legs. The autapomorphic species concept is one of many methods that scientists might use to define and distinguish species from one another; this definition assigns species on the basis of amount of divergence associated with reproductive incompatibility, measured by number of autapomorphies. This grouping method is referred to as the "monophyletic species concept" or the "phylospecies" concept and was popularized by D. E. Rosen in 1979. Within this definition, a species is seen as "the least inclusive monophyletic group definable by at least one autapomorphy". While this model of speciation is useful in that it avoids non-monophyletic groupings, it has its criticisms as well. N. I. Platnick, for example, believes the autapomorphic species concept to be inadequate because it allows for the possibility of reproductive isolation and speciation while revoking the "species" status of the mother population. In other words, if a peripheral population breaks away and becomes reproductively isolated, it would conceivably need to develop at least one autapomorphy to be recognized as a different species.
If this can happen without the larger mother population developing a new autapomorphy the mother population cannot remain a species under the autapomorphic species concept: it would no longer have any apomorphies not shared by the daughter species. Phylogenetic similarities: These phylogenetic terms are used to describe different patterns of ancestral and derived character or trait states as stated in the above diagram in association with synapomorphies. Homoplasy in biological systematics is when a trait has been gained or lost independently in separate lineages during evolution; this convergent evolution leads to species independently sharing a trait, different from the trait inferred to have been present in their common ancestor. Parallel Homoplasy – derived trait present in two groups or species without a common ancestor due to convergent evolution. Reverse Homoplasy – trait present in an ancestor but not in direct descendants that reappears in descendants. Apomorphy – a derived trait. Apomorphy shared by two or more taxa and inherited from a common ancestor is synapomorphy.
Apomorphy unique to a given taxon is autapomorphy. Synapomorphy/Homology – a derived trait, found in some or all terminal groups of a clade, inherited from a common ancestor, for which it was an autapomorphy. Underlying synapomorphy – a synapomorphy, lost again in many members of the clade. If lost in all but one, it can be hard to distinguish from an autapomorphy. Autapomorphy – a distinctive derived trait, unique to a given taxon or group. Symplesiomorphy – an ancestral trait shared by two or more taxa. Plesiomorphy – a symplesiomorphy discussed in reference to a more derived state. Pseudoplesiomorphy – is a trait that cannot be identified as neither a plesiomorphy nor an apomorphy, a reversal. Reversal – is a loss of derived trait present in ancestor and the reestablishment of a plesiomorphic trait. Convergence – independent evolution of a similar trait in two or more taxa. Hemiplasy