1.
Populus balsamifera
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The genus name Populus is from the Latin for poplar, and the specific epithet balsamifera from Latin for balsam-bearing. Other common names for the species include heartleaf balsam poplar, Populus balsamifera is the northernmost North American hardwood, growing transcontinentally on boreal and montane upland and flood plain sites, and attaining its best development on flood plains. It is a hardy, fast-growing tree which is short lived. The Balm-of-Gilead, also known as P. × gileadensis, is the hybrid between P. balsamifera and the eastern cottonwood, occurring occasionally where the two species ranges overlap. This hybrid is sometimes planted as a shade tree. The name Populus candicans has been used for either P. balsamifera or P. × jackii. The light, soft wood of Populus balsamifera is used for pulp, many kinds of animals use the twigs of Populus balsamifera for food. The leaves of the serve as food for caterpillars of various Lepidoptera. See List of Lepidoptera that feed on poplars, media related to Populus balsamifera at Wikimedia Commons
2.
Picea mariana
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Picea mariana is a North American species of spruce tree in the pine family. It is widespread across Canada, found in all 10 provinces and its range extends into northern parts of the United States, in Alaska, the Great Lakes region, and the upper Northeast. It is a frequent part of the known as taiga or boreal forest. Through much of its range it averages 5–15 m tall with a trunk 15–50 cm diameter at maturity, the bark is thin, scaly, and grayish brown. The leaves are needle-like, 6–15 mm long, stiff, four-sided, dark green on the upper sides. Natural hybridization occurs regularly with the closely related Picea rubens, numerous differences in details of its needle and pollen morphology also exist but require careful microscopic examination to detect. From true firs, such as Abies balsamea, it differs in having pendulous cones, persistent woody leaf-bases, older taxonomic synonyms include Abies mariana, Picea brevifolia, or Picea nigra. In swamp and muskeg it shows progressively slower growth rates from the edges toward the centre, the roots are shallow and wide spreading, resulting in susceptibility to windthrow. In the northern part of its range, ice pruned asymmetric black spruce are often seen with diminished foliage on the windward side, tilted trees colloquially called drunken trees are associated with thawing of permafrost. In the southern portion of its range it is primarily on wet organic soils. In the Great Lakes States it is most abundant in peat bogs and swamps, in these areas it is rare on uplands, except in isolated areas of northern Minnesota and the Upper Peninsula of Michigan. Most stands are even-aged due to frequent fire intervals in black spruce forests and it commonly grows in pure stands on organic soils and in mixed stands on mineral soils. It is tolerant of soils, and is commonly found on poorly drained acidic peatlands. The frequent fire return interval, a fire ecology, perpetuates numerous successional communities. Throughout boreal North America, Betula papyrifera and Populus tremuloides are successional hardwoods that frequently invade burns in black spruce, black spruce typically seeds in promptly after fire, and with the continued absence of fire, will eventually dominate the hardwoods. It is a pioneer that invades the sedge mat in filled-lake bogs, though often preceded slightly by Larix laricina, with which it may in time form a stable forest cover in swamps. The larvae of the spruce budworm moth cause defoliation which will lead to death if it occurs several years in a row, trees most at risk are those growing with balsam fir and white spruce. Numerous cultivars have been selected for use in parks and gardens, the cultivar P. mariana Nana is a dwarf form which has gained the Royal Horticultural Societys Award of Garden Merit
3.
Conservation status
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The conservation status of a group of organisms indicates whether the group still exists and how likely the group is to become extinct in the near future. Various systems of conservation status exist and are in use at international, multi-country, the IUCN Red List of Threatened Species is the best known worldwide conservation status listing and ranking system. Also included are species that have gone extinct since 500 AD, when discussing the IUCN Red List, the official term threatened is a grouping of three categories, critically endangered, endangered, and vulnerable. Widespread and abundant taxa are included in this category, Data deficient – Not enough data to make an assessment of its risk of extinction Not evaluated – Has not yet been evaluated against the criteria. The Convention on International Trade in Endangered Species of Wild Fauna and Flora aims to ensure that trade in specimens of wild animals. Many countries require CITES permits when importing plants and animals listed on CITES, in the European Union, the Birds and Habitats Directives are the legal instruments that evaluate the conservation status within the EU of species and habitats. NatureServe conservation status focuses on Latin America, United States, Canada, and it has been developed by scientists from NatureServe, The Nature Conservancy, and the network of natural heritage programs and data centers. It is increasingly integrated with the IUCN Red List system and its categories for species include, presumed extinct, possibly extinct, critically imperiled, imperiled, vulnerable, apparently secure, and secure. The system also allows ambiguous or uncertain ranks including inexact numeric ranks, NatureServe adds a qualifier for captive or cultivated only, which has a similar meaning to the IUCN Red List extinct in the wild status. The Red Data Book of the Russian Federation is used within the Russian Federation, in Australia, the Environment Protection and Biodiversity Conservation Act 1999 describes lists of threatened species, ecological communities and threatening processes. The categories resemble those of the 1994 IUCN Red List Categories & Criteria, prior to the EPBC Act, a simpler classification system was used by the Endangered Species Protection Act 1992. Some state and territory governments also have their own systems for conservation status, in Belgium, the Flemish Research Institute for Nature and Forest publishes an online set of more than 150 nature indicators in Dutch. In Canada, the Committee on the Status of Endangered Wildlife in Canada is a group of experts that assesses and designates which wild species are in danger of disappearing from Canada. Under the Species at Risk Act, it is up to the federal government, in China, the State, provinces and some counties have determined their key protected wildlife species. There is the China red data book, in Finland, a large number of species are protected under the Nature Conservation Act, and through the EU Habitats Directive and EU Birds Directive. In Germany, the Federal Agency for Nature Conservation publishes red lists of endangered species, india has the Wild Life Protection Act,1972, Amended 2003 and the Biological Diversity Act,2002. In Japan, the Ministry of Environment publishes a Threatened Wildlife of Japan Red Data Book, in the Netherlands, the Dutch Ministry of Agriculture, Nature and Food Quality publishes a list of threatened species, and conservation is enforced by the Nature Conservation Act 1998. Species are also protected through the Wild Birds and Habitats Directives, in New Zealand, the Department of Conservation publishes the New Zealand Threat Classification System lists
4.
Least Concern
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A least concern species is one which has been categorized by the International Union for Conservation of Nature as evaluated but not qualified for any other category. As such they do not qualify as threatened, near threatened, species cannot be assigned the Least Concern category unless they have had their population status evaluated. That is, adequate information is needed to make a direct, or indirect, since 2001 the category has had the abbreviation LC, following the IUCN2001 Categories & Criteria. However, around 20% of least concern taxa in the IUCN database use the code LR/lc, prior to 2001 least concern was a subcategory of the Lower Risk category and assigned the code LR/lc or. While least concern is not considered a red listed category by the IUCN, the number of animal species listed in this category totals 14033. There are also 101 animal subspecies listed and 1500 plant taxa, there are also two animal subpopulations listed, the Australasian and Southern African subpopulations of spiny dogfish. No fungi or protista have the classification, though only four species in those kingdoms have been evaluated by the IUCN, humans qualify for this category, and in 2008 were formally assessed as such by the IUCN
5.
IUCN Red List
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The IUCN Red List of Threatened Species, founded in 1964, is the worlds most comprehensive inventory of the global conservation status of biological species. The International Union for Conservation of Nature is the main authority on the conservation status of species. A series of Regional Red Lists are produced by countries or organizations, the IUCN Red List is set upon precise criteria to evaluate the extinction risk of thousands of species and subspecies. These criteria are relevant to all species and all regions of the world, the aim is to convey the urgency of conservation issues to the public and policy makers, as well as help the international community to try to reduce species extinction. Major species assessors include BirdLife International, the Institute of Zoology, the World Conservation Monitoring Centre, collectively, assessments by these organizations and groups account for nearly half the species on the Red List. The IUCN aims to have the category of every species re-evaluated every five years if possible, the 1964 IUCN Red List of Threatened Plants used the older pre-criteria Red List assessment system. Plants listed may not, therefore, appear in the current Red List, IUCN advise that is best to check both the online Red List and the 1997 plants Red List publication. The 2006 Red List, released on 4 May 2006 evaluated 40,168 species as a whole, plus an additional 2,160 subspecies, varieties, aquatic stocks, on 12 September 2007, the World Conservation Union released the 2007 IUCN Red List of Threatened Species. Russ Mittermeier, chief of Swiss-based IUCNs Primate Specialist Group, stated that 16,306 species are endangered with extinction,188 more than in 2006, the Red List includes the Sumatran orangutan in the Critically Endangered category and the Bornean orangutan in the Endangered category. The study shows at least 1,141 of the 5,487 mammals on Earth are known to be threatened with extinction, and 836 are listed as Data Deficient. The Red List of 2012 was released 19 July 2012 at Rio+20 Earth Summit, nearly 2,000 species were added, the IUCN assessed a total of 63,837 species which revealed 19,817 are threatened with extinction. With 3,947 described as endangered and 5,766 as endangered. At threat are 41% of amphibian species, 33% of reef-building corals, 30% of conifers, 25% of mammals, the IUCN Red List has listed 132 species of plants and animals from India as Critically Endangered. Extinct – No known individuals remaining, extinct in the wild – Known only to survive in captivity, or as a naturalized population outside its historic range. Critically endangered – Extremely high risk of extinction in the wild, Endangered – High risk of extinction in the wild. Vulnerable – High risk of endangerment in the wild, near threatened – Likely to become endangered in the near future. Does not qualify for a more at-risk category, widespread and abundant taxa are included in this category. Data deficient – Not enough data to make an assessment of its risk of extinction, Not evaluated – Has not yet been evaluated against the criteria
6.
Taxonomy (biology)
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Taxonomy is the science of defining groups of biological organisms on the basis of shared characteristics and giving names to those groups. The exact definition of taxonomy varies from source to source, but the core of the remains, the conception, naming. There is some disagreement as to whether biological nomenclature is considered a part of taxonomy, the broadest meaning of taxonomy is used here. The word taxonomy was introduced in 1813 by Candolle, in his Théorie élémentaire de la botanique, the term alpha taxonomy is primarily used today to refer to the discipline of finding, describing, and naming taxa, particularly species. In earlier literature, the term had a different meaning, referring to morphological taxonomy, ideals can, it may be said, never be completely realized. They have, however, a value of acting as permanent stimulants. Some of us please ourselves by thinking we are now groping in a beta taxonomy, turrill thus explicitly excludes from alpha taxonomy various areas of study that he includes within taxonomy as a whole, such as ecology, physiology, genetics, and cytology. He further excludes phylogenetic reconstruction from alpha taxonomy, thus, Ernst Mayr in 1968 defined beta taxonomy as the classification of ranks higher than species. This activity is what the term denotes, it is also referred to as beta taxonomy. How species should be defined in a group of organisms gives rise to practical and theoretical problems that are referred to as the species problem. The scientific work of deciding how to define species has been called microtaxonomy, by extension, macrotaxonomy is the study of groups at higher taxonomic ranks, from subgenus and above only, than species. While some descriptions of taxonomic history attempt to date taxonomy to ancient civilizations, earlier works were primarily descriptive, and focused on plants that were useful in agriculture or medicine. There are a number of stages in scientific thinking. Early taxonomy was based on criteria, the so-called artificial systems. Later came systems based on a complete consideration of the characteristics of taxa, referred to as natural systems, such as those of de Jussieu, de Candolle and Bentham. The publication of Charles Darwins Origin of Species led to new ways of thinking about classification based on evolutionary relationships and this was the concept of phyletic systems, from 1883 onwards. This approach was typified by those of Eichler and Engler, the advent of molecular genetics and statistical methodology allowed the creation of the modern era of phylogenetic systems based on cladistics, rather than morphology alone. Taxonomy has been called the worlds oldest profession, and naming and classifying our surroundings has likely been taking place as long as mankind has been able to communicate
7.
Plant
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Plants are mainly multicellular, predominantly photosynthetic eukaryotes of the kingdom Plantae. The term is generally limited to the green plants, which form an unranked clade Viridiplantae. This includes the plants, conifers and other gymnosperms, ferns, clubmosses, hornworts, liverworts, mosses and the green algae. Green plants have cell walls containing cellulose and obtain most of their energy from sunlight via photosynthesis by primary chloroplasts and their chloroplasts contain chlorophylls a and b, which gives them their green color. Some plants are parasitic and have lost the ability to produce amounts of chlorophyll or to photosynthesize. Plants are characterized by sexual reproduction and alternation of generations, although reproduction is also common. There are about 300–315 thousand species of plants, of which the great majority, green plants provide most of the worlds molecular oxygen and are the basis of most of Earths ecologies, especially on land. Plants that produce grains, fruits and vegetables form humankinds basic foodstuffs, Plants play many roles in culture. They are used as ornaments and, until recently and in variety, they have served as the source of most medicines. The scientific study of plants is known as botany, a branch of biology, Plants are one of the two groups into which all living things were traditionally divided, the other is animals. The division goes back at least as far as Aristotle, who distinguished between plants, which generally do not move, and animals, which often are mobile to catch their food. Much later, when Linnaeus created the basis of the system of scientific classification. Since then, it has become clear that the plant kingdom as originally defined included several unrelated groups, however, these organisms are still often considered plants, particularly in popular contexts. When the name Plantae or plant is applied to a group of organisms or taxon. The evolutionary history of plants is not yet settled. Those which have been called plants are in bold, the way in which the groups of green algae are combined and named varies considerably between authors. Algae comprise several different groups of organisms which produce energy through photosynthesis, most conspicuous among the algae are the seaweeds, multicellular algae that may roughly resemble land plants, but are classified among the brown, red and green algae. Each of these groups also includes various microscopic and single-celled organisms
8.
Pinophyta
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The Pinophyta, also known as Coniferophyta or Coniferae, or commonly as conifers, are a division of vascular land plants containing a single class, Pinopsida. They are gymnosperms, cone-bearing seed plants, all extant conifers are perennial woody plants with secondary growth. The great majority are trees, though a few are shrubs, examples include cedars, Douglas firs, cypresses, firs, junipers, kauri, larches, pines, hemlocks, redwoods, spruces, and yews. As of 1998, the division Pinophyta was estimated to contain eight families,68 genera, although the total number of species is relatively small, conifers are ecologically important. They are the dominant plants over large areas of land, most notably the taiga of the Northern Hemisphere, boreal conifers have many wintertime adaptations. The narrow conical shape of northern conifers, and their downward-drooping limbs, many of them seasonally alter their biochemistry to make them more resistant to freezing. While tropical rainforests have more biodiversity and turnover, the conifer forests of the world represent the largest terrestrial carbon sink. Conifers are of economic value for softwood lumber and paper production. The earliest conifers in the record date to the late Carboniferous period, possibly arising from Cordaites. Pinophytes, Cycadophytes, and Ginkgophytes all developed at this time, an important adaptation of these gymnosperms was allowing plants to live without being so dependent on water. Other adaptations are pollen and the seed, which allows the embryo to be transported and developed elsewhere, Conifers appear to be one of the taxa that benefited from the Permian–Triassic extinction event, and were the dominant land plants of the Mesozoic. They were overtaken by the plants, which first appeared in the Cretaceous. They were the food of herbivorous dinosaurs, and their resins and poisons would have given protection against herbivores. Reproductive features of modern conifers had evolved by the end of the Mesozoic era, Conifer is a Latin word, a compound of conus and ferre, meaning the one that bears cone. A descriptive name in use for the conifers is Coniferae. Alternatively, descriptive botanical names may also be used at any rank above family and this means that if conifers are considered a division, they may be called Pinophyta or Coniferae. As a class they may be called Pinopsida or Coniferae, as an order they may be called Pinales or Coniferae or Coniferales. Conifers are the largest and economically most important component group of the gymnosperms, the division Pinophyta consists of just one class, Pinopsida, which includes both living and fossil taxa
9.
Pinales
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The order Pinales in the division Pinophyta, class Pinopsida, comprises all the extant conifers. This order used to be known as the Coniferales, the distinguishing characteristic is the reproductive structure known as a cone produced by all Pinales. All of the extant conifers, such as cedar, celery-pine, cypress, fir, juniper, larch, pine, redwood, spruce, some fossil conifers, however, belong to other distinct orders within the division Pinophyta. The yews had been separated into an order of their own. The families included are the Araucariaceae, Cephalotaxaceae, Cupressaceae, Pinaceae, Podocarpaceae, Sciadopityaceae, and Taxaceae
10.
Pinaceae
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The Pinaceae are trees or shrubs, including many of the well-known conifers of commercial importance such as cedars, firs, hemlocks, larches, pines and spruces. The family is included in the order Pinales, formerly known as Coniferales, Pinaceae are supported as monophyletic by their protein-type sieve cell plastids, pattern of proembryogeny, and lack of bioflavonoids. The family often forms the dominant component of boreal, coastal, one species, Pinus merkusii, grows just south of the equator in Southeast Asia. Major centres of diversity are found in the mountains of southwest China, Mexico, central Japan, and California. Members of the family Pinaceae are trees growing from 2 to 100 m tall, mostly evergreen, resinous, monoecious, with subopposite or whorled branches, the embryos of Pinaceae have three to 24 cotyledons. The female cones are large and usually woody, 2–60 cm long, with spirally arranged scales. The male cones are small,0. 5–6.0 cm long, seed dispersal is mostly by wind, but some species have large seeds with reduced wings, and are dispersed by birds. Analysis of Pinaceae cones reveals how selective pressure has shaped the evolution of variable cone size, variation in cone size in the family has likely resulted from the variation of seed dispersal mechanisms available in their environments over time. All Pinaceae with seeds weighing less than 90 mg are seemingly adapted for wind dispersal, pines having seeds larger than 100 mg are more likely to have benefited from adaptations that promote animal dispersal, particularly by birds. Pinaceae that persist in areas where tree squirrels are abundant do not seem to have evolved adaptations for bird dispersal, boreal conifers have many adaptions for winter. Classification of the subfamilies and genera of Pinaceae has been subject to debate in the past, Pinaceae ecology, morphology, and history have all been used as the basis for methods of analyses of the family. An 1891 publication divided the family into two subfamiles, using the number and position of resin canals in the primary region of the young taproot as the primary consideration. In a 1910 publication, the family was divided into two based on the occurrence and type of long–short shoot dimorphism. A more recent classification divided the subfamilies and genera based on the consideration of features of ovulate cone anatomy among extant, below is an example of how the morphology has been used to classify Pinaceae. Sieve element plastids of Gymnospermae, their ultrastructure and relation to systematics, D. F. Greene & E. A. Johnson. The dispersal of winged fruits and seeds differing in autorotative behavior, a. Liston, D. S. Gernandt, T. F. Vining, C. S. Campbell & D. Piñero. Molecular phylogeny of Pinaceae and Pinus, proceedings of the Fourth International Conifer Conference. Brugge, International Society for Horticultural Science, cS1 maint, Multiple names, authors list Zsolt Debreczy, Istvan Racz
11.
Larch
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Larches are conifers in the genus Larix, in the family Pinaceae. Growing from 20 to 45 m tall, they are native to much of the temperate northern hemisphere, on lowlands in the north. Larches are among the dominant plants in the forests of Siberia. Although they are conifers, larches are deciduous trees lose their needles in the autumn. The leaves are needle-like, 2–5 centimetres long, slender, larches are among the few deciduous conifers, which are mostly evergreen. There are eleven accepted species of larch, Larix czekanowskii Szafer Larix decidua Mill, plains of central and eastern Siberia. Mountains of eastern Himalaya Larix kaempferi Carr, Larix laricina K. Koch Tamarack larch or American larch. Parts of Alaska and throughout Canada and the northern United States from the eastern Rocky Mountains to the Atlantic shore, Mountains of northwest United States and southwest Canada, at very high altitude. Larix mastersiana Rehder & E. H. Wilson Masters larch, Mountains of northwest United States and southwest Canada, at lower altitudes. Most if not all of the species can be hybridised in cultivation, currently-accepted hybrids are, Larix × lubarskii Sukaczev Larix × maritima Sukaczev Larix × polonica Racib. Larch is used as a plant by the larvae of a number of Lepidoptera species—see list of Lepidoptera that feed on larches. In Canada, this disease was first detected in 1980 and is harmful to an indigenous species larch. Larches are also vulnerable to Phytophthora ramorum, in late 2009 the disease was first found in Japanese larch trees in the English counties of Devon, Cornwall and Somerset, and has since spread to the south-west of Scotland. In August 2010 the disease was found in Japanese larch trees in counties Waterford and Tipperary in Ireland, larch wood is valued for its tough, waterproof and durable qualities. Top quality knot-free timber is in demand for building yachts and other small boats, for exterior cladding of buildings. The timber is resistant to rot when in contact with the ground, the hybrid Dunkeld larch is widely grown as a timber crop in Northern Europe, valued for its fast growth and disease resistance. Larch has also used in herbal medicine, see Bach flower remedies. In Central Europe larch is viewed as one of the best wood materials for the building of residences, planted on borders with birch, both tree species were used in pagan cremations
12.
Binomial nomenclature
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Such a name is called a binomial name, a binomen, binominal name or a scientific name, more informally it is also called a Latin name. The first part of the name identifies the genus to which the species belongs, for example, humans belong to the genus Homo and within this genus to the species Homo sapiens. The formal introduction of system of naming species is credited to Carl Linnaeus. But Gaspard Bauhin, in as early as 1623, had introduced in his book Pinax theatri botanici many names of genera that were adopted by Linnaeus. Although the general principles underlying binomial nomenclature are common to these two codes, there are differences, both in the terminology they use and in their precise rules. Similarly, both parts are italicized when a binomial name occurs in normal text, thus the binomial name of the annual phlox is now written as Phlox drummondii. In scientific works, the authority for a name is usually given, at least when it is first mentioned. In zoology Patella vulgata Linnaeus,1758, the original name given by Linnaeus was Fringilla domestica, the parentheses indicate that the species is now considered to belong in a different genus. The ICZN does not require that the name of the person who changed the genus be given, nor the date on which the change was made, in botany Amaranthus retroflexus L. – L. is the standard abbreviation used in botany for Linnaeus. – Linnaeus first named this bluebell species Scilla italica, Rothmaler transferred it to the genus Hyacinthoides, the ICN does not require that the dates of either publication be specified. Prior to the adoption of the binomial system of naming species. Together they formed a system of polynomial nomenclature and these names had two separate functions. First, to designate or label the species, and second, to be a diagnosis or description, such polynomial names may sometimes look like binomials, but are significantly different. For example, Gerards herbal describes various kinds of spiderwort, The first is called Phalangium ramosum, Branched Spiderwort, is aptly termed Phalangium Ephemerum Virginianum, Soon-Fading Spiderwort of Virginia. The Latin phrases are short descriptions, rather than identifying labels, the Bauhins, in particular Caspar Bauhin, took some important steps towards the binomial system, by pruning the Latin descriptions, in many cases to two words. The adoption by biologists of a system of binomial nomenclature is due to Swedish botanist and physician Carl von Linné. It was in his 1753 Species Plantarum that he first began using a one-word trivial name together with a generic name in a system of binomial nomenclature. This trivial name is what is now known as an epithet or specific name
13.
Synonym (taxonomy)
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For example, Linnaeus was the first to give a scientific name to the Norway spruce, which he called Pinus abies. This name is no longer in use, it is now a synonym of the current scientific name which is Picea abies, unlike synonyms in other contexts, in taxonomy a synonym is not interchangeable with the name of which it is a synonym. In taxonomy, synonyms are not equals, but have a different status, for any taxon with a particular circumscription, position, and rank, only one scientific name is considered to be the correct one at any given time. A synonym cannot exist in isolation, it is always an alternative to a different scientific name, given that the correct name of a taxon depends on the taxonomic viewpoint used a name that is one taxonomists synonym may be another taxonomists correct name. Synonyms may arise whenever the same taxon is described and named more than once, independently. They may also arise when existing taxa are changed, as when two taxa are joined to one, a species is moved to a different genus. To the general user of scientific names, in such as agriculture, horticulture, ecology, general science. A synonym is a name that was used as the correct scientific name but which has been displaced by another scientific name. Thus Oxford Dictionaries Online defines the term as a name which has the same application as another. In handbooks and general texts, it is useful to have mentioned as such after the current scientific name. Synonyms used in this way may not always meet the strict definitions of the synonym in the formal rules of nomenclature which govern scientific names. Changes of scientific name have two causes, they may be taxonomic or nomenclatural, a name change may be caused by changes in the circumscription, position or rank of a taxon, representing a change in taxonomic, scientific insight. A name change may be due to purely nomenclatural reasons, that is, based on the rules of nomenclature, the earliest such name is called the senior synonym, while the later name is the junior synonym. One basic principle of zoological nomenclature is that the earliest correctly published name, synonyms are important because if the earliest name cannot be used, then the next available junior synonym must be used for the taxon. Objective synonyms refer to taxa with the type and same rank. For example, John Edward Gray published the name Antilocapra anteflexa in 1855 for a species of pronghorn, however, it is now commonly accepted that his specimen was an unusual individual of the species Antilocapra americana published by George Ord in 1815. Ords name thus takes precedence, with Antilocapra anteflexa being a subjective synonym. Objective synonyms are common at the level of genera, because for various reasons two genera may contain the type species, these are objective synonyms
14.
Canada
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Canada is a country in the northern half of North America. Canadas border with the United States is the worlds longest binational land border, the majority of the country has a cold or severely cold winter climate, but southerly areas are warm in summer. Canada is sparsely populated, the majority of its territory being dominated by forest and tundra. It is highly urbanized with 82 per cent of the 35.15 million people concentrated in large and medium-sized cities, One third of the population lives in the three largest cities, Toronto, Montreal and Vancouver. Its capital is Ottawa, and other urban areas include Calgary, Edmonton, Quebec City, Winnipeg. Various aboriginal peoples had inhabited what is now Canada for thousands of years prior to European colonization. Pursuant to the British North America Act, on July 1,1867, the colonies of Canada, New Brunswick and this began an accretion of provinces and territories to the mostly self-governing Dominion to the present ten provinces and three territories forming modern Canada. With the Constitution Act 1982, Canada took over authority, removing the last remaining ties of legal dependence on the Parliament of the United Kingdom. Canada is a parliamentary democracy and a constitutional monarchy, with Queen Elizabeth II being the head of state. The country is officially bilingual at the federal level and it is one of the worlds most ethnically diverse and multicultural nations, the product of large-scale immigration from many other countries. Its advanced economy is the eleventh largest in the world, relying chiefly upon its abundant natural resources, Canadas long and complex relationship with the United States has had a significant impact on its economy and culture. Canada is a country and has the tenth highest nominal per capita income globally as well as the ninth highest ranking in the Human Development Index. It ranks among the highest in international measurements of government transparency, civil liberties, quality of life, economic freedom, Canada is an influential nation in the world, primarily due to its inclusive values, years of prosperity and stability, stable economy, and efficient military. While a variety of theories have been postulated for the origins of Canada. In 1535, indigenous inhabitants of the present-day Quebec City region used the word to direct French explorer Jacques Cartier to the village of Stadacona, from the 16th to the early 18th century Canada referred to the part of New France that lay along the St. Lawrence River. In 1791, the area became two British colonies called Upper Canada and Lower Canada collectively named The Canadas, until their union as the British Province of Canada in 1841. Upon Confederation in 1867, Canada was adopted as the name for the new country at the London Conference. The transition away from the use of Dominion was formally reflected in 1982 with the passage of the Canada Act, later that year, the name of national holiday was changed from Dominion Day to Canada Day
15.
Yukon
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Yukon is the smallest and westernmost of Canadas three federal territories. The territory has the smallest population of any province or territory in Canada, Whitehorse is the territorial capital and Yukons only city. The territory was split from the Northwest Territories in 1898 and was named the Yukon Territory, though officially bilingual, the Yukon Government also recognizes First Nations languages. At 5,959 m, Yukons Mount Logan, in Kluane National Park and Reserve, is the highest mountain in Canada, most of Yukon has a subarctic climate, characterized by long cold winters and brief warm summers. The Arctic Ocean coast has a tundra climate, notable rivers include the Yukon River, after which the territory was named, as well as the Pelly, Stewart, Peel, White and Tatshenshini rivers. Long before the arrival of Europeans, central and southern Yukon was populated by First Nations people, sites of archeological significance in Yukon hold some of the earliest evidence of the presence of human occupation in North America. The sites safeguard the history of the first people and the earliest First Nations of the Yukon, the volcanic eruption of Mount Churchill in approximately 800 AD in what is now the U. S. Coastal and inland First Nations had extensive trading networks, European incursions into the area only began early in the 19th century with the fur trade, followed by missionaries. By the 1870s and 1880s gold miners began to arrive and this drove a population increase that justified the establishment of a police force, just in time for the start of the Klondike Gold Rush in 1897. The increased population coming with the gold led to the separation of the Yukon district from the Northwest Territories. Its northern coast is on the Beaufort Sea and its ragged eastern boundary mostly follows the divide between the Yukon Basin and the Mackenzie River drainage basin to the east in the Mackenzie mountains. Most of the territory is in the watershed of its namesake, the southern Yukon is dotted with a large number of large, long and narrow glacier-fed alpine lakes, most of which flow into the Yukon River system. The larger lakes include Teslin Lake, Atlin Lake, Tagish Lake, Marsh Lake, Lake Laberge, Kusawa Lake, bennett Lake on the Klondike Gold Rush trail is a lake flowing into Nares Lake, with the greater part of its area within Yukon. Canadas highest point, Mount Logan, is in the territorys southwest, Mount Logan and a large part of the Yukons southwest are in Kluane National Park and Reserve, a UNESCO World Heritage Site. Other national parks include Ivvavik National Park and Vuntut National Park in the north, other watersheds include the Mackenzie River, the Peel Watershed and the Alsek–Tatshenshini, and a number of rivers flowing directly into the Beaufort Sea. The two main Yukon rivers flowing into the Mackenzie in the Northwest Territories are the Liard River in the southeast, notable widespread tree species within Yukon are the black spruce and white spruce. Many trees are stunted because of the growing season and severe climate. The capital, Whitehorse, is also the largest city, with about three-quarters of the population, the second largest is Dawson City, which was the capital until 1952
16.
Inuvik
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Inuvik /ᵻˈnuːvɪk/ is a town in the Northwest Territories of Canada and is the administrative centre for the Inuvik Region. The population as of the 2011 Census was 3,484, the two previous census counts show wide fluctuations due to economic conditions,2,894 in 2001 and 3,296 in 1996. In 2012 the Government of the Northwest Territories reported that the population was 3,321 with a yearly growth rate of 0. 2% from 2001. Inuvik was conceived in 1953 as a replacement administrative centre for the hamlet of Aklavik on the west of the Mackenzie Delta, initially called New Aklavik, it was renamed Inuvik in 1958. The school was built in 1959 and the hospital, government offices and staff residences in 1960, naval Radio Station Inuvik, callsign CFV, was commissioned on 10 September 1963 after operations had been successfully transferred from NRS Aklavik. Station CFV was part of the SUPRAD network of intercept and direction finding stations, CFS Inuvik closed on 1 April 1986 and the site was transferred to the Dept. of Transport for use as a telecommunications station. Nothing remains of CFS Inuvik today, even now, many people of all backgrounds still lament its closing. The Navy Operations base at the end of Navy Road was completely dismantled and removed, Inuvik achieved village status in 1967 and became a full town in 1979 with an elected mayor and council. This all collapsed in 1990 for a variety of reasons, including disappearing government subsidies, local resistance to petroleum exploration, on 10 February 2010, the Google Street View imaging service uploaded images of most Inuvik streets. Local CBC Radio broadcasts an hour of programming a day in each of these languages, there are also about 100 Muslims, most of whom came there for economic opportunities. A small mosque was established in 2010, Inuvik is located on the East Channel of the Mackenzie Delta, approximately 100 km from the Arctic Ocean and approximately 200 km north of the Arctic Circle. Due to its location, Inuvik experiences an average of 56 days of continuous sunlight every summer and 30 days of polar night every winter. Access is via the Dempster Highway for the majority of the year, the highway relies on ferries and ice bridges to get across the rivers. It is thus closed during the time of freeze-up, for ice to form and allow ice bridges, at these times, there is air access only. The main airport is Inuvik Airport and there is a general aviation airport, when the Mackenzie River is ice-free, Northern Transportation Company Limited provides a commercial barge service from Hay River, on Great Slave Lake to the regional terminal in Inuvik. The annual sealift moves supplies as far east as Taloyoak, Nunavut and west to Barrow, a distinct feature of Inuvik is the use of utilidors – above-ground utility conduits carrying water and sewage – which are covered by corrugated steel. They run throughout town connecting most buildings, and as a result there are small bridges and underpasses. The utilidors are necessary because of the permafrost underlying the town, another feature is an inukhuk placed outside the Mackenzie Hotel, which was rebuilt in 2006
17.
Northwest Territories
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The Northwest Territories is a territory of Canada. At a land area of approximately 1,144,000 km2 and its estimated population as of 2016 is 44,291. Yellowknife became the capital in 1967, following recommendations by the Carrothers Commission. The Northwest Territories are bordered by Canadas two other territories, Nunavut to the east and Yukon to the west, and by the provinces of British Columbia, Alberta, and Saskatchewan to the south. The name is descriptive, adopted by the British government during the era to indicate where it lay in relation to Ruperts Land. It is shortened from North-Western Territory, in Inuktitut, the Northwest Territories are referred to as ᓄᓇᑦᓯᐊᖅ, beautiful land. There was some discussion of changing the name of the Northwest Territories after the splitting off of Nunavut, one proposal was Denendeh, as advocated by the former premier Stephen Kakfwi, among others. One of the most popular proposals for a new name – one to name the territory Bob – began as a prank, in the end a poll conducted prior to division showed that strong support remained to keep the name Northwest Territories. This name arguably became more appropriate following division than it had been when the territories extended far into Canadas north-central and it possibly meets Manitoba at a quadripoint to the extreme southeast, though surveys have not been completed. It has an area of 1,183,085 km2. Territorial islands in the Canadian Arctic Archipelago include Banks Island, Borden Island, Prince Patrick Island and its highest point is Mount Nirvana near the border with Yukon at an elevation of 2,773 m. The Northwest Territories extends for more than 1,300,000 km2 and has a large climate variant from south to north, the southern part of the territory has a subarctic climate, while the islands and northern coast have a polar climate. Summers in the north are short and cool, with highs in the mid teens Celsius. Winters are long and harsh, daytime highs in the mid −20 °C, extremes are common with summer highs in the south reaching 36 °C and lows reaching into the negatives. In winter in the south, it is not uncommon for the temperatures to reach −40 °C, in the north, temperatures can reach highs of 30 °C, and lows can reach into the low negatives. In winter in the north it is not uncommon for the temperatures to reach −50 °C, thunderstorms are not rare in the south. In the north they are rare, but do occur. Tornadoes are extremely rare but have happened with the most notable one happening just outside Yellowknife that destroyed a communications tower, the Territory has a fairly dry climate due to the mountains in the west
18.
Newfoundland (island)
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Newfoundland is a large Canadian island off the east coast of the North American mainland, and the most populous part of the Canadian province of Newfoundland and Labrador. It has 29 percent of the land area. The island is separated from the Labrador Peninsula by the Strait of Belle Isle and it blocks the mouth of the Saint Lawrence River, creating the Gulf of Saint Lawrence, the worlds largest estuary. Newfoundlands nearest neighbour is the French overseas community of Saint-Pierre and Miquelon, with an area of 108,860 square kilometres, Newfoundland is the worlds 16th-largest island, Canadas fourth-largest island, and the largest Canadian island outside the North. The provincial capital, St. Johns, is located on the southeastern coast of the island, Cape Spear, just south of the capital, is the easternmost point of North America, excluding Greenland. It is common to consider all directly neighbouring islands such as New World, Twillingate, Fogo, by that classification, Newfoundland and its associated small islands have a total area of 111,390 square kilometres. Additionally 6. 1% claimed at least one parent of French ancestry, the islands total population as of the 2006 census was 479,105. Long settled by peoples of the Dorset culture, the island was visited by the Icelandic Viking Leif Eriksson in the 11th century. The next European visitors to Newfoundland were Portuguese, Basque, Spanish, French, the island was visited by the Genoese navigator John Cabot, working under contract to King Henry VII of England on his expedition from Bristol in 1497. In 1501, Portuguese explorers Gaspar Corte-Real and his brother Miguel Corte-Real charted part of the coast of Newfoundland in a attempt to find the Northwest Passage. Newfoundland is considered Britains oldest colony, at the time of English settlement, the Beothuk inhabited the island. While there is evidence of ancient indigenous peoples on the island. LAnse aux Meadows was a Norse settlement near the northernmost tip of Newfoundland, the site is considered the only undisputed evidence of Pre-Columbian contact between the Old and New Worlds, if the Norse-Inuit contact on Greenland is not counted. There is a second suspected Norse site in Point Rosee, the island is a likely location of Vinland, mentioned in the Viking Chronicles, although this has been disputed. The indigenous people on the island at the time of European settlement were the Beothuk, later immigrants developed a variety of dialects associated with settlement on the island, Newfoundland English, Newfoundland French. In the 19th century, it also had a dialect of Irish known as Newfoundland Irish, Scottish Gaelic was spoken on the island during the 19th and early 20th centuries, particularly in the Codroy Valley area, chiefly by settlers from Cape Breton Island, Nova Scotia. The Gaelic names reflected the association with fishing, in Scottish Gaelic, it was called Eilean a Trosg, or literally, similarly, the Irish Gaelic name Talamh an Éisc means Land of the Fish. The first inhabitants of Newfoundland were the Paleo-Eskimo, who have no link to other groups in Newfoundland history
19.
United States
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Forty-eight of the fifty states and the federal district are contiguous and located in North America between Canada and Mexico. The state of Alaska is in the northwest corner of North America, bordered by Canada to the east, the state of Hawaii is an archipelago in the mid-Pacific Ocean. The U. S. territories are scattered about the Pacific Ocean, the geography, climate and wildlife of the country are extremely diverse. At 3.8 million square miles and with over 324 million people, the United States is the worlds third- or fourth-largest country by area, third-largest by land area. It is one of the worlds most ethnically diverse and multicultural nations, paleo-Indians migrated from Asia to the North American mainland at least 15,000 years ago. European colonization began in the 16th century, the United States emerged from 13 British colonies along the East Coast. Numerous disputes between Great Britain and the following the Seven Years War led to the American Revolution. On July 4,1776, during the course of the American Revolutionary War, the war ended in 1783 with recognition of the independence of the United States by Great Britain, representing the first successful war of independence against a European power. The current constitution was adopted in 1788, after the Articles of Confederation, the first ten amendments, collectively named the Bill of Rights, were ratified in 1791 and designed to guarantee many fundamental civil liberties. During the second half of the 19th century, the American Civil War led to the end of slavery in the country. By the end of century, the United States extended into the Pacific Ocean. The Spanish–American War and World War I confirmed the status as a global military power. The end of the Cold War and the dissolution of the Soviet Union in 1991 left the United States as the sole superpower. The U. S. is a member of the United Nations, World Bank, International Monetary Fund, Organization of American States. The United States is a developed country, with the worlds largest economy by nominal GDP. It ranks highly in several measures of performance, including average wage, human development, per capita GDP. While the U. S. economy is considered post-industrial, characterized by the dominance of services and knowledge economy, the United States is a prominent political and cultural force internationally, and a leader in scientific research and technological innovations. In 1507, the German cartographer Martin Waldseemüller produced a map on which he named the lands of the Western Hemisphere America after the Italian explorer and cartographer Amerigo Vespucci
20.
Minnesota
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Minnesota is a state in the midwestern and northern regions of the United States. Minnesota was admitted as the 32nd U. S. state on May 11,1858, the state has a large number of lakes, and is known by the slogan Land of 10,000 Lakes. Its official motto is LÉtoile du Nord, Minnesota is the 12th largest in area and the 21st most populous of the U. S. Minnesota is known for its progressive political orientation and its high rate of civic participation and voter turnout. Until European settlement, Minnesota was inhabited by the Dakota and Ojibwe/Anishinaabe, in recent decades, immigration from Asia, the Horn of Africa, and Latin America has broadened its historic demographic and cultural composition. Minnesotas standard of living index is among the highest in the United States, Native Americans demonstrated the name to early settlers by dropping milk into water and calling it mnisota. Many places in the state have similar names, such as Minnehaha Falls, Minneiska, Minneota, Minnetonka, Minnetrista, and Minneapolis, a combination of mni and polis, Minnesota is the second northernmost U. S. state. Its isolated Northwest Angle in Lake of the Woods county is the part of the 48 contiguous states lying north of the 49th parallel. The state is part of the U. S. region known as the Upper Midwest and it shares a Lake Superior water border with Michigan and a land and water border with Wisconsin to the east. Iowa is to the south, North Dakota and South Dakota are to the west, with 86,943 square miles, or approximately 2.25 percent of the United States, Minnesota is the 12th-largest state. Minnesota has some of the Earths oldest rocks, gneisses that are about 3.6 billion years old. About 2.7 billion years ago, basaltic lava poured out of cracks in the floor of the primordial ocean, the roots of these volcanic mountains and the action of Precambrian seas formed the Iron Range of northern Minnesota. Following a period of volcanism 1, in more recent times, massive ice sheets at least one kilometer thick ravaged the landscape of the state and sculpted its terrain. The Wisconsin glaciation left 12,000 years ago and these glaciers covered all of Minnesota except the far southeast, an area characterized by steep hills and streams that cut into the bedrock. This area is known as the Driftless Zone for its absence of glacial drift, much of the remainder of the state outside the northeast has 50 feet or more of glacial till left behind as the last glaciers retreated. Gigantic Lake Agassiz formed in the northwest 13,000 years ago and its bed created the fertile Red River valley, and its outflow, glacial River Warren, carved the valley of the Minnesota River and the Upper Mississippi downstream from Fort Snelling. Minnesota is geologically quiet today, it experiences earthquakes infrequently, the states high point is Eagle Mountain at 2,301 feet, which is only 13 miles away from the low of 601 feet at the shore of Lake Superior. Notwithstanding dramatic local differences in elevation, much of the state is a rolling peneplain. Two major drainage divides meet in Minnesotas northeast in rural Hibbing, forming a triple watershed, precipitation can follow the Mississippi River south to the Gulf of Mexico, the Saint Lawrence Seaway east to the Atlantic Ocean, or the Hudson Bay watershed to the Arctic Ocean
21.
Cranesville Swamp Preserve
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Cranesville Swamp Preserve is a 1, 600-acre preserve situated in Preston County, West Virginia and Garrett County, Maryland. It is one of the few remaining boreal bogs in the southern United States, unusual in harboring many plants, the Nature Conservancy purchased the 1,600 acres beginning in 1960. In October 1964, the site was designated as one of the first National Natural Landmarks in the country, cranesville Swamp is situated in a natural bowl, or frost pocket, creating a climate that is more consistent with more northerly regions. Cranesville Swamps unusual setting allows 19 different plant communities to flourish, with the most dominant species being, among others, sphagnum moss, speckled alder, bog species include round-leaved sundew and narrowleaf gentian. The swamp is home to the southernmost natural community of larches. In addition to bear, porcupine, snowshoe hare and a wide variety of birds. Birders have also noted the golden-crowned kinglet and Nashville warbler, rarely seen in this area
22.
Maryland
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The states largest city is Baltimore, and its capital is Annapolis. Among its occasional nicknames are Old Line State, the Free State, the state is named after Henrietta Maria of France, the wife of Charles I of England. George Calvert was the first Lord of Baltimore and the first English proprietor of the colonial grant. Maryland was the state to ratify the United States Constitution. Maryland is one of the smallest U. S. states in terms of area, as well as one of the most densely populated, Maryland has an area of 12,406.68 square miles and is comparable in overall area with Belgium. It is the 42nd largest and 9th smallest state and is closest in size to the state of Hawaii, the next largest state, its neighbor West Virginia, is almost twice the size of Maryland. Maryland possesses a variety of topography within its borders, contributing to its nickname America in Miniature. The mid-portion of this border is interrupted by Washington, D. C. which sits on land that was part of Montgomery and Prince Georges counties and including the town of Georgetown. This land was ceded to the United States Federal Government in 1790 to form the District of Columbia, the Chesapeake Bay nearly bisects the state and the counties east of the bay are known collectively as the Eastern Shore. Close to the town of Hancock, in western Maryland, about two-thirds of the way across the state. This geographical curiosity makes Maryland the narrowest state, bordered by the Mason–Dixon line to the north, portions of Maryland are included in various official and unofficial geographic regions. Much of the Baltimore–Washington corridor lies just south of the Piedmont in the Coastal Plain, earthquakes in Maryland are infrequent and small due to the states distance from seismic/earthquake zones. The M5.8 Virginia earthquake in 2011 was felt moderately throughout Maryland, buildings in the state are not well-designed for earthquakes and can suffer damage easily. The lack of any glacial history accounts for the scarcity of Marylands natural lakes, laurel Oxbow Lake is an over one-hundred-year-old 55-acre natural lake two miles north of Maryland City and adjacent to Russett. Chews Lake is a natural lake two miles south-southeast of Upper Marlboro. There are numerous lakes, the largest of them being the Deep Creek Lake. Maryland has shale formations containing natural gas, where fracking is theoretically possible, as is typical of states on the East Coast, Marylands plant life is abundant and healthy. Middle Atlantic coastal forests, typical of the southeastern Atlantic coastal plain, grow around Chesapeake Bay, moving west, a mixture of Northeastern coastal forests and Southeastern mixed forests cover the central part of the state
23.
Alaska
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Alaska is a U. S. state located in the northwest extremity of North America. To the north are the Chukchi and Beaufort seas–the southern parts of the Arctic Ocean, the Pacific Ocean lies to the south and southwest. Alaska is the largest state in the United States by area, the 3rd least populous, approximately half of Alaskas residents live within the Anchorage metropolitan area. Alaskas economy is dominated by the fishing, natural gas, military bases and tourism are also a significant part of the economy. The United States purchased Alaska from the Russian Empire on March 30,1867, the area went through several administrative changes before becoming organized as a territory on May 11,1912. It was admitted as the 49th state of the U. S. on January 3,1959, the name Alaska was introduced in the Russian colonial period when it was used to refer to the peninsula. It was derived from an Aleut, or Unangam idiom, which refers to the mainland of Alaska. Literally, it means object to which the action of the sea is directed, Alaska is the northernmost and westernmost state in the United States and has the most easterly longitude in the United States because the Aleutian Islands extend into the Eastern Hemisphere. Alaska is the only non-contiguous U. S. state on continental North America and it is technically part of the continental U. S. but is sometimes not included in colloquial use, Alaska is not part of the contiguous U. S. often called the Lower 48. The capital city, Juneau, is situated on the mainland of the North American continent but is not connected by road to the rest of the North American highway system. Alaskas territorial waters touch Russias territorial waters in the Bering Strait, as the Russian Big Diomede Island, Alaska has a longer coastline than all the other U. S. states combined. Alaska is the largest state in the United States by total area at 663,268 square miles, over twice the size of Texas, Alaska is larger than all but 18 sovereign countries. Counting territorial waters, Alaska is larger than the area of the next three largest states, Texas, California, and Montana. It is also larger than the area of the 22 smallest U. S. states. Also referred to as the Panhandle or Inside Passage, this is the region of Alaska closest to the rest of the United States, as such, this was where most of the initial non-indigenous settlement occurred in the years following the Alaska Purchase. The region is dominated by the Alexander Archipelago as well as the Tongass National Forest and it contains the state capital Juneau, the former capital Sitka, and Ketchikan, at one time Alaskas largest city. The Alaska Marine Highway provides a vital transportation link throughout the area. The Interior is the largest region of Alaska, much of it is uninhabited wilderness, Fairbanks is the only large city in the region
24.
Algonquian languages
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The Algonquian languages are a subfamily of Native American languages which includes most of the languages in the Algic language family. The term Algonquin has been suggested to derive from the Maliseet word elakómkwik, a number of Algonquian languages, like many of the Iroquoian languages of the hereditary enemies of the Algonquian peoples, have already become extinct. Speakers of Algonquian languages stretch from the east coast of North America to the Rocky Mountains, the proto-language from which all of the languages of the family descend, Proto-Algonquian, was spoken around 2,500 to 3,000 years ago. There is no consensus about where this language was spoken. This subfamily of around 30 languages is divided into three groups according to geography, Plains, Central, and Eastern Algonquian, only Eastern Algonquian constitutes a true genetic subgroup. The languages are listed below, following the classifications of Goddard, extinct languages are marked with †, and endangered languages are noted as such. For dialects and subdialects, consult the separate articles for each of the three divisions. Shinnecock Eastern Algonquian is a true genetic subgrouping, the Plains Algonquian and the Central Algonquian groups are not genetic groupings but rather areal groupings. However, these groups often do share linguistic features. Paul Proulx has argued that this view is incorrect. However, this scheme has failed to gain acceptance from other specialists in the Algonquian languages. Instead, the commonly accepted subgrouping scheme is that proposed by Ives Goddard, by this scenario, Blackfoot was the first language to branch off, which coincides well with its being the most divergent language of Algonquian. In west-to-east order, the subsequent branchings were, Arapaho-Gros Ventre, Cree-Montagnais, Menominee, and Cheyenne, then the core Great Lakes languages, and finally, Proto-Eastern Algonquian. Goddard also points out there is clear evidence for pre-historical contact between Eastern Algonquian and Cree-Montagnais, as well as between Cheyenne and Arapaho-Gros Ventre. There has long been especially extensive back-and-forth influence between Cree and Ojibwe and they share certain intriguing lexical and phonological innovations. But, this theory has not yet been fully fleshed out and is still considered conjectural, linguistic evidence is scarce and poorly recorded however, and it is unlikely that reliable evidence of a connection can be found. The Algonquian language family is known for its complex polysynthetic morphology, statements that take many words to say in English can be expressed with a single word. Ex, paehtāwāēwesew He is heard by higher powers or kāstāhikoyahk it frightens us and these languages have been extensively studied by Leonard Bloomfield, Ives Goddard, and others
25.
Taiga
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Taiga also known as boreal forest or snow forest, is a biome characterized by coniferous forests consisting mostly of pines, spruces and larches. The taiga is the worlds largest biome apart from the oceans, in North America it covers most of inland Canada and Alaska as well as parts of the extreme northern continental United States, where it is known as the Northwoods or North woods. However, the tree species, the length of the growing season. Hoffman discusses the origin of this use in North America. Although at high elevations taiga grades into alpine tundra through Krummholz, it is not an alpine biome only like subalpine forest, Taiga is the worlds largest land biome, and makes up 29% of the worlds forest cover, the largest areas are located in Russia and Canada. The taiga is the terrestrial biome with the lowest annual average temperatures after the tundra, extreme winter minimums in the northern taiga are typically lower than those of the tundra. The lowest reliably recorded temperatures in the Northern Hemisphere were recorded in the taiga of northeastern Russia, the taiga or boreal forest has a subarctic climate with very large temperature range between seasons, but the long and cold winter is the dominant feature. This climate is classified as Dfc, Dwc, Dsc, Dfd and Dwd in the Köppen climate classification scheme, meaning that the summer lasts 1–3 months. In Siberian taiga the average temperature of the coldest month is between −6 °C and −50 °C, the mean annual temperature generally varies from -5 °C to 5 °C, but there are taiga areas in eastern Siberia and interior Alaska-Yukon where the mean annual reaches down to -10 °C. According to some sources, the boreal forest grades into a mixed forest when mean annual temperature reaches about 3 °C. The winters, with temperatures below freezing, last five to seven months. Temperatures vary from −54 °C to 30 °C throughout the whole year, the summers, while short, are generally warm and humid. In much of the taiga, -20 °C would be a winter day temperature and 18 °C an average summer day. In Canada, Scandinavia and Finland, the season is often estimated by using the period of the year when the 24-hour average temperature is +5 °C or more. For the Taiga Plains in Canada, growing season varies from 80 to 150 days, some sources claim 130 days growing season as typical for the taiga. Other sources mention that 50–100 frost-free days are characteristic, data for locations in southwest Yukon gives 80–120 frost-free days. The closed canopy boreal forest in Kenozersky National Park near Plesetsk, Arkhangelsk Province, Russia, the longest growing season is found in the smaller areas with oceanic influences, in coastal areas of Scandinavia and Finland, the growing season of the closed boreal forest can be 145–180 days. High latitudes mean that the sun not rise far above the horizon
26.
Deciduous
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In a more general sense, deciduous means the dropping of a part that is no longer needed or falling away after its purpose is finished. In plants it is the result of natural processes, in botany and horticulture, deciduous plants, including trees, shrubs and herbaceous perennials, are those that lose all of their leaves for part of the year. In some cases leaf loss coincides with winter—namely in temperate or polar climates, in other parts of the world, including tropical, subtropical, and arid regions, plants lose their leaves during the dry season or other seasons, depending on variations in rainfall. The converse of deciduous is coniferous, where foliage is shed on a different schedule from deciduous trees, plants that are intermediate may be called semi-deciduous, they lose old foliage as new growth begins. Other plants are semi-evergreen and lose their leaves before the growing season. Many deciduous plants flower during the period when they are leafless, the absence of leaves improves wind transmission of pollen for wind-pollinated plants and increases the visibility of the flowers to insects in insect-pollinated plants. This strategy is not without risks, as the flowers can be damaged by frost or, in dry season regions, leaf drop or abscission involves complex physiological signals and changes within plants. The process of photosynthesis steadily degrades the supply of chlorophylls in foliage, the brightest leaf colors are produced when days grow short and nights are cool, but remain above freezing. These other pigments include carotenoids that are yellow, brown, anthocyanin pigments produce red and purple colors, though they are not always present in the leaves. Rather, they are produced in the foliage in late summer, parts of the world that have showy displays of bright autumn colors are limited to locations where days become short and nights are cool. In other parts of the world, the leaves of deciduous trees simply fall off without turning the bright colors produced from the accumulation of anthocyanin pigments, the beginnings of leaf drop starts when an abscission layer is formed between the leaf petiole and the stem. This layer is formed in the spring during active new growth of the leaf, the cells are sensitive to a plant hormone called auxin that is produced by the leaf and other parts of the plant. The elongation of cells break the connection between the different cell layers, allowing the leaf to break away from the plant. It also forms a layer that seals the break, so the plant does not lose sap, in the spring, these proteins are used as a nitrogen source during the growth of new leaves or flowers. Plants with deciduous foliage have advantages and disadvantages compared to plants with evergreen foliage, evergreens suffer greater water loss during the winter and they also can experience greater predation pressure, especially when small. Losing leaves in winter may reduce damage from insects, repairing leaves, removing leaves also reduces cavitation which can damage xylem vessels in plants. This then allows deciduous plants to have xylem vessels with larger diameters, the deciduous characteristic has developed repeatedly among woody plants. Trees include maple, many oaks and nothofagus, elm, aspen, Deciduous shrubs include honeysuckle, viburnum, and many others
27.
Tree
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In botany, a tree is a perennial plant with an elongated stem, or trunk, supporting branches and leaves in most species. In some usages, the definition of a tree may be narrower, including only woody plants with secondary growth, plants that are usable as lumber or plants above a specified height. Trees are not a group but include a variety of plant species that have independently evolved a woody trunk. In looser senses, the palms, the tree ferns, bananas. Trees tend to be long-lived, some reaching several thousand years old, the tallest known tree, a coast redwood named Hyperion, stands 115.6 m high. Trees have been in existence for 370 million years and it is estimated that there are just over 3 trillion mature trees in the world. A tree typically has many secondary branches supported clear of the ground by the trunk and this trunk typically contains woody tissue for strength, and vascular tissue to carry materials from one part of the tree to another. For most trees it is surrounded by a layer of bark which serves as a protective barrier, below the ground, the roots branch and spread out widely, they serve to anchor the tree and extract moisture and nutrients from the soil. Above ground, the divide into smaller branches and shoots. The shoots typically bear leaves, which light energy and convert it into sugars by photosynthesis, providing the food for the trees growth. Flowers and fruit may also be present, but some trees, such as conifers, instead have pollen cones and seed cones, others, such as tree ferns, trees play a significant role in reducing erosion and moderating the climate. They remove carbon dioxide from the atmosphere and store large quantities of carbon in their tissues, trees and forests provide a habitat for many species of animals and plants. Tropical rainforests are one of the most biodiverse habitats in the world, trees provide shade and shelter, timber for construction, fuel for cooking and heating, and fruit for food as well as having many other uses. In parts of the world, forests are shrinking as trees are cleared to increase the amount of available for agriculture. Because of their longevity and usefulness, trees have always revered, with sacred groves in various cultures. Although tree is a term of common parlance, there is no universally recognised precise definition of what a tree is, either botanically or in common language. In its broadest sense, a tree is any plant with the form of an elongated stem, or trunk. Trees are also defined by height, with smaller plants from 0.5 to 10 m being called shrubs
28.
Leaf
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A leaf is an organ of a vascular plant and is the principal lateral appendage of the stem. The leaves and stem together form the shoot, Leaves are collectively referred to as foliage, as in autumn foliage. Although leaves can be seen in different shapes, sizes and textures, typically a leaf is a thin, dorsiventrally flattened organ, borne above ground. Most leaves have distinctive upper surface and lower surface that differ in colour, hairiness, broad, flat leaves with complex venation are known as megaphylls and the species that bear them, the majority, as broad-leaved or megaphyllous plants. In others, such as the clubmosses, with different evolutionary origins, some leaves, such as bulb scales are not above ground, and in many aquatic species the leaves are submerged in water. Succulent plants often have thick juicy leaves, but some leaves are without major photosynthetic function and may be dead at maturity, as in some cataphylls, furthermore, several kinds of leaf-like structures found in vascular plants are not totally homologous with them. Examples include flattened plant stems called phylloclades and cladodes, and flattened leaf stems called phyllodes which differ from both in their structure and origin. Many structures of plants, such as the phyllids of mosses and liverworts and even of some foliose lichens. Leaves are the most important organs of most vascular plants and these are then further processed by chemical synthesis into more complex organic molecules such as cellulose, the basic structural material in plant cell walls. The plant must therefore bring these three together in the leaf for photosynthesis to take place. Once sugar has been synthesized, it needs to be transported to areas of growth such as the plant shoots and roots. Vascular plants transport sucrose in a tissue called the phloem. The phloem and xylem are parallel to each other but the transport of materials is usually in opposite directions. Within the leaf these vascular systems branch to form veins which supply as much as the leaf as possible and they are arranged on the plant so as to expose their surfaces to light as efficiently as possible without shading each other, but there are many exceptions and complications. For instance plants adapted to windy conditions may have pendent leaves, such as in many willows, the flat, or laminar, shape also maximises thermal contact with the surrounding air, promoting cooling. Functionally, in addition to photosynthesis the leaf is the site of transpiration and guttation. Many gymnosperms have thin needle-like or scale-like leaves that can be advantageous in cold climates with frequent snow and these are interpreted as reduced from megaphyllous leaves of their Devonian ancestors. For xerophytes the major constraint is not light flux or intensity, some window plants such as Fenestraria species and some Haworthia species such as Haworthia tesselata and Haworthia truncata are examples of xerophytes. and Bulbine mesembryanthemoides
29.
Conifer cone
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A cone is an organ on plants in the division Pinophyta that contains the reproductive structures The familiar woody cone is the female cone, which produces seeds. The male cones, which produce pollen, are usually herbaceous, the name cone derives from the fact that the shape in some species resembles a geometric cone. The individual plates of a cone are known as scales, the male cone is structurally similar across all conifers, differing only in small ways from species to species. Extending out from an axis are microsporophylls. Under each microsporophyll is one or several microsporangia, the female cone contains ovules which, when fertilized by pollen, become seeds. The female cone structure varies more markedly between the different conifer families, and is crucial for the identification of many species of conifers. The members of the family have cones that are imbricate. The scales are arranged in fibonacci number ratios. The female cone has two types of scale, the scales, derived from a modified leaf, and the seed scales, one subtended by each bract scale. On the upper-side base of each seed scale are two ovules that develop into seeds after fertilization by pollen grains. The bract scales develop first, and are conspicuous at the time of pollination, the scales open temporarily to receive gametophytes, then close during fertilization and maturation, and then re-open again at maturity to allow the seed to escape. Maturation takes 6–8 months from pollination in most Pinaceae genera, but 12 months in cedars, the cones open either by the seed scales flexing back when they dry out, or by the cones disintegrating with the seed scales falling off. The cones are conic, cylindrical or ovoid, and small to large, from 2–60 cm long. After ripening, the opening of non-serotinous pine cones is associated with their moisture content—cones are open when dry and this assures that the small, wind disseminated seeds will be dispersed during relatively dry weather, and thus, the distance traveled from the parent tree will be enhanced. A pine cone will go through cycles of opening and closing during its life span. This process occurs with older cones while attached to branches and even after the cones have fallen to the forest floor. The condition of fallen pine cones is an indication of the forest floors moisture content. Closed cones indicate damp conditions while open cones indicate the forest floor is dry, members of the Araucariaceae have the bract and seed scales fully fused, and have only one ovule on each scale
30.
Seed
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A seed is an embryonic plant enclosed in a protective outer covering. The formation of the seed is part of the process of reproduction in seed plants, Seeds are the product of the ripened ovule, after fertilization by pollen and some growth within the mother plant. The embryo is developed from the zygote and the coat from the integuments of the ovule. Seed plants now dominate biological niches on land, from forests to both in hot and cold climates. The term seed also has a meaning that antedates the above—anything that can be sown, e. g. seed potatoes. In the case of sunflower and corn seeds, what is sown is the seed enclosed in a shell or husk, many structures commonly referred to as seeds are actually dry fruits. Plants producing berries are called baccate, sunflower seeds are sometimes sold commercially while still enclosed within the hard wall of the fruit, which must be split open to reach the seed. Different groups of plants have other modifications, the stone fruits have a hardened fruit layer fused to. Nuts are the one-seeded, hard-shelled fruit of plants with an indehiscent seed. Seeds are produced in several related groups of plants, and their manner of production distinguishes the angiosperms from the gymnosperms, angiosperm seeds are produced in a hard or fleshy structure called a fruit that encloses the seeds, hence the name. Some fruits have layers of hard and fleshy material. In gymnosperms, no special structure develops to enclose the seeds, however, the seeds do become covered by the cone scales as they develop in some species of conifer. Seed production in natural plant populations varies widely from year-to-year in response to weather variables, insects and diseases, over a 20-year period, for example, forests composed of loblolly pine and shortleaf pine produced from 0 to nearly 5 million sound pine seeds per hectare. Over this period, there were six bumper, five poor, and nine good seed crops, right after fertilization, the zygote is mostly inactive, but the primary endosperm divides rapidly to form the endosperm tissue. This tissue becomes the food the young plant will consume until the roots have developed after germination, after fertilization the ovules develop into the seeds. The ovule consists of a number of components, The funicle or seed stalk which attaches the ovule to the placenta and hence ovary or fruit wall, the nucellus, the remnant of the megasporangium and main region of the ovule where the megagametophyte develops. The micropyle, a pore or opening in the apex of the integument of the ovule where the pollen tube usually enters during the process of fertilization. The chalaza, the base of the ovule opposite the micropyle, the shape of the ovules as they develop often affects the final shape of the seeds
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Pollination
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Pollination is the process by which pollen is transferred to the female reproductive organs of a plant, thereby enabling fertilization to take place. Like all living organisms, seed plants have a major goal. The reproductive unit is the seed, and pollination is a step in the production of seeds in all spermatophytes. The process is different in angiosperms from what it is in gymnosperms. In angiosperms, after the grain has landed on the stigma. Sperm cells from the pollen grain then move along the tube, enter the egg cell through the micropyle and fertilise it. A successful angiosperm pollen grain containing the gametes is transported to the stigma. Its two gametes travel down the tube to where the containing the female gametes are held within the carpel. One nucleus fuses with the bodies to produce the endosperm tissues. In gymnosperms, the ovule is not contained in a carpel, details of the process vary according to the division of gymnosperms in question. Two main modes of fertilization are found in gymnosperms, the study of pollination brings together many disciplines, such as botany, horticulture, entomology, and ecology. The pollination process as an interaction between flower and pollen vector was first addressed in the 18th century by Christian Konrad Sprengel and it is important in horticulture and agriculture, because fruiting is dependent on fertilization, the result of pollination. The study of pollination by insects is known as anthecology, pollen germination has three stages, hydration, activation and pollen tube emergence. The pollen grain is severely dehydrated so that its mass is reduced enabling it to be easily transported from flower to flower. Germination only takes place after rehydration, ensuring that premature germination does not take place in the anther, hydration allows the plasma membrane of the pollen grain to reform into its normal bilayer organization providing an effective osmotic membrane. Activation involves the development of actin filaments throughout the cytoplasm of the cell, hydration and activation continue as the pollen tube begins to grow. In conifers, the structures are borne on cones. The cones are either pollen cones or ovulate cones, but some species are monoecious, a pollen cone contains hundreds of microsporangia carried on reproductive structures called sporophylls
32.
Arctic
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The Arctic is a polar region located at the northernmost part of Earth. The Arctic consists of the Arctic Ocean, adjacent seas, and parts of Alaska, Canada, Finland, Greenland, Iceland, Norway, Russia, land within the Arctic region has seasonally varying snow and ice cover, with predominantly treeless permafrost-containing tundra. Arctic seas contain seasonal sea ice in many places, the Arctic region is a unique area among Earths ecosystems. For example, the cultures in the region and the Arctic indigenous peoples have adapted to its cold, in recent years, Arctic sea ice decline has been caused by global warming. Life in the Arctic includes organisms living in the ice, zooplankton and phytoplankton, fish and marine mammals, birds, land animals, plants, Arctic land is bordered by the subarctic. The word Arctic comes from the Greek word ἀρκτικός, near the Bear, northern, there are a number of definitions of what area is contained within the Arctic. The area can be defined as north of the Arctic Circle, the southern limit of the midnight sun. The Arctics climate is characterized by cold winters and cool summers and its precipitation mostly comes in the form of snow and is low, with most of the area receiving less than 50 cm. High winds often stir up snow, creating the illusion of continuous snowfall, average winter temperatures can be as low as −40 °C, and the coldest recorded temperature is approximately −68 °C. Coastal Arctic climates are moderated by oceanic influences, having generally warmer temperatures, the Arctic is affected by current global warming, leading to Arctic sea ice shrinkage, diminished ice in the Greenland ice sheet, and Arctic methane release as the permafrost thaws. Due to the migration of the planets isotherms, the Arctic region is currently shrinking. Perhaps the most spectacular result of Arctic shrinkage is sea ice loss, there is a large variance in predictions of Arctic sea ice loss, with models showing near-complete to complete loss in September from 2040 to some time well beyond 2100. About half of the models show near-complete to complete sea ice loss in September by the year 2100. Arctic life is characterized by adaptation to short growing seasons with long periods of sunlight, Arctic vegetation is composed of plants such as dwarf shrubs, graminoids, herbs, lichens and mosses, which all grow relatively close to the ground, forming tundra. As one moves northward, the amount of available for plant growth decreases considerably. Colder summer temperatures cause the size, abundance, productivity and variety of plants to decrease, trees cannot grow in the Arctic, but in its warmest parts, shrubs are common and can reach 2 m in height, sedges, mosses and lichens can form thick layers. In the coldest parts of the Arctic, much of the ground is bare, non-vascular plants such as lichens and mosses predominate, along with a few scattered grasses, herbivores on the tundra include the Arctic hare, lemming, muskox, and caribou. They are preyed on by the owl, Arctic fox, Grizzly bear
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Tree line
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The tree line is the edge of the habitat at which trees are capable of growing. It is found at high elevations and in frigid environments, beyond the tree line, trees cannot tolerate the environmental conditions. The tree line should not be confused with a lower timberline or forest line, at the tree line, tree growth is often sparse and stunted, with the last trees forming densely matted bushes, known as krummholz. The tree line, like other natural lines, appears well-defined from a distance. Trees grow shorter towards the inhospitable climate until they stop growing. The climate above the line of mountains is called an alpine climate. In the northern hemisphere treelines on north-facing slopes are lower than on south-facing slopes because the increased shade on north-facing slopes means the snowpack takes longer to melt and this shortens the growing season for trees. In the southern hemisphere, the slopes have the shorter growing season. The alpine tree line boundary is seldom abrupt, it forms a transition zone between closed forest below and treeless alpine tundra above. Environmentally dwarfed shrubs commonly forms the upper limit, the decrease in air temperature due to increasing elevation causes the alpine climate. Skin effects and topography can create microclimates that alter the general cooling trend, however, the number of degree days calculated from leaf temperatures may be very similar in the two kinds of timberlines. A series of warm summers in the 1940s seems to have permitted the establishment of “significant numbers” of spruce seedlings above the treeline in the hills near Fairbanks. Survival depends on a sufficiency of new growth to support the tree, the windiness of high-elevation sites is also a potent determinant of the distribution of tree growth. However, snow accumulation in sheltered gullies in the Selkirk Mountains of southeastern British Columbia causes timberline to be 400 metres lower than on exposed intervening shoulders. In a desert, the line marks the driest places where trees can grow. These tend to be called the tree line and occur below about 5,000 ft elevation in the Desert Southwestern United States. In some mountainous areas, higher elevations above the line or on equator-facing and leeward slopes can result in low rainfall. This dries out the soil, resulting in an arid environment unsuitable for trees
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Tundra
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In physical geography, tundra is a type of biome where the tree growth is hindered by low temperatures and short growing seasons. The term tundra comes through Russian тундра from the Kildin Sami word tūndâr uplands, there are three types of tundra, Arctic tundra, alpine tundra, and Antarctic tundra. In tundra, the vegetation is composed of shrubs, sedges and grasses, mosses. Scattered trees grow in tundra regions. The ecotone between the tundra and the forest is known as the line or timberline. Arctic tundra occurs in the far Northern Hemisphere, north of the taiga belt, the word tundra usually refers only to the areas where the subsoil is permafrost, or permanently frozen soil. Permafrost tundra includes vast areas of northern Russia and Canada, the polar tundra is home to several peoples who are mostly nomadic reindeer herders, such as the Nganasan and Nenets in the permafrost area. Arctic tundra contains areas of landscape and is frozen for much of the year. The soil there is frozen from 25–90 cm down, and it is impossible for trees to grow, instead, bare and sometimes rocky land can only support low growing plants such as moss, heath, and lichen. There are two seasons, winter and summer, in the polar tundra areas. During the winter it is cold and dark, with the average temperature around −28 °C. However, extreme temperatures on the tundra do not drop as low as those experienced in taiga areas further south. During the summer, temperatures rise somewhat, and the top layer of seasonally-frozen soil melts, the tundra is covered in marshes, lakes, bogs and streams during the warm months. Generally daytime temperatures during the rise to about 12 °C. Arctic tundras are sometimes the subject of conservation programs. In Canada and Russia, many of areas are protected through a national Biodiversity Action Plan. Tundra tends to be windy, with winds often blowing upwards of 50–100 km/h, however, in terms of precipitation, it is desert-like, with only about 15–25 cm falling per year. Although precipitation is light, evaporation is also relatively minimal, there is a natural pattern of accumulation of fuel and wildfire which varies depending on the nature of vegetation and terrain
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Swamp
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A swamp is a wetland that is forested. Many swamps occur along rivers where they are critically dependent upon natural water level fluctuations. Other swamps occur on the shores of large lakes, some swamps have hammocks, or dry-land protrusions, covered by aquatic vegetation, or vegetation that tolerates periodic inundation. The two main types of swamp are true or swamp forests and transitional or shrub swamps, in the boreal regions of Canada, the word swamp is colloquially used for what is more correctly termed a bog or muskeg. The water of a swamp may be water, brackish water or seawater. Some of the worlds largest swamps are found along rivers such as the Amazon, the Mississippi. Swamps are characterized by slow-moving to stagnant waters and they are usually associated with adjacent rivers or lakes. Swamps are features of areas with very low topographic relief, historically, humans have drained swamps to provide additional land for agriculture and to reduce the threat of diseases borne by swamp insects and similar animals. Many swamps have also undergone intensive logging, requiring the construction of drainage ditches and these ditches and canals contributed to drainage and, along the coast, allowed salt water to intrude, converting swamps to marsh or even to open water. Large areas of swamp were therefore lost or degraded, louisiana provides a classic example of wetland loss from these combined factors. Europe has probably lost nearly half its wetlands, New Zealand lost 90 percent of its wetlands over a period of 150 years. Ecologists recognise that swamps provide valuable ecological services including flood control, fish production, water purification, carbon storage, in many parts of the world authorities protect swamps. In parts of Europe and North America, swamp restoration projects are becoming widespread, often the simplest steps to restoring swamps involve plugging drainage ditches and removing levees. Swamps and other wetlands have traditionally held a very low property value compared to fields, prairies and they have a reputation for being unproductive land that cannot easily be utilized for human activities, other than perhaps hunting and trapping. Farmers, for example, typically drained swamps next to their fields so as to more land usable for planting crops. Many societies now realize that swamps are important to providing fresh water and oxygen to all life. Indeed, floodplain swamps are important in fish production. Government environmental agencies are taking steps to protect and preserve swamps, in Europe, major effort is being invested in the restoration of swamp forests along rivers
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Bog
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A bog is a wetland that accumulates peat, a deposit of dead plant material—often mosses, and in a majority of cases, sphagnum moss. It is one of the four types of wetlands. Other names for bogs include mire, quagmire, and muskeg and they are frequently covered in ericaceous shrubs rooted in the sphagnum moss and peat. The gradual accumulation of decayed plant material in a bog functions as a carbon sink, Bogs occur where the water at the ground surface is acidic and low in nutrients. In some cases, the water is derived entirely from precipitation, water flowing out of bogs has a characteristic brown colour, which comes from dissolved peat tannins. In general, the low fertility and cool climate results in relatively slow plant growth, large areas of landscape can be covered many metres deep in peat. Bogs have distinctive assemblages of animal, fungal and plant species, Bogs are widely distributed in cold, temperate climes, mostly in boreal ecosystems in the Northern Hemisphere. The worlds largest wetland is the bogs of the Western Siberian Lowlands in Russia. Large peat bogs also occur in North America, particularly the Hudson Bay Lowland and they are less common in the Southern Hemisphere, with the largest being the Magellanic moorland, comprising some 44,000 square kilometres. Sphagnum bogs were widespread in northern Europe but have often been cleared and drained for agriculture, a 2014 expedition leaving from Itanga village, Republic of the Congo discovered a peat bog as big as England which stretches into neighboring Democratic Republic of Congo. There are many highly specialised animals, fungi and plants associated with bog habitat, most are capable of tolerating the combination of low nutrient levels and waterlogging. Sphagnum moss is generally abundant, along with ericaceous shrubs, the shrubs are often evergreen, which is understood to assist in conservation of nutrients. In drier locations, evergreen trees can occur, in case the bog blends into the surrounding expanses of boreal evergreen forest. Sedges are one of the more common herbaceous species, carnivorous plants such as sundews and pitcher plants have adapted to the low-nutrient conditions by using invertebrates as a nutrient source. Orchids have adapted to these conditions through the use of fungi to extract nutrients. Some shrubs such as Myrica gale have root nodules in which nitrogen fixation occurs, Bogs are recognized as a significant/specific habitat type by a number of governmental and conservation agencies. They can provide habitat for mammals, such as caribou, moose, the United Kingdom in its Biodiversity Action Plan establishes bog habitats as a priority for conservation. Russia has a reserve system in the West Siberian Lowland
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Muskeg
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Muskeg is an acidic soil type common in Arctic and boreal areas, although it is found in other northern climates as well. Muskeg is approximately synonymous with bogland, but muskeg is the term in Western Canada and Alaska. The term became common in these areas because it is of Cree origin, large tracts of this soil existing in Siberia may be called muskeg or bogland interchangeably. Muskeg consists of plants in various states of decomposition, ranging from fairly intact sphagnum moss, to sedge peat. Pieces of wood can make up five to 15 percent of the peat soil, Muskeg tends to have a water table near the surface. The sphagnum moss forming it can hold 15 to 30 times its own weight in water, Muskeg patches are ideal habitats for beavers, pitcher plants, agaric mushrooms and a variety of other organisms. Muskeg forms because permafrost, clay or bedrock prevents water drainage, the water from rain and snow collects, forming permanently waterlogged vegetation and stagnant pools. It needs two conditions to develop, abundant rain and cool summers, a dead plant that falls on dry soil is normally attacked by bacteria and fungi and quickly rots. If the same plant lands in water or on saturated soil, less oxygen is available under water, so aerobic bacteria and fungi fail to colonize the submerged debris effectively. In addition, cool temperatures retard bacterial and fungal growth and this causes slow decomposition, and thus the plant debris gradually accumulates to form peat and eventually muskeg. Depending on the topography of the land, muskeg can reach depths greater than 30 metres. Although at first glance resembles a plain covered with short grasses. Small stands of stunted and often dead trees, which vaguely resemble Bonsai trees, grow where land protrudes above the water table and its grassland appearance invites the unwary to walk on it, but even the most solid muskeg is spongy and waterlogged. Traveling through muskeg is a strange and dangerous experience for the unaccustomed, Muskeg can grow atop bodies of water, especially small ponds and streams. Because of the water beneath, the muskeg surface sometimes ripples underfoot, thinner patches allow large animals to fall through, becoming trapped under the muskeg and drowning. Moose are at a disadvantage in muskeg due to their long legs, minimal hoof area. Hunters and hikers may occasionally encounter young moose in muskeg-covered ponds submerged up to their torsos or necks, Muskeg can be a significant impediment to transportation. During the 1870s, muskeg in Northern Ontario was reported to have swallowed a railroad engine whole when a track was laid on muskeg instead of clearing down to bedrock
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Sphagnum
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Sphagnum is a genus of approximately 380 accepted species of mosses, commonly known as peat moss. The empty cells help retain water in drier conditions, hence, as sphagnum moss grows, it can slowly spread into drier conditions, forming larger mires, both raised bogs and blanket bogs. These peat accumulations then provide habitat for an array of peatland plants, including sedges and ericaceous shrubs, as well as orchids. Sphagnum and the formed from it do not decay readily because of the phenolic compounds embedded in the mosss cell walls. In addition, bogs, like all wetlands, develop anaerobic soil conditions, Peat moss can also acidify its surroundings by taking up cations, such as calcium and magnesium, and releasing hydrogen ions. Under the right conditions, peat can accumulate to a depth of many meters, different species of Sphagnum have different tolerance limits for flooding and pH, so any one peatland may have a number of different Sphagnum species. Individual peat moss plants consist of a stem, with tightly arranged clusters of branch fascicles usually consisting of two or three spreading branches and two to four hanging branches. The top of the plant, or capitulum, has compact clusters of young branches, along the stem are scattered leaves of various shapes, named stem leaves, the shape varies according to species. The leaves consist of two kinds of cells, small, green, living cells, and large, clear, structural, the latter have the large water-holding capacity. Sphagnum, like all other plants, has an alternation of generations, like other bryophytes. Unlike other mosses, the long-lived gametophytes do not rely upon rhizoids to assist in water uptake, Sphagnum species can be unisexual or bisexual, In North America, 80% of Sphagnum species are unisexual. Gametophytes have substantial asexual reproduction by fragmentation, producing much of the material in sphagnum peatlands. Swimming sperm fertilize eggs contained in archegonia that remain attached to the female gametophyte, the sporophyte is relatively short-lived, and consists almost entirely of a shiny green, spherical spore capsule that becomes black with spores. Sporophytes are raised on stalks to facilitate spore dispersal, but unlike other mosses, the spores germinate to produce minute protonemae, which start as filaments, can become thalloid, and can produce a few rhizoids. Soon afterwards the protonema develops buds and these differentiate into its characteristic, erect, leafy, branched gametophyte with chlorophyllose cells and this stage dominates the environment where Sphagnum grows, obliterating and burying the protonema and eventually building up into layers of dead moss called peat. Carpets of living Sphagnum may be attacked by fungi, and one fungus that is also a mushroom, Sphagnurus paluster. It is unknown whether the stage can produce such gemmae. Peat moss can be distinguished from other species by its unique branch clusters
39.
Peat
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Peat, also called turf, is an accumulation of partially decayed vegetation or organic matter that is unique to natural areas called peatlands, bogs, mires, moors, or muskegs. The peatland ecosystem is the most efficient carbon sink on the planet because peatland plants capture the CO2 which is released from the peat. Sphagnum moss is one of the most common components in peat, soils that contain mostly peat are known as histosols. Peat forms in wetland conditions, where flooding obstructs flows of oxygen from the atmosphere, Peatlands, particularly bogs, are the most important source of peat. That said, other less common types, including fens, pocosins. Landscapes covered in peat are home to specific kinds of plants including Sphagnum moss, ericaceous shrubs, because organic matter accumulates over thousands of years, peat deposits also provide records of past vegetation and climates stored in plant remains like pollen. This allows humans to reconstruct past environments and study changes in land use. Peat is harvested as an important source of fuel in certain parts of the world, by volume, there are about 4 trillion cubic metres of peat in the world, covering a total of around 2% of the global land area, containing about 8 billion terajoules of energy. Over time, the formation of peat is often the first step in the formation of other fossil fuels such as coal. This is also the commonly used in the peat industry. At 106 g CO2/MJ, the carbon dioxide emission intensity of peat is higher than that of coal and natural gas, lastly, peat fires have been responsible for large public health disasters including the 1997 Southeast Asian haze. Peat forms when plant material does not fully decay in acidic and anaerobic conditions and it is composed mainly of wetland vegetation, principally bog plants including mosses, sedges, and shrubs. As it accumulates, the peat holds water and this slowly creates wetter conditions that allow the area of wetland to expand. Peatland features can include ponds, ridges, and raised bogs, most modern peat bogs formed 12,000 years ago in high latitudes after the glaciers retreated at the end of the last ice age. Peat usually accumulates slowly at the rate of about a millimetre per year, using Carbon-dating, scientists found that Peat in peatlands started forming 360 million years ago based on it currently containing 550 Gt of carbon. Peat material is either fibric, hemic, or sapric, fibric peats are the least decomposed and consist of intact fiber. Hemic peats are partially decomposed and sapric are the most decomposed, Phragmites peat is one composed of reed grass, Phragmites australis, and other grasses. It is denser than other types of peat
40.
Calcareous
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Calcareous is an adjective meaning mostly or partly composed of calcium carbonate, in other words, containing lime or being chalky. The term is used in a variety of scientific disciplines. The term also applies to the calcium carbonate tests of often more or less microscopic Foraminifera, note that not all tests are calcareous, diatoms and radiolaria have siliceous tests. The molluscs are calcareous, as are calcareous sponges, that have spicules which are made of calcium carbonate, calcareous grassland is a form of grassland characteristic of soils containing much calcium carbonate from underlying chalk or limestone rock. Species of algae such as the green-segmented genus Halimeda are calcareous, the term is used in pathology, for example in calcareous conjunctivitis, and when referring to calcareous metastasis or calcareous deposits, which may both be removed surgically. The term calcareous can be applied to a sediment, sedimentary rock, or soil type which is formed from, or contains a proportion of. Calcareous sediments are deposited in shallow water near land, since the carbonate is precipitated by marine organisms that need land-derived nutrients. Generally speaking, the farther from land sediments fall, the less calcareous they are, some areas can have interbedded calcareous sediments due to storms, or changes in ocean currents. Calcareous ooze is a form of calcium derived from planktonic organisms that accumulates on the sea floor. This can only occur if the ocean is shallower than the carbonate compensation depth, below this depth, calcium carbonate begins to dissolve in the ocean, and only non-calcareous sediments are stable, such as siliceous ooze or pelagic red clay. Calcareous soils are alkaline, in other words they have a high pH. This is because of the very weak acidity of carbonic acid, note that this is not the only reason for a high soil pH
41.
Limestone
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Limestone is a sedimentary rock, composed mainly of skeletal fragments of marine organisms such as coral, forams and molluscs. Its major materials are the minerals calcite and aragonite, which are different crystal forms of calcium carbonate, about 10% of sedimentary rocks are limestones. The solubility of limestone in water and weak acid solutions leads to karst landscapes, most cave systems are through limestone bedrock. The first geologist to distinguish limestone from dolomite was Belsazar Hacquet in 1778, like most other sedimentary rocks, most limestone is composed of grains. Most grains in limestone are skeletal fragments of organisms such as coral or foraminifera. Other carbonate grains comprising limestones are ooids, peloids, intraclasts and these organisms secrete shells made of aragonite or calcite, and leave these shells behind when they die. Limestone often contains variable amounts of silica in the form of chert or siliceous skeletal fragment, some limestones do not consist of grains at all, and are formed completely by the chemical precipitation of calcite or aragonite, i. e. travertine. Secondary calcite may be deposited by supersaturated meteoric waters and this produces speleothems, such as stalagmites and stalactites. Another form taken by calcite is oolitic limestone, which can be recognized by its granular appearance, the primary source of the calcite in limestone is most commonly marine organisms. Some of these organisms can construct mounds of rock known as reefs, below about 3,000 meters, water pressure and temperature conditions cause the dissolution of calcite to increase nonlinearly, so limestone typically does not form in deeper waters. Limestones may also form in lacustrine and evaporite depositional environments, calcite can be dissolved or precipitated by groundwater, depending on several factors, including the water temperature, pH, and dissolved ion concentrations. Calcite exhibits a characteristic called retrograde solubility, in which it becomes less soluble in water as the temperature increases. Impurities will cause limestones to exhibit different colors, especially with weathered surfaces, Limestone may be crystalline, clastic, granular, or massive, depending on the method of formation. Crystals of calcite, quartz, dolomite or barite may line small cavities in the rock, when conditions are right for precipitation, calcite forms mineral coatings that cement the existing rock grains together, or it can fill fractures. Travertine is a banded, compact variety of limestone formed along streams, particularly there are waterfalls. Calcium carbonate is deposited where evaporation of the leaves a solution supersaturated with the chemical constituents of calcite. Tufa, a porous or cellular variety of travertine, is found near waterfalls, coquina is a poorly consolidated limestone composed of pieces of coral or shells. During regional metamorphism that occurs during the building process, limestone recrystallizes into marble
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Ontario
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Ontario, one of the 13 provinces and territories of Canada, is located in east-central Canada. It is Canadas most populous province by a margin, accounting for nearly 40 percent of all Canadians. Ontario is fourth-largest in total area when the territories of the Northwest Territories and it is home to the nations capital city, Ottawa, and the nations most populous city, Toronto. There is only about 1 km of land made up of portages including Height of Land Portage on the Minnesota border. Ontario is sometimes divided into two regions, Northern Ontario and Southern Ontario. The great majority of Ontarios population and arable land is located in the south, in contrast, the larger, northern part of Ontario is sparsely populated with cold winters and is heavily forested. The province is named after Lake Ontario, a thought to be derived from Ontarí, io, a Huron word meaning great lake, or possibly skanadario. Ontario has about 250,000 freshwater lakes, the province consists of three main geographical regions, The thinly populated Canadian Shield in the northwestern and central portions, which comprises over half the land area of Ontario. Although this area mostly does not support agriculture, it is rich in minerals and in part covered by the Central and Midwestern Canadian Shield forests, studded with lakes, Northern Ontario is subdivided into two sub-regions, Northwestern Ontario and Northeastern Ontario. The virtually unpopulated Hudson Bay Lowlands in the north and northeast, mainly swampy. Southern Ontario which is further sub-divided into four regions, Central Ontario, Eastern Ontario, Golden Horseshoe, the highest point is Ishpatina Ridge at 693 metres above sea level located in Temagami, Northeastern Ontario. In the south, elevations of over 500 m are surpassed near Collingwood, above the Blue Mountains in the Dundalk Highlands, the Carolinian forest zone covers most of the southwestern region of the province. A well-known geographic feature is Niagara Falls, part of the Niagara Escarpment, the Saint Lawrence Seaway allows navigation to and from the Atlantic Ocean as far inland as Thunder Bay in Northwestern Ontario. Northern Ontario occupies roughly 87 percent of the area of the province. Point Pelee is a peninsula of Lake Erie in southwestern Ontario that is the southernmost extent of Canadas mainland, Pelee Island and Middle Island in Lake Erie extend slightly farther. All are south of 42°N – slightly farther south than the border of California. The climate of Ontario varies by season and location, the effects of these major air masses on temperature and precipitation depend mainly on latitude, proximity to major bodies of water and to a small extent, terrain relief. In general, most of Ontarios climate is classified as humid continental, Ontario has three main climatic regions
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Pioneer species
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Some lichens grow on rocks without soil, so may be among the first of life forms, and break down the rock into soil for other plants. Clear water - algae, mosses, freshwater eel grass, soil fauna, ranging from microscopic protists to larger invertebrates, have a role in soil formation and nutrient cycling. Bacteria and fungi are the most important groups in the breakdown of organic detritus left by primary producing plants such as soil, moss. Soil invertebrates enhance fungal activity by breaking down detritus, as soil develops, earthworms and ants alter soil characteristics. Worm burrows aerate soil and ant hills alter sediment particle size dispersal, though vertebrates in general would not be considered pioneer species, there are exceptions. Natterjack toads are specialists in open, sparsely vegetated habitats which may be at an early seral stage, wide-ranging generalists visit early succession stage habitats, but are not obligate species of those habitats because they use a mosaic of different habitats. Vertebrates can effect early seral stages, fossorial mammals could alter soil and plant community development. In a profound example, a seabird colony transfers considerable nitrogen into infertile soils, a keystone species may facilitate the introduction of pioneer species by creating new niches. For example, beavers may flood an area, allowing new species to immigrate, common examples of the plants in such areas include, Birch - Betula spp. Graminoids, forbs, and wildflowers - native, introduced, and invasive species, such as fire dependent seed, cone, and resprouter chaparral genera
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The Maritimes
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The Maritimes, also called the Maritime provinces or the Canadian Maritimes, is a region of Eastern Canada consisting of three provinces, New Brunswick, Nova Scotia, and Prince Edward Island. The Maritimes had a population of 1,813,606 in 2016, the Maritimes, along with a fourth province – Canadas easternmost province, Newfoundland and Labrador – make up Atlantic Canada. Located along the Atlantic coast, various aquatic sub-basins are located in the Maritimes, such as the Gulf of Maine, the region is located northeast of New England, southeast of Quebecs Gaspé Peninsula, and southwest of the island of Newfoundland. All three provinces are entirely south of the southernmost extremity of Western Canada, and are the provinces of Canada without large. The Mikmaq, Maliseet and Passamaquoddy people are indigenous to the Maritimes, while Acadian, the word maritime is an adjective that simply means of the sea, thus any land associated with the sea can be considered a maritime state or province. Nonetheless, the term Maritimes has historically been applied to New Brunswick, Nova Scotia and Prince Edward Island. The Middle Period, starting 6,000 years ago, and this is also when what is called the Laurentian tradition started among Archaic Indians, existing First Nations peoples of the time. Evidence of Archaic Indian burial mounds and other ceremonial sites existing in the Saint John River valley has been uncovered, the primarily agrarian Maliseet Nation settled throughout the Saint John River and Allagash River valleys of present-day New Brunswick and Maine. The Passamaquoddy Nation inhabited the coastal regions of the present-day Bay of Fundy. The Maritimes were the area in Canada to be settled by Europeans. Both Giovanni Caboto and Giovanni da Verrazzano are reported to have sailed in or near Maritime waters during their voyages of discovery for England, several Portuguese explorers/cartographers have also documented various parts of the Maritimes, namely Diogo Homem. However, it was French explorer Jacques Cartier who made the first detailed reconnaissance of the region for a European power, and in so doing, claimed the region for the King of France. Champlain went on to fame as the founder of New Frances province of Canada which comprises much of the present-day lower St. Lawrence River valley in the province of Quebec. Most Acadian fishing activities were overshadowed by the comparatively enormous seasonal European fishing fleets based out of Newfoundland which took advantage of proximity to the Grand Banks. In 1613, Virginian raiders captured Port-Royal, and in 1621 Acadia was ceded to Scotlands Sir William Alexander who renamed it Nova Scotia. By 1632, Acadia was returned from Scotland to France under the Treaty of Saint-Germain-en-Laye, colonial administration by France throughout the history of Acadia was contemptuous at best. Frances priorities were in settling and strengthening its claim on New France and the exploration and settlement of interior North America, over 74 years there were six colonial wars, which involved continuous warfare between New England and Acadia. Throughout these wars, New England was allied with the Iroquois Confederacy, in the first war, King Williams War, natives from the Maritime region participated in numerous attacks with the French on the Acadia/ New England border in southern Maine
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Society of American Foresters
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The Society of American Foresters is a scientific and educational 501 non-profit organization, representing the forestry profession in the United States of America. Its headquarters are located in Bethesda, Maryland, the SAF was founded in 1900 by Gifford Pinchot, the first Chief of the United States Forest Service. Accreditation is ongoing, with a review of all universities occurring every ten years,2007 being the latest. Currently, only Michigan State University, Mississippi State University, additionally, it reviews two-year programs leading to an Associate degree in forest technology and other related fields, full accreditation of technical programs will commence as of August,2009. The SAF also offers a continuing education credit system, in hopes of keeping foresters, the SAF produces five periodic journals, which are all archived through the members-only, Roots of Forestry. The Journal of Forestry and Forest Science are the most comprehensive journals of general scholarly forestry research, additionally, SAF also publishes a variety of technical publications. The Forestry Source is published monthly, and the newsletter, E-Forester. Both provide a blend of articles on current events, applied science and technology, and social happenings
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Abies balsamea
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Abies balsamea or balsam fir is a North American fir, native to most of eastern and central Canada and the northeastern United States. Balsam fir is a small to medium-size evergreen tree typically 14–20 metres tall, rarely to 27 metres tall, the bark on young trees is smooth, grey, and with resin blisters, becoming rough and fissured or scaly on old trees. The leaves are flat and needle-like,15 to 30 mm long, dark green above often with a patch of stomata near the tip. They are arranged spirally on the shoot, but with the leaf bases twisted so that the appear to be in two more-or-less horizontal rows on either side of the shoot. The seed cones are erect,40 to 80 mm long, dark purple, ripening brown, there are two varieties, Abies balsamea var. balsamea – bracts subtending seed scales short, not visible on the closed cones. Abies balsamea var. phanerolepis – bracts subtending seed scales longer, the southeast of the species range, from southernmost Quebec to West Virginia. The name Canaan fir derives from one of its native localities, some botanists regard this variety as a natural hybrid between balsam fir and Fraser fir, which occurs further south in the Appalachian mountains. On mountain tops, stands of balsam fir occasionally develop fir waves, often found in association with black spruce, white spruce and trembling aspen. The needles are eaten by some caterpillars, for example the Io moth. Both varieties of the species are popular as Christmas trees. The wood is milled for framing lumber, siding and pulped for paper manufacture, balsam fir oil is an EPA approved nontoxic rodent repellent. The balsam fir is used as an air freshener and as incense. Prior to the availability of foam rubber and air mattresses, balsam fir boughs were a preferred mattress in places where trees greatly outnumbered campers, many fir limbs are vertically bowed from alternating periods of downward deformation from snow loading and new growth reaching upward for sunlight. Layers of inverted freshly cut limbs from small trees created a pleasantly fragrant mattress lifting bedding off the wet ground, upper layers of limbs were placed with the cut ends of the limbs touching the earth to avoid uncomfortably sharp spots and sap. The cultivar A. balsamea Hudsonia group, has gained the Royal Horticultural Societys Award of Garden Merit, the Native Americans used it for a variety of medicinal purposes. Balsam fir is the tree of New Brunswick. Balsam of Peru Hunt, Richard S. Abies balsamea, IUCN Red List of Threatened Species. International Union for Conservation of Nature, gymnosperm Database, Abies balsamea Karen Legasy, Shayna LaBelle-Beadman & Brenda Chambers
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Populus tremuloides
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Populus tremuloides is a deciduous tree native to cooler areas of North America, one of several species referred to by the common name aspen. It is commonly called quaking aspen, trembling aspen, American aspen, Quakies, mountain or golden aspen, trembling poplar, white poplar, popple, the trees have tall trunks, up to 25 meters tall, with smooth pale bark, scarred with black. The glossy green leaves, dull beneath, become golden to yellow, rarely red, the species often propagates through its roots to form large groves originating from a shared system of rhizomes. Populus tremuloides is the most widely distributed tree in North America and it is the defining species of the aspen parkland biome in the Prairie Provinces of Canada and extreme northwest Minnesota. The quaking or trembling of the leaves that is referred to in the names is due to the flexible flattened petioles. The specific epithet, tremuloides, evokes this trembling behavior and can be translated as like tremula. Some species of Populus have petioles flattened partially along their length, while the aspens and some other poplars have them flattened from side to side along the entire length of the petiole. Quaking aspen is a tall, fast growing tree, usually 20–25 m at maturity, with a trunk 20 to 80 cm in diameter, the bark is relatively smooth, colored greenish-white to gray, and is marked by thick black horizontal scars and prominent black knots. Parallel vertical scars are tell-tale signs of elk, which strip off bark with their front teeth. The leaves on trees are nearly round, 4–8 centimeters in diameter with small rounded teeth. Young trees and root sprouts have much larger nearly triangular leaves, aspens are dioecious, with separate male and female clones. The quaking aspen is the State Tree of Utah, quaking aspen occurs across Canada in all provinces and territories, with the possible exception of Nunavut. It occurs at low elevations as far south as northern Nebraska, in the western United States, this tree rarely survives at elevations lower than 1,500 feet due to hot summers experienced below that elevation, and is generally found at 5, 000–12,000 feet. It grows at altitudes as far south as Guanajuato, Mexico. Quaking aspen grows in a variety of climatic conditions. January and July average temperatures range from −30 °C and 16 °C in the Alaska Interior to −3 °C and 23 °C in Fort Wayne, average annual precipitation ranges from 1,020 mm in Gander, Newfoundland and Labrador to as little as 180 mm in the Alaska Interior. The southern limit of the range roughly follows the 24 °C mean July isotherm. Shrub-like dwarf clones exist in environments too cold and dry to be hospitable to full-size trees
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