The Nematocera are a suborder of elongated flies with thin, segmented antennae and mostly aquatic larvae. This group is paraphyletic and contains all flies but species from suborder Brachycera, which includes more commonly known species such as the housefly or the common fruit fly. The equivalent clade to Nematocera is the whole Diptera, with Brachycera as a subclade. Families in Nematocera include mosquitoes, crane flies, gnats, black flies, and multiple families commonly known as midges. The Nematocera typically have fairly long, fine, finely-jointed antennae. In many species, such as most mosquitoes, the female antennae are more or less threadlike, but the males have spectacularly plumose antennae.
Nematocera
Flies are insects of the order Diptera, the name being derived from the Greek δι- di- "two", and πτερόν pteron "wing". Insects of this order use only a single pair of wings to fly, the hindwings having evolved into advanced mechanosensory organs known as halteres, which act as high-speed sensors of rotational movement and allow dipterans to perform advanced aerobatics. Diptera is a large order containing an estimated 1,000,000 species including horse-flies, crane flies, hoverflies, mosquitoes and others, although only about 125,000 species have been described.
Fly
An Anthomyiidae species showing characteristic dipteran features: large eyes, small antennae, sucking mouthparts, single pair of flying wings, hindwings reduced to clublike halteres
Fossil brachyceran in Baltic amber. Lower Eocene, c. 50 million years ago
Fossil nematoceran in Dominican amber. Sandfly, Lutzomyia adiketis (Psychodidae), Early Miocene, c. 20 million years ago