1.
Onager (weapon)
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The onager was a Roman siege engine that was a type of catapult that uses a torsional force, generally from twisted rope, to store energy for the shot. The onager consisted of a frame placed on the ground to whose front end a vertical frame of solid timber was rigidly fixed. A vertical spoke that passed through a rope bundle fastened to the frame had a sling attached which contained a projectile. To fire it, the spoke or arm was forced down, against the tension of twisted ropes or other springs, by a windlass, as the sling swung outwards, one end would release, as with a staff-sling, and the projectile would be hurled forward. The arm would then be caught by a beam or bed. The onagers framework is made out of two beams from oak, which curve into humps, in the middle they have quite large holes in them, in which strong sinew ropes are stretched and twisted. A long arm is inserted between the bundle of rope, at its end it has a pin and a pouch. It strikes on a huge buffer with a sack stuffed with fine chaff, when it comes to combat, a round stone is put in the pouch and the arm is winched down. Then, the master artilleryman strikes the pin with a hammer, and with a big blow, the onagers of the Roman Empire were mainly used for besieging forts or settlements. They would often be armed with stones that could be covered with a combustible substance. They could be outranged by bows, in the Middle Ages a less powerful version of the onager was used that held the projectile in a fixed bowl instead of a sling. Some have speculated that contemporary drawings depicting fixed bowls were intentionally misleading, such deception would occur because, at the time, catapults were state of the art weapons of war and their exact operation and construction would be a closely guarded secret. It derived its name from the action of the machine. This engine was called the mangonel, although that name may have also been used for a variety of siege engines
2.
Megaannum
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A year is the orbital period of the Earth moving in its orbit around the Sun. Due to the Earths axial tilt, the course of a year sees the passing of the seasons, marked by changes in weather, the hours of daylight, and, consequently, vegetation and soil fertility. In temperate and subpolar regions around the globe, four seasons are recognized, spring, summer, autumn. In tropical and subtropical regions several geographical sectors do not present defined seasons, but in the seasonal tropics, a calendar year is an approximation of the number of days of the Earths orbital period as counted in a given calendar. The Gregorian, or modern, calendar, presents its calendar year to be either a common year of 365 days or a year of 366 days, as do the Julian calendars. For the Gregorian calendar the average length of the year across the complete leap cycle of 400 years is 365.2425 days. The ISO standard ISO 80000-3, Annex C, supports the symbol a to represent a year of either 365 or 366 days, in English, the abbreviations y and yr are commonly used. In astronomy, the Julian year is a unit of time, it is defined as 365.25 days of exactly 86400 seconds, totalling exactly 31557600 seconds in the Julian astronomical year. The word year is used for periods loosely associated with, but not identical to, the calendar or astronomical year, such as the seasonal year, the fiscal year. Similarly, year can mean the period of any planet, for example. The term can also be used in reference to any long period or cycle, west Saxon ġēar, Anglian ġēr continues Proto-Germanic *jǣran. Cognates are German Jahr, Old High German jār, Old Norse ár and Gothic jer, all the descendants of the Proto-Indo-European noun *yeh₁rom year, season. Cognates also descended from the same Proto-Indo-European noun are Avestan yārǝ year, Greek ὥρα year, season, period of time, Old Church Slavonic jarŭ, Latin annus is from a PIE noun *h₂et-no-, which also yielded Gothic aþn year. Both *yeh₁-ro- and *h₂et-no- are based on verbal roots expressing movement, *h₁ey- and *h₂et- respectively, the Greek word for year, ἔτος, is cognate with Latin vetus old, from the PIE word *wetos- year, also preserved in this meaning in Sanskrit vat-sa- yearling and vat-sa-ras year. Derived from Latin annus are a number of English words, such as annual, annuity, anniversary, etc. per annum means each year, anno Domini means in the year of the Lord. No astronomical year has an number of days or lunar months. Financial and scientific calculations often use a 365-day calendar to simplify daily rates, in the Julian calendar, the average length of a year is 365.25 days. In a non-leap year, there are 365 days, in a year there are 366 days
3.
Precambrian
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The Precambrian is the earliest period of Earths history, set before the current Phanerozoic Eon. The Precambrian is a supereon that is subdivided into three eons of the time scale. It spans from the formation of Earth about 4.6 billion years ago to the beginning of the Cambrian Period, about 541 million years ago, the Precambrian accounts for 89% of geologic time. Relatively little is known about the Precambrian, despite it making up roughly seven-eighths of the Earths history, the Precambrian fossil record is poorer than that of the succeeding Phanerozoic, and fossils from that time are of limited biostratigraphic use. This is because many Precambrian rocks have been metamorphosed, obscuring their origins, while others have been destroyed by erosion. A stable crust was apparently in place by 4,412 Ma, the term Precambrian is recognized by the International Commission on Stratigraphy as a general term including the Archean and Proterozoic eons. It is still used by geologists and paleontologists for general discussions not requiring the more specific eon names and it was briefly called the Cryptozoic eon. A specific date for the origin of life has not been determined, carbon found in 3.8 billion year old rocks from islands off western Greenland may be of organic origin. Well-preserved microscopic fossils of bacteria older than 3.46 billion years have found in Western Australia. Probable fossils 100 million years older have been found in the same area, there is a fairly solid record of bacterial life throughout the remainder of the Precambrian. The oldest fossil evidence from that era of such complex life comes from the Lantian formation of the Ediacarian period, a very diverse collection of soft-bodied forms is found in a variety of locations worldwide and date to between 635 and 542 Ma. These are referred to as Ediacaran or Vendian biota, hard-shelled creatures appeared toward the end of that time span, marking the beginning of the Phanerozoic era. By the middle of the following Cambrian period, a diverse fauna is recorded in the Burgess Shale. The explosion in diversity of lifeforms during the early Cambrian is called the Cambrian explosion of life, while land seems to have been devoid of plants and animals, cyanobacteria and other microbes formed prokaryotic mats that covered terrestrial areas. Evidence of the details of plate motions and other activity in the Precambrian has been poorly preserved. It is generally believed that small proto-continents existed prior to 4280 Ma, the supercontinent, known as Rodinia, broke up around 750 Ma. A number of glacial periods have been identified going as far back as the Huronian epoch, one of the best studied is the Sturtian-Varangian glaciation, around 850–635 Ma, which may have brought glacial conditions all the way to the equator, resulting in a Snowball Earth. The atmosphere of the early Earth is not well understood, most geologists believe it was composed primarily of nitrogen, carbon dioxide, and other relatively inert gases, and was lacking in free oxygen
4.
Cambrian
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The Cambrian Period was the first geological period of the Paleozoic Era, of the Phanerozoic Eon. The Cambrian lasted 55.6 million years from the end of the preceding Ediacaran Period 541 million years ago to the beginning of the Ordovician Period 485.4 mya and its subdivisions, and its base, are somewhat in flux. The period was established by Adam Sedgwick, who named it after Cambria, the Latinised form of Cymru, the Welsh name for Wales, as a result, our understanding of the Cambrian biology surpasses that of some later periods. The rapid diversification of lifeforms in the Cambrian, known as the Cambrian explosion, most of the continents were probably dry and rocky due to a lack of vegetation. Shallow seas flanked the margins of several continents created during the breakup of the supercontinent Pannotia, the seas were relatively warm, and polar ice was absent for much of the period. The United States Federal Geographic Data Committee uses a barred capital C ⟨Є⟩ character similar to the capital letter Ukrainian Ye ⟨Є⟩ to represent the Cambrian Period, the proper Unicode character is U+A792 Ꞓ LATIN CAPITAL LETTER C WITH BAR. Despite the long recognition of its distinction from younger Ordovician Period rocks and older Supereon Precambrian rocks, the base of the Cambrian lies atop a complex assemblage of trace fossils known as the Treptichnus pedum assemblage. Pedum in Namibia, Spain and Newfoundland, and possibly, in the western USA, the stratigraphic range of T. pedum overlaps the range of the Ediacaran fossils in Namibia, and probably in Spain. The Cambrian Period followed the Ediacaran Period and was followed by the Ordovician Period, the Cambrian is divided into four epochs and ten ages. Currently only two series and five stages are named and have a GSSP, because the international stratigraphic subdivision is not yet complete, many local subdivisions are still widely used. In some of these subdivisions the Cambrian is divided into three epochs with locally differing names – the Early Cambrian, Middle Cambrian and Furongian, rocks of these epochs are referred to as belonging to the Lower, Middle, or Upper Cambrian. Trilobite zones allow biostratigraphic correlation in the Cambrian, each of the local epochs is divided into several stages. The International Commission on Stratigraphy list the Cambrian period as beginning at 541 million years ago, the lower boundary of the Cambrian was originally held to represent the first appearance of complex life, represented by trilobites. The recognition of small shelly fossils before the first trilobites, and Ediacara biota substantially earlier and this formal designation allowed radiometric dates to be obtained from samples across the globe that corresponded to the base of the Cambrian. Early dates of 570 million years ago quickly gained favour, though the used to obtain this number are now considered to be unsuitable. A more precise date using modern radiometric dating yield a date of 541 ±0.3 million years ago, most continental land was clustered in the Southern Hemisphere at this time, but was drifting north. Large, high-velocity rotational movement of Gondwana appears to have occurred in the Early Cambrian, the sea levels fluctuated somewhat, suggesting there were ice ages, associated with pulses of expansion and contraction of a south polar ice cap. In Baltoscandia a Lower Cambrian transgression transformed large swathes of the Sub-Cambrian peneplain into a epicontinental sea, the Earth was generally cold during the early Cambrian, probably due to the ancient continent of Gondwana covering the South Pole and cutting off polar ocean currents
5.
Ordovician
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The Ordovician is a geologic period and system, the second of six periods of the Paleozoic Era. The Ordovician spans 41.2 million years from the end of the Cambrian Period 485.4 million years ago to the start of the Silurian Period 443.8 Mya. Lapworth recognized that the fauna in the disputed strata were different from those of either the Cambrian or the Silurian periods. It received international sanction in 1960, when it was adopted as a period of the Paleozoic Era by the International Geological Congress. Life continued to flourish during the Ordovician as it did in the earlier Cambrian period, invertebrates, namely molluscs and arthropods, dominated the oceans. The Great Ordovician Biodiversification Event considerably increased the diversity of life, fish, the worlds first true vertebrates, continued to evolve, and those with jaws may have first appeared late in the period. Life had yet to diversify on land, about 100 times as many meteorites struck the Earth during the Ordovician compared with today. The Ordovician Period began with a major extinction called the Cambrian–Ordovician extinction event and it lasted for about 42 million years and ended with the Ordovician–Silurian extinction event, about 443.8 Mya which wiped out 60% of marine genera. The dates given are recent radiometric dates and vary slightly from those found in other sources and this second period of the Paleozoic era created abundant fossils that became major petroleum and gas reservoirs. The boundary chosen for the beginning of both the Ordovician Period and the Tremadocian stage is highly significant and it correlates well with the occurrence of widespread graptolite, conodont, and trilobite species. The base of the Tremadocian allows scientists to relate these species not only to each other and this makes it easier to place many more species in time relative to the beginning of the Ordovician Period. A number of terms have been used to subdivide the Ordovician Period. In 2008, the ICS erected an international system of subdivisions. There exist Baltoscandic, British, Siberian, North American, Australian, the Ordovician Period in Britain was traditionally broken into Early, Middle and Late epochs. The corresponding rocks of the Ordovician System are referred to as coming from the Lower, Middle, the Floian corresponds to the lower Arenig, the Arenig continues until the early Darriwilian, subsuming the Dapingian. The Llanvirn occupies the rest of the Darriwilian, and terminates with it at the base of the Late Ordovician. The Sandbian represents the first half of the Caradoc, the Caradoc ends in the mid-Katian, during the Ordovician, the southern continents were collected into Gondwana. Gondwana started the period in equatorial latitudes and, as the period progressed, drifted toward the South Pole, the small continent Avalonia separated from Gondwana and began to move north towards Baltica and Laurentia, opening the Rheic Ocean between Gondwana and Avalonia
6.
Silurian
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The Silurian is a geologic period and system spanning 24.6 million years from the end of the Ordovician Period, at 443.8 million years ago, to the beginning of the Devonian Period,419.2 Mya. As with other periods, the rock beds that define the periods start and end are well identified. The base of the Silurian is set at a major Ordovician-Silurian extinction event when 60% of marine species were wiped out, a significant evolutionary milestone during the Silurian was the diversification of jawed and bony fish. However, terrestrial life would not greatly diversify and affect the landscape until the Devonian, the Silurian system was first identified by British geologist Sir Roderick Impey Murchison, who was examining fossil-bearing sedimentary rock strata in south Wales in the early 1830s. He named the sequences for a Celtic tribe of Wales, the Silures, inspired by his friend Adam Sedgwick and this naming does not indicate any correlation between the occurrence of the Silurian rocks and the land inhabited by the Silures. As it was first identified, the Silurian series when traced farther afield quickly came to overlap Sedgwicks Cambrian sequence, however, charles Lapworth resolved the conflict by defining a new Ordovician system including the contested beds. An early alternative name for the Silurian was Gotlandian after the strata of the Baltic island of Gotland, the French geologist Joachim Barrande, building on Murchisons work, used the term Silurian in a more comprehensive sense than was justified by subsequent knowledge. He divided the Silurian rocks of Bohemia into eight stages and his interpretation was questioned in 1854 by Edward Forbes, and the later stages of Barrande, F, G and H, have since been shown to be Devonian. Despite these modifications in the groupings of the strata, it is recognized that Barrande established Bohemia as a classic ground for the study of the earliest fossils. The epoch is named for the town of Llandovery in Carmarthenshire, the Wenlock, which lasted from 433.4 ±1.5 to 427.4 ±2.8 mya, is subdivided into the Sheinwoodian and Homerian ages. It is named after Wenlock Edge in Shropshire, England, during the Wenlock, the oldest known tracheophytes of the genus Cooksonia, appear. The first terrestrial animals also appear in the Wenlock, represented by air-breathing millipedes from Scotland. The Ludlow, lasting from 427.4 ±1.5 to 423 ±2.8 mya, comprises the Gorstian stage, lasting until 425.6 million years ago, and it is named for the town of Ludlow in Shropshire, England. The Pridoli, lasting from 423 ±1.5 to 419.2 ±2.8 mya, is the final and it is named after one locality at the Homolka a Přídolí nature reserve near the Prague suburb Slivenec in the Czech Republic. Přídolí is the old name of a field area. The high sea levels of the Silurian and the flat land resulted in a number of island chains. The southern continents remained united during this period, the melting of icecaps and glaciers contributed to a rise in sea level, recognizable from the fact that Silurian sediments overlie eroded Ordovician sediments, forming an unconformity. The continents of Avalonia, Baltica, and Laurentia drifted together near the equator and this event is the Caledonian orogeny, a spate of mountain building that stretched from New York State through conjoined Europe and Greenland to Norway
7.
Devonian
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The Devonian is a geologic period and system of the Paleozoic, spanning 60 million years from the end of the Silurian,419.2 million years ago, to the beginning of the Carboniferous,358.9 Mya. It is named after Devon, England, where rocks from this period were first studied, the first significant adaptive radiation of life on dry land occurred during the Devonian. Free-sporing vascular plants began to spread across dry land, forming extensive forests which covered the continents, by the middle of the Devonian, several groups of plants had evolved leaves and true roots, and by the end of the period the first seed-bearing plants appeared. Various terrestrial arthropods also became well-established, Fish reached substantial diversity during this time, leading the Devonian to often be dubbed the Age of Fish. The first ray-finned and lobe-finned bony fish appeared, while the placodermi began dominating almost every aquatic environment. The ancestors of all four-limbed vertebrates began adapting to walking on land, as their strong pectoral, in the oceans, primitive sharks became more numerous than in the Silurian and Late Ordovician. The first ammonites, species of molluscs, appeared, trilobites, the mollusk-like brachiopods and the great coral reefs, were still common. The Late Devonian extinction which started about 375 million years ago severely affected marine life, killing off all placodermi, and all trilobites, save for a few species of the order Proetida. The palaeogeography was dominated by the supercontinent of Gondwana to the south, the continent of Siberia to the north, while the rock beds that define the start and end of the Devonian period are well identified, the exact dates are uncertain. According to the International Commission on Stratigraphy, the Devonian extends from the end of the Silurian 419.2 Mya, another common term is Age of the Fishes, referring to the evolution of several major groups of fish that took place during the period. Older literature on the Anglo-Welsh basin divides it into the Downtonian, Dittonian, Breconian and Farlovian stages, in the Late Devonian, by contrast, arid conditions were less prevalent across the world and temperate climates were more common. The Devonian Period is formally broken into Early, Middle and Late subdivisions, the rocks corresponding to those epochs are referred to as belonging to the Lower, Middle and Upper parts of the Devonian System. Early Devonian The Early Devonian lasted from 419.2 ±2.8 to 393.3 ±2.5 and began with the Lochkovian stage, which lasted until the Pragian. It spanned from 410.8 ±2.8 to 407.6 ±2.5, and was followed by the Emsian, which lasted until the Middle Devonian began,393. 3±2.7 million years ago. Middle Devonian The Middle Devonian comprised two subdivisions, first the Eifelian, which gave way to the Givetian 387. 7±2.7 million years ago. Late Devonian Finally, the Late Devonian started with the Frasnian,382.7 ±2.8 to 372.2 ±2.5, during which the first forests took shape on land. The first tetrapods appeared in the record in the ensuing Famennian subdivision. This lasted until the end of the Devonian,358. 9±2.5 million years ago, the Devonian was a relatively warm period, and probably lacked any glaciers
8.
Carboniferous
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The Carboniferous is a geologic period and system that spans 60 million years from the end of the Devonian Period 358.9 million years ago, to the beginning of the Permian Period,298.9 Mya. The name Carboniferous means coal-bearing and derives from the Latin words carbō and ferō, and was coined by geologists William Conybeare and William Phillips in 1822. Based on a study of the British rock succession, it was the first of the system names to be employed. The Carboniferous is often treated in North America as two periods, the earlier Mississippian and the later Pennsylvanian. Terrestrial life was established by the Carboniferous period. Amphibians were the dominant land vertebrates, of one branch would eventually evolve into amniotes. Arthropods were also common, and many were much larger than those of today. Vast swaths of forest covered the land, which would eventually be laid down, the atmospheric content of oxygen also reached their highest levels in geological history during the period, 35% compared with 21% today, allowing terrestrial invertebrates to evolve to great size. A major marine and terrestrial extinction event, the Carboniferous rainforest collapse, occurred in the middle of the period, the later half of the period experienced glaciations, low sea level, and mountain building as the continents collided to form Pangaea. In the United States the Carboniferous is usually broken into Mississippian and Pennsylvanian subperiods, the Silesian is roughly contemporaneous with the late Mississippian Serpukhovian plus the Pennsylvanian. In Britain the Dinantian is traditionally known as the Carboniferous Limestone, the Namurian as the Millstone Grit, and the Westphalian as the Coal Measures and Pennant Sandstone. There was also a drop in south polar temperatures, southern Gondwanaland was glaciated throughout the period and these conditions apparently had little effect in the deep tropics, where lush swamps, later to become coal, flourished to within 30 degrees of the northernmost glaciers. Mid-Carboniferous, a drop in sea level precipitated a major extinction, one that hit crinoids. This sea level drop and the unconformity in North America separate the Mississippian subperiod from the Pennsylvanian subperiod. This happened about 323 million years ago, at the onset of the Permo-Carboniferous Glaciation, the Carboniferous was a time of active mountain-building, as the supercontinent Pangaea came together. The southern continents remained tied together in the supercontinent Gondwana, which collided with North America–Europe along the present line of eastern North America, in the same time frame, much of present eastern Eurasian plate welded itself to Europe along the line of the Ural mountains. Most of the Mesozoic supercontinent of Pangea was now assembled, although North China, the Late Carboniferous Pangaea was shaped like an O. There were two major oceans in the Carboniferous—Panthalassa and Paleo-Tethys, which was inside the O in the Carboniferous Pangaea, other minor oceans were shrinking and eventually closed - Rheic Ocean, the small, shallow Ural Ocean and Proto-Tethys Ocean
9.
Permian
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The Permian is a geologic period and system which spans 46.7 million years from the end of the Carboniferous Period 298.9 million years ago, to the beginning of the Triassic Period 252.2 Mya. It is the last period of the Paleozoic Era, the following Triassic Period belongs to the Mesozoic Era, the concept of the Permian was introduced in 1841 by geologist Sir Roderick Murchison, who named it after the city of Perm. The Permian witnessed the diversification of the early amniotes into the groups of the mammals, turtles, lepidosaurs. The world at the time was dominated by two known as Pangaea and Siberia, surrounded by a global ocean called Panthalassa. The Carboniferous rainforest collapse left behind vast regions of desert within the continental interior, amniotes, who could better cope with these drier conditions, rose to dominance in place of their amphibian ancestors. The Permian ended with the Permian–Triassic extinction event, the largest mass extinction in Earths history, in which nearly 90% of marine species and it would take well into the Triassic for life to recover from this catastrophe. Recovery from the Permian-Triassic extinction event was protracted, on land, the term Permian was introduced into geology in 1841 by Sir R. I. Murchison, president of the Geological Society of London, who identified typical strata in extensive Russian explorations undertaken with Edouard de Verneuil, the region now lies in the Perm Krai of Russia. This could have in part caused the extinctions of marine species at the end of the period by severely reducing shallow coastal areas preferred by many marine organisms. During the Permian, all the Earths major landmasses were collected into a supercontinent known as Pangaea. The Cimmeria continent rifted away from Gondwana and drifted north to Laurasia, a new ocean was growing on its southern end, the Tethys Ocean, an ocean that would dominate much of the Mesozoic Era. Large continental landmass interiors experience climates with extreme variations of heat and cold, deserts seem to have been widespread on Pangaea. Such dry conditions favored gymnosperms, plants with seeds enclosed in a cover, over plants such as ferns that disperse spores in a wetter environment. The first modern trees appeared in the Permian, the climate in the Permian was quite varied. At the start of the Permian, the Earth was still in an Ice Age, glaciers receded around the mid-Permian period as the climate gradually warmed, drying the continents interiors. In the late Permian period, the drying continued although the temperature cycled between warm and cool cycles, Permian marine deposits are rich in fossil mollusks, echinoderms, and brachiopods. By the close of the Permian, trilobites and a host of other groups became extinct. Terrestrial life in the Permian included diverse plants, fungi, arthropods, the period saw a massive desert covering the interior of Pangaea
10.
Triassic
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The Triassic is a geologic period and system which spans 50.9 million years from the end of the Permian Period 252.17 million years ago, to the beginning of the Jurassic Period 201.3 Mya. The Triassic is the first period of the Mesozoic Era, both the start and end of the period are marked by major extinction events. Therapsids and archosaurs were the terrestrial vertebrates during this time. A specialized subgroup of archosaurs, called dinosaurs, first appeared in the Late Triassic, the vast supercontinent of Pangaea existed until the mid-Triassic, after which it began to gradually rift into two separate landmasses, Laurasia to the north and Gondwana to the south. The global climate during the Triassic was mostly hot and dry, however, the climate shifted and became more humid as Pangaea began to drift apart. The end of the period was marked by yet another mass extinction, the Triassic-Jurassic extinction event. The Triassic is usually separated into Early, Middle, and Late Triassic Epochs, from the east, along the equator, the Tethys sea penetrated Pangaea, causing the Paleo-Tethys Ocean to be closed. Later in the mid-Triassic a similar sea penetrated along the equator from the west, the remaining shores were surrounded by the world-ocean known as Panthalassa. All the deep-ocean sediments laid down during the Triassic have disappeared through subduction of oceanic plates, thus, the supercontinent Pangaea was rifting during the Triassic—especially late in that period—but had not yet separated. In North America, for example, marine deposits are limited to a few exposures in the west, thus Triassic stratigraphy is mostly based on organisms that lived in lagoons and hypersaline environments, such as Estheria crustaceans. At the beginning of the Mesozoic Era, Africa was joined with Earths other continents in Pangaea, Africa shared the supercontinents relatively uniform fauna which was dominated by theropods, prosauropods and primitive ornithischians by the close of the Triassic period. Late Triassic fossils are found throughout Africa, but are common in the south than north. The time boundary separating the Permian and Triassic marks the advent of an event with global impact. At Paleorrota geopark, located in Rio Grande do Sul, Brazil, in these formations, one of the earliest dinosaurs, Staurikosaurus, as well as the mammal ancestors Brasilitherium and Brasilodon have been discovered. The Triassic continental interior climate was hot and dry, so that typical deposits are red bed sandstones and evaporites. Pangaeas large size limited the effect of the global ocean, its continental climate was highly seasonal, with very hot summers. The strong contrast between the Pangea supercontinent and the global ocean triggered intense cross-equatorial monsoons, the best studied of such episodes of humid climate, and probably the most intense and widespread, was the Carnian Pluvial Event. On land, the vascular plants included the lycophytes, the dominant cycadophytes, ginkgophyta, ferns, horsetails
11.
Jurassic
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The Jurassic is a geologic period and system that spans 56.3 million years from the end of the Triassic Period 201.3 million years ago to the beginning of the Cretaceous Period 145 Mya. The Jurassic constitutes the middle period of the Mesozoic Era, also known as the Age of Reptiles, the start of the period is marked by the major Triassic–Jurassic extinction event. The Jurassic is named after the Jura Mountains within the European Alps, by the beginning of the Jurassic, the supercontinent Pangaea had begun rifting into two landmasses, Laurasia to the north and Gondwana to the south. This created more coastlines and shifted the continental climate from dry to humid, on land, the fauna transitioned from the Triassic fauna, dominated by both dinosauromorph and crocodylomorph archosaurs, to one dominated by dinosaurs alone. The first birds also appeared during the Jurassic, having evolved from a branch of theropod dinosaurs, other major events include the appearance of the earliest lizards, and the evolution of therian mammals, including primitive placentals. Crocodilians made the transition from a terrestrial to a mode of life. The oceans were inhabited by marine reptiles such as ichthyosaurs and plesiosaurs, the chronostratigraphic term Jurassic is directly linked to the Jura Mountains. The name Jura is derived from the Celtic root jor, which was Latinised into juria, the Jurassic period is divided into the Early Jurassic, Middle, and Late Jurassic epochs. The Jurassic System, in stratigraphy, is divided into the Lower Jurassic, Middle, the separation of the term Jurassic into three sections goes back to Leopold von Buch. The Jurassic North Atlantic Ocean was relatively narrow, while the South Atlantic did not open until the following Cretaceous period, the Tethys Sea closed, and the Neotethys basin appeared. Climates were warm, with no evidence of glaciation, as in the Triassic, there was apparently no land over either pole, and no extensive ice caps existed. In contrast, the North American Jurassic record is the poorest of the Mesozoic, the Jurassic was a time of calcite sea geochemistry in which low-magnesium calcite was the primary inorganic marine precipitate of calcium carbonate. Carbonate hardgrounds were thus very common, along with calcitic ooids, calcitic cements, the first of several massive batholiths were emplaced in the northern American cordillera beginning in the mid-Jurassic, marking the Nevadan orogeny. Important Jurassic exposures are found in Russia, India, South America, Japan, Australasia. As the Jurassic proceeded, larger and more groups of dinosaurs like sauropods and ornithopods proliferated in Africa. Middle Jurassic strata are well represented nor well studied in Africa. Late Jurassic strata are also poorly represented apart from the spectacular Tendaguru fauna in Tanzania, the Late Jurassic life of Tendaguru is very similar to that found in western North Americas Morrison Formation. During the Jurassic period, the primary living in the sea were fish
12.
Cretaceous
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The Cretaceous is a geologic period and system that spans 79 million years from the end of the Jurassic Period 145 million years ago to the beginning of the Paleogene Period 66 Mya. It is the last period of the Mesozoic Era, the Cretaceous Period is usually abbreviated K, for its German translation Kreide. The Cretaceous was a period with a warm climate, resulting in high eustatic sea levels that created numerous shallow inland seas. These oceans and seas were populated with now-extinct marine reptiles, ammonites and rudists, during this time, new groups of mammals and birds, as well as flowering plants, appeared. The Cretaceous ended with a mass extinction, the Cretaceous–Paleogene extinction event, in which many groups, including non-avian dinosaurs, pterosaurs. The end of the Cretaceous is defined by the abrupt Cretaceous–Paleogene boundary, the name Cretaceous was derived from Latin creta, meaning chalk. The Cretaceous is divided into Early and Late Cretaceous epochs, or Lower and Upper Cretaceous series, in older literature the Cretaceous is sometimes divided into three series, Neocomian, Gallic and Senonian. A subdivision in eleven stages, all originating from European stratigraphy, is now used worldwide, in many parts of the world, alternative local subdivisions are still in use. As with other geologic periods, the rock beds of the Cretaceous are well identified. No great extinction or burst of diversity separates the Cretaceous from the Jurassic and this layer has been dated at 66.043 Ma. A140 Ma age for the Jurassic-Cretaceous boundary instead of the usually accepted 145 Ma was proposed in 2014 based on a study of Vaca Muerta Formation in Neuquén Basin. Víctor Ramos, one of the authors of the study proposing the 140 Ma boundary age sees the study as a first step toward formally changing the age in the International Union of Geological Sciences, due to the high sea level there was extensive space for such sedimentation. Because of the young age and great thickness of the system. Chalk is a type characteristic for the Cretaceous. It consists of coccoliths, microscopically small calcite skeletons of coccolithophores, the group is found in England, northern France, the low countries, northern Germany, Denmark and in the subsurface of the southern part of the North Sea. Chalk is not easily consolidated and the Chalk Group still consists of sediments in many places. The group also has other limestones and arenites, among the fossils it contains are sea urchins, belemnites, ammonites and sea reptiles such as Mosasaurus. In southern Europe, the Cretaceous is usually a marine system consisting of competent limestone beds or incompetent marls
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Paleogene
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The Paleogene is a geologic period and system that spans 43 million years from the end of the Cretaceous Period 66 million years ago to the beginning of the Neogene Period 23.03 Mya. It is the beginning of the Cenozoic Era of the present Phanerozoic Eon and this period consists of the Paleocene, Eocene and Oligocene epochs. The terms Paleogene System and lower Tertiary System are applied to the rocks deposited during the Paleogene Period. By dividing the Tertiary Period into two periods instead of directly into five epochs, the periods are more comparable to the duration of periods of the preceding Mesozoic and Paleozoic Eras. The trend was caused by the formation of the Antarctic Circumpolar Current. During the Paleogene, the continued to drift closer to their current positions. India was in the process of colliding with Asia, subsequently forming the Himalayas, the Atlantic Ocean continued to widen by a few centimeters each year. Africa was moving north to meet with Europe and form the Mediterranean, inland seas retreated from North America early in the period. Australia had also separated from Antarctica and was drifting towards Southeast Asia, mammals began a rapid diversification during this period. Some of these mammals would evolve into forms that would dominate the land, while others would become capable of living in marine, specialized terrestrial. Those that took to the oceans became modern cetaceans, while those that took to the trees became primates, the group to which humans belong. Birds, which were well established by the end of the Cretaceous. In comparison to birds and mammals, most other branches of life remained unchanged during this period. As the Earth cooled, tropical plants became less numerous and were now restricted to equatorial regions, deciduous plants, which could survive through the seasonal climates the world was now experiencing, became more common. The Paleogene is notable in the context of offshore oil drilling, and especially in Gulf of Mexico oil exploration and these rock formations represent the current cutting edge of deep-water oil discovery. Lower Tertiary explorations to date include, Kaskida Oil Field Tiber Oil Field Jack 2 Paleogene Microfossils, 180+ images of Foraminifera
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Neogene
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The Neogene is a geologic period and system that spans 20.45 million years from the end of the Paleogene Period 23.03 million years ago to the beginning of the present Quaternary Period 2.58 Mya. The Neogene is sub-divided into two epochs, the earlier Miocene and the later Pliocene, some geologists assert that the Neogene cannot be clearly delineated from the modern geological period, the Quaternary. During this period, mammals and birds continued to evolve into modern forms. Early hominids, the ancestors of humans, appeared in Africa near the end of the period, some continental movement took place, the most significant event being the connection of North and South America at the Isthmus of Panama, late in the Pliocene. This cut off the ocean currents from the Pacific to the Atlantic ocean. The global climate cooled considerably over the course of the Neogene, the terms Neogene System and upper Tertiary System describe the rocks deposited during the Neogene Period. The continents in the Neogene were very close to their current positions, the Isthmus of Panama formed, connecting North and South America. The Indian subcontinent continued to collide with Asia, forming the Himalayas, sea levels fell, creating land bridges between Africa and Eurasia and between Eurasia and North America. The global climate became seasonal and continued an overall drying and cooling trend which began at the start of the Paleogene. The ice caps on both poles began to grow and thicken, and by the end of the period the first of a series of glaciations of the current Ice Age began, marine and continental flora and fauna have a modern appearance. The reptile group Choristodera became extinct in the part of the period. Mammals and birds continued to be the dominant terrestrial vertebrates, the first hominids, the ancestors of humans, appeared in Africa and spread into Eurasia. In response to the cooler, seasonal climate, tropical plant species gave way to deciduous ones, grasses therefore greatly diversified, and herbivorous mammals evolved alongside it, creating the many grazing animals of today such as horses, antelope, and bison. The Neogene traditionally ended at the end of the Pliocene Epoch, just before the definition of the beginning of the Quaternary Period. However, there was a movement amongst geologists to also include ongoing geological time in the Neogene, by dividing the Cenozoic Era into three periods instead of seven epochs, the periods are more closely comparable to the duration of periods in the Mesozoic and Paleozoic eras. The International Commission on Stratigraphy once proposed that the Quaternary be considered a sub-era of the Neogene, with a date of 2.58 Ma. In the 2004 proposal of the ICS, the Neogene would have consisted of the Miocene and Pliocene epochs, thus the Neogene Period ends bounding the succeeding Quaternary Period at 2.58 Mya. Digital Atlas of Neogene Life for the Southeastern United States — by San Jose State University via Web Archive
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Pliocene
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The Pliocene Epoch is the epoch in the geologic timescale that extends from 5.333 million to 2.58 million years BP. It is the second and youngest epoch of the Neogene Period in the Cenozoic Era, the Pliocene follows the Miocene Epoch and is followed by the Pleistocene Epoch.588 to 1.806 million years ago, and is now included in the Pleistocene. As with other geologic periods, the geological strata that define the start and end are well identified but the exact dates of the start. The boundaries defining the Pliocene are not set at an easily identified worldwide event, the upper boundary was set at the start of the Pleistocene glaciations. The Pliocene was named by Sir Charles Lyell, the name comes from the Greek words πλεῖον and καινός and means roughly continuation of the recent, referring to the essentially modern marine mollusc faunas. H. W. Fowler called the term a regrettable barbarism, in the official timescale of the ICS, the Pliocene is subdivided into two stages. From youngest to oldest they are, Piacenzian Zanclean The Piacenzian is sometimes referred to as the Late Pliocene, in the system of North American Land Mammal Ages include Hemphillian, and Blancan. The Blancan extends forward into the Pleistocene, South American Land Mammal Ages include Montehermosan, Chapadmalalan and Uquian. In the Paratethys area the Pliocene contains the Dacian and Romanian stages, as usual in stratigraphy, there are many other regional and local subdivisions in use. In Britain the Pliocene is divided into the stages, Gedgravian, Waltonian, Pre-Ludhamian, Ludhamian, Thurnian, Bramertonian or Antian, Pre-Pastonian or Baventian, Pastonian and Beestonian. The exact correlations between these stages and the ICS stages is still a matter of detail. The formation of an Arctic ice cap is signaled by a shift in oxygen isotope ratios and ice-rafted cobbles in the North Atlantic. Mid-latitude glaciation was probably underway before the end of the epoch, the global cooling that occurred during the Pliocene may have spurred on the disappearance of forests and the spread of grasslands and savannas. Continents continued to drift, moving from positions possibly as far as 250 km from their present locations to positions only 70 km from their current locations, africas collision with Europe formed the Mediterranean Sea, cutting off the remnants of the Tethys Ocean. The border between the Miocene and the Pliocene is also the time of the Messinian salinity crisis, Sea level changes exposed the land-bridge between Alaska and Asia. Pliocene marine rocks are exposed in the Mediterranean, India. Elsewhere, they are exposed largely near shores, the change to a cooler, dry, seasonal climate had considerable impacts on Pliocene vegetation, reducing tropical species worldwide. Deciduous forests proliferated, coniferous forests and tundra covered much of the north, tropical forests were limited to a tight band around the equator, and in addition to dry savannahs, deserts appeared in Asia and Africa
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Holocene
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The Holocene is the geological epoch that began after the Pleistocene at approximately 11,700 years before present. The term Recent has often used as an exact synonym of Holocene. The Holocene is part of the Quaternary period and its name comes from the Ancient Greek words ὅλος and καινός, meaning entirely recent. It has been identified with the current warm period, known as MIS1, given these, a new term, Anthropocene, is specifically proposed and used informally only for the very latest part of modern history involving significant human impact. It is accepted by the International Commission on Stratigraphy that the Holocene started approximately 11,700 years ago, the epoch follows the Pleistocene and the last glacial period. The Holocene can be subdivided into five time intervals, or chronozones, based on climatic fluctuations, Preboreal, Boreal, Atlantic, Subboreal and they find a general correspondence across Eurasia and North America, though the method was once thought to be of no interest. The scheme was defined for Northern Europe, but the changes were claimed to occur more widely. The periods of the include a few of the final pre-Holocene oscillations of the last glacial period. Paleontologists have not defined any faunal stages for the Holocene, if subdivision is necessary, periods of human technological development, such as the Mesolithic, Neolithic, and Bronze Age, are usually used. However, the time periods referenced by these terms vary with the emergence of those technologies in different parts of the world, climatically, the Holocene may be divided evenly into the Hypsithermal and Neoglacial periods, the boundary coincides with the start of the Bronze Age in Europe. According to some scholars, a division, the Anthropocene, has now begun. Continental motions due to plate tectonics are less than a kilometre over a span of only 10,000 years, however, ice melt caused world sea levels to rise about 35 m in the early part of the Holocene. The sea level rise and temporary land depression allowed temporary marine incursions into areas that are now far from the sea, Holocene marine fossils are known, for example, from Vermont and Michigan. Other than higher-latitude temporary marine incursions associated with depression, Holocene fossils are found primarily in lakebed, floodplain. Holocene marine deposits along low-latitude coastlines are rare because the rise in sea levels during the period exceeds any likely tectonic uplift of non-glacial origin, post-glacial rebound in the Scandinavia region resulted in the formation of the Baltic Sea. The region continues to rise, still causing weak earthquakes across Northern Europe, the equivalent event in North America was the rebound of Hudson Bay, as it shrank from its larger, immediate post-glacial Tyrrell Sea phase, to near its present boundaries. Climate has been stable over the Holocene. It appears that this was influenced by the glacial ice remaining in the Northern Hemisphere until the later date
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Persian onager
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The Persian onager, also called the Persian wild ass or Persian zebra, is a subspecies of onager native to Iran. It is listed as Endangered, with no more than 600 individuals left in the wild, the Persian onager is also simply named gur, meaning swift in the Persian language, in which the word gur has preserved as the second syllable for ono, which meant donkey. Sometimes, the onager is reserved specifically for this subspecies. However, as the species of the Asiatic wild ass is known simply as onager, it now also serves as the Persian wild asss scientific name. Information on the biology of the subspecies and how it differs from others is lacking. Onagers used to be numerous from the Middle East to China, however, until the 19th century, their population has been reduced from several thousand to a few thousand. Currently, more than 600 Persian onagers are living in the wild, Persian wild asses are known to inhabit mountain steppes, semidesert, or desert plains. They are usually found in desert steppes and their largest population is found in Khar Turan National Park. The Persian onager is listed as endangered by IUCN Red List. Currently, poaching for meat and hides, competition with livestock, Asiatic wild asses are highly and legally protected, hunting them is forbidden. A few Persian onagers are breeding in various zoos of Europe and the Middle East, such as Chester, Whipsnade, the Smithsonian Conservation Biology Institute in Front Royal, Virginia, also breeds Persian onagers, including two born in June 2015. Since 2003, Persian onagers have been introduced in Saudi Arabia, introduced Persian onagers live in deserts foraging on grasses and branches or woodier plant material in dry seasons. In 1968,11 Persian and Turkmenian onagers were flown from their countries to Israel in exchange for Israeli gazelles and these were bred in captivity at the Hai Bar Yotvata wildlife sanctuary. Together, they bred a few Persian/Turkmenian hybrids in Israel, offspring were introduced into the wild in the Negev Mountains area, intended to replace the local subspecies gone extinct. The introduced onagers have since established a population around 200 individuals
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Russia
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Russia, also officially the Russian Federation, is a country in Eurasia. The European western part of the country is more populated and urbanised than the eastern. Russias capital Moscow is one of the largest cities in the world, other urban centers include Saint Petersburg, Novosibirsk, Yekaterinburg, Nizhny Novgorod. Extending across the entirety of Northern Asia and much of Eastern Europe, Russia spans eleven time zones and incorporates a range of environments. It shares maritime borders with Japan by the Sea of Okhotsk, the East Slavs emerged as a recognizable group in Europe between the 3rd and 8th centuries AD. Founded and ruled by a Varangian warrior elite and their descendants, in 988 it adopted Orthodox Christianity from the Byzantine Empire, beginning the synthesis of Byzantine and Slavic cultures that defined Russian culture for the next millennium. Rus ultimately disintegrated into a number of states, most of the Rus lands were overrun by the Mongol invasion. The Soviet Union played a role in the Allied victory in World War II. The Soviet era saw some of the most significant technological achievements of the 20th century, including the worlds first human-made satellite and the launching of the first humans in space. By the end of 1990, the Soviet Union had the second largest economy, largest standing military in the world. It is governed as a federal semi-presidential republic, the Russian economy ranks as the twelfth largest by nominal GDP and sixth largest by purchasing power parity in 2015. Russias extensive mineral and energy resources are the largest such reserves in the world, making it one of the producers of oil. The country is one of the five recognized nuclear weapons states and possesses the largest stockpile of weapons of mass destruction, Russia is a great power as well as a regional power and has been characterised as a potential superpower. The name Russia is derived from Rus, a state populated mostly by the East Slavs. However, this name became more prominent in the later history, and the country typically was called by its inhabitants Русская Земля. In order to distinguish this state from other states derived from it, it is denoted as Kievan Rus by modern historiography, an old Latin version of the name Rus was Ruthenia, mostly applied to the western and southern regions of Rus that were adjacent to Catholic Europe. The current name of the country, Россия, comes from the Byzantine Greek designation of the Kievan Rus, the standard way to refer to citizens of Russia is Russians in English and rossiyane in Russian. There are two Russian words which are translated into English as Russians
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Conservation status
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The conservation status of a group of organisms indicates whether the group still exists and how likely the group is to become extinct in the near future. Various systems of conservation status exist and are in use at international, multi-country, the IUCN Red List of Threatened Species is the best known worldwide conservation status listing and ranking system. Also included are species that have gone extinct since 500 AD, when discussing the IUCN Red List, the official term threatened is a grouping of three categories, critically endangered, endangered, and vulnerable. Widespread and abundant taxa are included in this category, Data deficient – Not enough data to make an assessment of its risk of extinction Not evaluated – Has not yet been evaluated against the criteria. The Convention on International Trade in Endangered Species of Wild Fauna and Flora aims to ensure that trade in specimens of wild animals. Many countries require CITES permits when importing plants and animals listed on CITES, in the European Union, the Birds and Habitats Directives are the legal instruments that evaluate the conservation status within the EU of species and habitats. NatureServe conservation status focuses on Latin America, United States, Canada, and it has been developed by scientists from NatureServe, The Nature Conservancy, and the network of natural heritage programs and data centers. It is increasingly integrated with the IUCN Red List system and its categories for species include, presumed extinct, possibly extinct, critically imperiled, imperiled, vulnerable, apparently secure, and secure. The system also allows ambiguous or uncertain ranks including inexact numeric ranks, NatureServe adds a qualifier for captive or cultivated only, which has a similar meaning to the IUCN Red List extinct in the wild status. The Red Data Book of the Russian Federation is used within the Russian Federation, in Australia, the Environment Protection and Biodiversity Conservation Act 1999 describes lists of threatened species, ecological communities and threatening processes. The categories resemble those of the 1994 IUCN Red List Categories & Criteria, prior to the EPBC Act, a simpler classification system was used by the Endangered Species Protection Act 1992. Some state and territory governments also have their own systems for conservation status, in Belgium, the Flemish Research Institute for Nature and Forest publishes an online set of more than 150 nature indicators in Dutch. In Canada, the Committee on the Status of Endangered Wildlife in Canada is a group of experts that assesses and designates which wild species are in danger of disappearing from Canada. Under the Species at Risk Act, it is up to the federal government, in China, the State, provinces and some counties have determined their key protected wildlife species. There is the China red data book, in Finland, a large number of species are protected under the Nature Conservation Act, and through the EU Habitats Directive and EU Birds Directive. In Germany, the Federal Agency for Nature Conservation publishes red lists of endangered species, india has the Wild Life Protection Act,1972, Amended 2003 and the Biological Diversity Act,2002. In Japan, the Ministry of Environment publishes a Threatened Wildlife of Japan Red Data Book, in the Netherlands, the Dutch Ministry of Agriculture, Nature and Food Quality publishes a list of threatened species, and conservation is enforced by the Nature Conservation Act 1998. Species are also protected through the Wild Birds and Habitats Directives, in New Zealand, the Department of Conservation publishes the New Zealand Threat Classification System lists
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Near Threatened
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The IUCN notes the importance of re-evaluating near-threatened taxa at appropriate intervals. The rationale used for near-threatened taxa usually includes the criteria of vulnerable which are plausible or nearly met, additionally, the 402 conservation-dependent taxa may also be considered near-threatened. Prior to 2001, the IUCN used the version 2.3 Categories and Criteria, with this category system, Near Threatened and Conservation Dependent were both subcategories of the category Lower Risk. IUCN Red List near threatened species, ordered by taxonomic rank, category, IUCN Red List near threatened species, ordered alphabetically
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IUCN Red List
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The IUCN Red List of Threatened Species, founded in 1964, is the worlds most comprehensive inventory of the global conservation status of biological species. The International Union for Conservation of Nature is the main authority on the conservation status of species. A series of Regional Red Lists are produced by countries or organizations, the IUCN Red List is set upon precise criteria to evaluate the extinction risk of thousands of species and subspecies. These criteria are relevant to all species and all regions of the world, the aim is to convey the urgency of conservation issues to the public and policy makers, as well as help the international community to try to reduce species extinction. Major species assessors include BirdLife International, the Institute of Zoology, the World Conservation Monitoring Centre, collectively, assessments by these organizations and groups account for nearly half the species on the Red List. The IUCN aims to have the category of every species re-evaluated every five years if possible, the 1964 IUCN Red List of Threatened Plants used the older pre-criteria Red List assessment system. Plants listed may not, therefore, appear in the current Red List, IUCN advise that is best to check both the online Red List and the 1997 plants Red List publication. The 2006 Red List, released on 4 May 2006 evaluated 40,168 species as a whole, plus an additional 2,160 subspecies, varieties, aquatic stocks, on 12 September 2007, the World Conservation Union released the 2007 IUCN Red List of Threatened Species. Russ Mittermeier, chief of Swiss-based IUCNs Primate Specialist Group, stated that 16,306 species are endangered with extinction,188 more than in 2006, the Red List includes the Sumatran orangutan in the Critically Endangered category and the Bornean orangutan in the Endangered category. The study shows at least 1,141 of the 5,487 mammals on Earth are known to be threatened with extinction, and 836 are listed as Data Deficient. The Red List of 2012 was released 19 July 2012 at Rio+20 Earth Summit, nearly 2,000 species were added, the IUCN assessed a total of 63,837 species which revealed 19,817 are threatened with extinction. With 3,947 described as endangered and 5,766 as endangered. At threat are 41% of amphibian species, 33% of reef-building corals, 30% of conifers, 25% of mammals, the IUCN Red List has listed 132 species of plants and animals from India as Critically Endangered. Extinct – No known individuals remaining, extinct in the wild – Known only to survive in captivity, or as a naturalized population outside its historic range. Critically endangered – Extremely high risk of extinction in the wild, Endangered – High risk of extinction in the wild. Vulnerable – High risk of endangerment in the wild, near threatened – Likely to become endangered in the near future. Does not qualify for a more at-risk category, widespread and abundant taxa are included in this category. Data deficient – Not enough data to make an assessment of its risk of extinction, Not evaluated – Has not yet been evaluated against the criteria
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Taxonomy (biology)
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Taxonomy is the science of defining groups of biological organisms on the basis of shared characteristics and giving names to those groups. The exact definition of taxonomy varies from source to source, but the core of the remains, the conception, naming. There is some disagreement as to whether biological nomenclature is considered a part of taxonomy, the broadest meaning of taxonomy is used here. The word taxonomy was introduced in 1813 by Candolle, in his Théorie élémentaire de la botanique, the term alpha taxonomy is primarily used today to refer to the discipline of finding, describing, and naming taxa, particularly species. In earlier literature, the term had a different meaning, referring to morphological taxonomy, ideals can, it may be said, never be completely realized. They have, however, a value of acting as permanent stimulants. Some of us please ourselves by thinking we are now groping in a beta taxonomy, turrill thus explicitly excludes from alpha taxonomy various areas of study that he includes within taxonomy as a whole, such as ecology, physiology, genetics, and cytology. He further excludes phylogenetic reconstruction from alpha taxonomy, thus, Ernst Mayr in 1968 defined beta taxonomy as the classification of ranks higher than species. This activity is what the term denotes, it is also referred to as beta taxonomy. How species should be defined in a group of organisms gives rise to practical and theoretical problems that are referred to as the species problem. The scientific work of deciding how to define species has been called microtaxonomy, by extension, macrotaxonomy is the study of groups at higher taxonomic ranks, from subgenus and above only, than species. While some descriptions of taxonomic history attempt to date taxonomy to ancient civilizations, earlier works were primarily descriptive, and focused on plants that were useful in agriculture or medicine. There are a number of stages in scientific thinking. Early taxonomy was based on criteria, the so-called artificial systems. Later came systems based on a complete consideration of the characteristics of taxa, referred to as natural systems, such as those of de Jussieu, de Candolle and Bentham. The publication of Charles Darwins Origin of Species led to new ways of thinking about classification based on evolutionary relationships and this was the concept of phyletic systems, from 1883 onwards. This approach was typified by those of Eichler and Engler, the advent of molecular genetics and statistical methodology allowed the creation of the modern era of phylogenetic systems based on cladistics, rather than morphology alone. Taxonomy has been called the worlds oldest profession, and naming and classifying our surroundings has likely been taking place as long as mankind has been able to communicate
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Animal
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Animals are multicellular, eukaryotic organisms of the kingdom Animalia. The animal kingdom emerged as a clade within Apoikozoa as the group to the choanoflagellates. Animals are motile, meaning they can move spontaneously and independently at some point in their lives and their body plan eventually becomes fixed as they develop, although some undergo a process of metamorphosis later in their lives. All animals are heterotrophs, they must ingest other organisms or their products for sustenance, most known animal phyla appeared in the fossil record as marine species during the Cambrian explosion, about 542 million years ago. Animals can be divided broadly into vertebrates and invertebrates, vertebrates have a backbone or spine, and amount to less than five percent of all described animal species. They include fish, amphibians, reptiles, birds and mammals, the remaining animals are the invertebrates, which lack a backbone. These include molluscs, arthropods, annelids, nematodes, flatworms, cnidarians, ctenophores, the study of animals is called zoology. The word animal comes from the Latin animalis, meaning having breath, the biological definition of the word refers to all members of the kingdom Animalia, encompassing creatures as diverse as sponges, jellyfish, insects, and humans. Aristotle divided the world between animals and plants, and this was followed by Carl Linnaeus, in the first hierarchical classification. In Linnaeuss original scheme, the animals were one of three kingdoms, divided into the classes of Vermes, Insecta, Pisces, Amphibia, Aves, and Mammalia. Since then the last four have all been subsumed into a single phylum, in 1874, Ernst Haeckel divided the animal kingdom into two subkingdoms, Metazoa and Protozoa. The protozoa were later moved to the kingdom Protista, leaving only the metazoa, thus Metazoa is now considered a synonym of Animalia. Animals have several characteristics that set apart from other living things. Animals are eukaryotic and multicellular, which separates them from bacteria and they are heterotrophic, generally digesting food in an internal chamber, which separates them from plants and algae. They are also distinguished from plants, algae, and fungi by lacking cell walls. All animals are motile, if only at life stages. In most animals, embryos pass through a stage, which is a characteristic exclusive to animals. With a few exceptions, most notably the sponges and Placozoa and these include muscles, which are able to contract and control locomotion, and nerve tissues, which send and process signals
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Chordate
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Chordates are deuterostomes, as during the embryo development stage the anus forms before the mouth. They are also bilaterally symmetric coelomates, in the case of vertebrate chordates, the notochord is usually replaced by a vertebral column during development, and they may have body plans organized via segmentation. There are also additional extinct taxa, the Vertebrata are sometimes considered as a subgroup of the clade Craniata, consisting of chordates with a skull, the Craniata and Tunicata compose the clade Olfactores. Of the more than 65,000 living species of chordates, the worlds largest and fastest animals, the blue whale and peregrine falcon respectively, are chordates, as are humans. Fossil chordates are known from at least as early as the Cambrian explosion, Hemichordata, which includes the acorn worms, has been presented as a fourth chordate subphylum, but it now is usually treated as a separate phylum. The Hemichordata, along with the Echinodermata, form the Ambulacraria, the Chordata and Ambulacraria form the superphylum Deuterostomia, composed of the deuterostomes. Attempts to work out the relationships of the chordates have produced several hypotheses. All of the earliest chordate fossils have found in the Early Cambrian Chengjiang fauna. Because the fossil record of early chordates is poor, only molecular phylogenetics offers a prospect of dating their emergence. However, the use of molecular phylogenetics for dating evolutionary transitions is controversial and it has also proved difficult to produce a detailed classification within the living chordates. Attempts to produce family trees shows that many of the traditional classes are paraphyletic. While this has been known since the 19th century, an insistence on only monophyletic taxa has resulted in vertebrate classification being in a state of flux. Although the name Chordata is attributed to William Bateson, it was already in prevalent use by 1880, ernst Haeckel described a taxon comprising tunicates, cephalochordates, and vertebrates in 1866. Though he used the German vernacular form, it is allowed under the ICZN code because of its subsequent latinization, among the vertebrate sub-group of chordates the notochord develops into the spine, and in wholly aquatic species this helps the animal to swim by flexing its tail. In fish and other vertebrates, this develops into the spinal cord, the pharynx is the part of the throat immediately behind the mouth. In fish, the slits are modified to form gills, a muscular tail that extends backwards behind the anus. This is a groove in the wall of the pharynx. In filter-feeding species it produces mucus to gather food particles, which helps in transporting food to the esophagus and it also stores iodine, and may be a precursor of the vertebrate thyroid gland
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Mammal
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Mammals are any vertebrates within the class Mammalia, a clade of endothermic amniotes distinguished from reptiles by the possession of a neocortex, hair, three middle ear bones and mammary glands. All female mammals nurse their young with milk, secreted from the mammary glands, Mammals include the largest animals on the planet, the great whales. The basic body type is a quadruped, but some mammals are adapted for life at sea, in the air, in trees. The largest group of mammals, the placentals, have a placenta, Mammals range in size from the 30–40 mm bumblebee bat to the 30-meter blue whale. With the exception of the five species of monotreme, all modern mammals give birth to live young, most mammals, including the six most species-rich orders, belong to the placental group. The largest orders are the rodents, bats and Soricomorpha, the next three biggest orders, depending on the biological classification scheme used, are the Primates, the Cetartiodactyla, and the Carnivora. Living mammals are divided into the Yinotheria and Theriiformes There are around 5450 species of mammal, in some classifications, extant mammals are divided into two subclasses, the Prototheria, that is, the order Monotremata, and the Theria, or the infraclasses Metatheria and Eutheria. The marsupials constitute the group of the Metatheria, and include all living metatherians as well as many extinct ones. Much of the changes reflect the advances of cladistic analysis and molecular genetics, findings from molecular genetics, for example, have prompted adopting new groups, such as the Afrotheria, and abandoning traditional groups, such as the Insectivora. The mammals represent the only living Synapsida, which together with the Sauropsida form the Amniota clade, the early synapsid mammalian ancestors were sphenacodont pelycosaurs, a group that produced the non-mammalian Dimetrodon. At the end of the Carboniferous period, this group diverged from the line that led to todays reptiles. Some mammals are intelligent, with some possessing large brains, self-awareness, Mammals can communicate and vocalize in several different ways, including the production of ultrasound, scent-marking, alarm signals, singing, and echolocation. Mammals can organize themselves into fission-fusion societies, harems, and hierarchies, most mammals are polygynous, but some can be monogamous or polyandrous. They provided, and continue to provide, power for transport and agriculture, as well as commodities such as meat, dairy products, wool. Mammals are hunted or raced for sport, and are used as model organisms in science, Mammals have been depicted in art since Palaeolithic times, and appear in literature, film, mythology, and religion. Defaunation of mammals is primarily driven by anthropogenic factors, such as poaching and habitat destruction, Mammal classification has been through several iterations since Carl Linnaeus initially defined the class. No classification system is accepted, McKenna & Bell and Wilson & Reader provide useful recent compendiums. Though field work gradually made Simpsons classification outdated, it remains the closest thing to a classification of mammals
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Odd-toed ungulate
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Members of the order Perissodactyla, otherwise known as odd-toed ungulates, are mammals characterized by an odd number of toes and by hindgut fermentation with somewhat simple stomachs. Perissodactyla comes from the Ancient Greek περισσός + δάκτυλος, unlike the even-toed ungulates, they digest plant cellulose in their intestines rather than in one or more stomach chambers. The order includes three extant families, Equidae, Rhinocerotidae, and Tapiridae, with a total of about 17 species. In spite of their different appearances, they were recognized as related families in the 19th century by the zoologist Richard Owen. The largest odd-toed ungulates are rhinoceroses, and the extinct Paraceratherium, at the other extreme, an early member of the order, the prehistoric horse Hyracotherium, had a shoulder height of only 30 to 60 cm. Apart from dwarf varieties of the horse and donkey, perissodactyls reach a body length of 180–420 cm. While rhinos have only sparse hair and exhibit a thick epidermis, tapirs and horses have dense, most species are grey or brown, although zebras and young tapirs are striped. The main axes of both the front and rear feet pass through the toe, which is always the largest. The remaining toes have been reduced in size to varying degrees, tapirs, which are adapted to walking on soft ground, have four toes on their fore feet and three on their hind feet. Living rhinos have three toes on both the front and hind feet, modern equines possess only a single toe, however, their feet are equipped with hooves, which almost completely cover the toe. Rhinos and tapirs, by contrast, have hooves covering only the edge of the toes. The ulnae and fibulae are reduced in horses, a common feature that clearly distinguishes this group from other mammals is the saddle-shaped ankle between the astragalus and the scaphoid, which greatly restricts the mobility of the foot. The thigh is relatively short, and the clavicle is absent, odd-toed ungulates have a long upper jaw with an extended diastema between the front and cheek teeth, giving them an elongated head. The various forms of snout between families are due to differences in the form of the premaxilla, the lacrimal bone has projecting cusps in the eye sockets and a wide contact with the nasal bone. The temporomandibular joint is high and the mandible is enlarged, rhinos have one or two horns made of agglutinated keratin, unlike the horns of even-toed ungulates, which have a bony core. The number and form of the teeth vary according to diet, the incisors and canines can be very small or completely absent, as in the two African species of rhinoceros. In the horses, usually only the males possess canines, the surface shape and height of the molars is heavily dependent on whether soft leaves or hard grass makes up the main component of their diets. Three or four teeth are present on each jaw half, so the dental formula of odd-toed ungulates is
27.
Equidae
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Equidae is the taxonomic family of horses and related animals, including the extant horses, donkeys, and zebras, and many other species known only from fossils. All extant species are in the genus Equus, Equidae belongs to the order Perissodactyla, which includes the extant tapirs and rhinoceros, and several extinct families. The term equid refers to any member of family, including any equine. The oldest known fossils assigned to Equidae date from the early Eocene,54 million years ago and they used to be assigned to the genus Hyracotherium, but the type species of that genus now is regarded to be not a member of this family. The other species have split off into different genera. These early Equidae were fox-sized animals with three toes on the feet, and four on the front feet. They were herbivorous browsers on relatively soft plants, and already adapted for running, the complexity of their brains suggest that they already were alert and intelligent animals. Later species reduced the number of toes, and developed teeth more suited for grinding up grasses, the family became relatively diverse during the Miocene, with many new species appearing. By this time, equids were more truly horse-like, having developed the body shape of the modern animals. Many of these bore the main weight of their bodies on their central, third, toe, with the others becoming reduced. The sole surviving genus, Equus, had evolved by the early Pleistocene, order Perissodactyla † indicates extinct taxa
28.
Equus (genus)
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Equus is a genus of mammals in the family Equidae, which includes horses, asses, and zebras. Within Equidae, Equus is the only recognized extant genus, comprising seven living species, the term equine refers to any member of this genus, including horses. Like Equidae more broadly, Equus has numerous species known only from fossils. The genus most likely originated in North America and spread quickly to the Old World, Equines are odd-toed ungulates with slender legs, long heads, relatively long necks, manes and long tails. All species are herbivorous, and mostly grazers with simpler digestive systems than ruminants, while the domestic horse and donkey exist worldwide, wild equine populations are limited to Africa and Asia. In both systems, females take care of their offspring but males may play a role as well, Equines communicate with each other both visually and vocally. Human activities have threatened wild equine populations and out of the seven living species, only the plains zebra remains widespread and abundant. The word equus is Latin for horse, and is cognate with the Greek ἵππος, horse, and Mycenaean Greek i-qo /ikkʷos/, the genus Equus was first described by Carl Linnaeus in 1758. It is the only recognized extant genus in the family Equidae, the first equids were small, dog-sized mammals adapted for browsing on shrubs during the Eocene, around 54 million years ago. These animals had three toes on the feet and four on the front feet with small hooves in place of claws. Equids developed into larger, three-toed animals during the Oligocene and Miocene, from there, the tridactyl toes became progressively smaller through the Pleistocene until the emergence of the single-toed Equus. The genus Equus, which all extant equines, is believed to have evolved from Dinohippus. One of the oldest species is Equus simplicidens, described as zebra-like with a head shape. The oldest material to date was found in Idaho, USA, the genus appears to have spread quickly into the Old World, with the similarly aged E. livenzovensis documented from western Europe and Russia. Molecular phylogenies indicate that the most recent common ancestor of all modern equines lived ~5.6 mya, mitochondrial evidence supports the division of Equus species into non-caballoid and caballoids or true horses. Of the extant equine species, the lineage of the asses may have diverged first, zebras appear to be monophyletic and differentiated in Africa where they are endemic. Molecular dating indicates the caballoid lineage diverged from the non-caballoids 4 mya, genetic results suggest that all North American fossils of caballine equines, as well as South American fossils traditionally placed in the subgenus E. belong to E. ferus. Remains attributed to a variety of species and lumped together as New World stilt-legged horses probably all belong to a species that was endemic to North America
29.
Asinus
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The donkey or ass is a domesticated member of the horse family, Equidae. The wild ancestor of the donkey is the African wild ass, the donkey has been used as a working animal for at least 5000 years. There are more than 40 million donkeys in the world, mostly in underdeveloped countries, working donkeys are often associated with those living at or below subsistence levels. Small numbers of donkeys are kept for breeding or as pets in developed countries, a male donkey or ass is called a jack, a female a jenny or jennet, a young donkey is a foal. Jack donkeys are used to mate with female horses to produce mules. Asses were first domesticated around 3000 BC, probably in Egypt or Mesopotamia and they continue to fill important roles in many places today. While domesticated species are increasing in numbers, the African wild ass is an endangered species, as beasts of burden and companions, asses and donkeys have worked together with humans for millennia. Traditionally, the name for the donkey is Equus asinus asinus based on the principle of priority used for scientific names of animals. This means that the scientific name for the donkey is Equus africanus asinus when it is considered a subspecies. At one time, the ass was the more common term for the donkey. The first recorded use of donkey was in either 1784 or 1785, while the word ass has cognates in most other Indo-European languages, donkey is an etymologically obscure word for which no credible cognate has been identified. From the 18th century, donkey gradually replaced ass, and jenny replaced she-ass, by the end of the 17th century, changes in pronunciation of both ass and arse had caused them to become homophones. Donkeys vary considerably in size, depending on breed and management, the height at the withers ranges from 7.3 to 15.3 hands, and the weight from 80 to 480 kg. Working donkeys in the poorest countries have an expectancy of 12 to 15 years, in more prosperous countries. Donkeys are adapted to desert lands. Unlike wild and feral horses, wild donkeys in dry areas are solitary, each adult donkey establishes a home range, breeding over a large area may be dominated by one jack. Donkeys have large ears, which may pick up more distant sounds, donkeys can defend themselves by biting, striking with the front hooves or kicking with the hind legs. A jenny is normally pregnant for about 12 months, though the period varies from 11 to 14 months
30.
Binomial nomenclature
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Such a name is called a binomial name, a binomen, binominal name or a scientific name, more informally it is also called a Latin name. The first part of the name identifies the genus to which the species belongs, for example, humans belong to the genus Homo and within this genus to the species Homo sapiens. The formal introduction of system of naming species is credited to Carl Linnaeus. But Gaspard Bauhin, in as early as 1623, had introduced in his book Pinax theatri botanici many names of genera that were adopted by Linnaeus. Although the general principles underlying binomial nomenclature are common to these two codes, there are differences, both in the terminology they use and in their precise rules. Similarly, both parts are italicized when a binomial name occurs in normal text, thus the binomial name of the annual phlox is now written as Phlox drummondii. In scientific works, the authority for a name is usually given, at least when it is first mentioned. In zoology Patella vulgata Linnaeus,1758, the original name given by Linnaeus was Fringilla domestica, the parentheses indicate that the species is now considered to belong in a different genus. The ICZN does not require that the name of the person who changed the genus be given, nor the date on which the change was made, in botany Amaranthus retroflexus L. – L. is the standard abbreviation used in botany for Linnaeus. – Linnaeus first named this bluebell species Scilla italica, Rothmaler transferred it to the genus Hyacinthoides, the ICN does not require that the dates of either publication be specified. Prior to the adoption of the binomial system of naming species. Together they formed a system of polynomial nomenclature and these names had two separate functions. First, to designate or label the species, and second, to be a diagnosis or description, such polynomial names may sometimes look like binomials, but are significantly different. For example, Gerards herbal describes various kinds of spiderwort, The first is called Phalangium ramosum, Branched Spiderwort, is aptly termed Phalangium Ephemerum Virginianum, Soon-Fading Spiderwort of Virginia. The Latin phrases are short descriptions, rather than identifying labels, the Bauhins, in particular Caspar Bauhin, took some important steps towards the binomial system, by pruning the Latin descriptions, in many cases to two words. The adoption by biologists of a system of binomial nomenclature is due to Swedish botanist and physician Carl von Linné. It was in his 1753 Species Plantarum that he first began using a one-word trivial name together with a generic name in a system of binomial nomenclature. This trivial name is what is now known as an epithet or specific name
31.
Peter Simon Pallas
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Peter Simon Pallas was a German zoologist and botanist who worked in Russia. Pallas was born in Berlin, the son of Professor of Surgery Simon Pallas and he studied with private tutors and took an interest in natural history, later attending the University of Halle and the University of Göttingen. In 1760, he moved to the University of Leiden and passed his doctors degree at the age of 19, Pallas traveled throughout the Netherlands and to London, improving his medical and surgical knowledge. He then settled at The Hague, and his new system of classification was praised by Georges Cuvier. Pallas wrote Miscellanea Zoologica, which included descriptions of several new to science which he had discovered in the Dutch museum collections. A planned voyage to southern Africa and the East Indies fell through when his father recalled him to Berlin, there, he began work on his Spicilegia Zoologica. He explored the Caspian Sea, the Ural and Altai Mountains, the regular reports which Pallas sent to St Petersburg were collected and published as Reise durch verschiedene Provinzen des Russischen Reichs. They covered a range of topics, including geology and mineralogy, reports on the native peoples and their religions. In 1776, Pallas was elected a member of the Royal Swedish Academy of Sciences. Pallas settled in St Petersburg, becoming a favourite of Catherine II and teaching history to the Grand Dukes Alexander. He was provided with the plants collected by other naturalists to compile the Flora Rossica, a Russian flora, and started work on his Zoographica Rosso-Asiatica and he also published an account of Johann Anton Güldenstädts travels in the Caucasus. The Empress bought Pallass large natural history collection for 2,000 rubles,500 more than his asking price, during this period, Pallas helped plan the Mulovsky expedition, which was cancelled in October 1787. Between 1793 and 1794, Pallas led an expedition to southern Russia, visiting the Crimea. He was accompanied by his daughter and his new wife, an artist, servants, in February 1793, they travelled to Saratov and then downriver to Tsaritsyn. They spent the spring exploring the country to the east, and in August travelled along the banks of the Caspian Sea, in September, they travelled to the Crimea, wintering in Simferopol. Pallas spent the spring of 1794 exploring to the southeast, and in July travelled up the valley of the Dnieper, Pallas gave his account of the journey in his P. S. Pallas Bemerkungen auf einer Reise in die Südlichen Statthalterschaften des Russischen Reichs. Catherine II gave him an estate at Simferopol, where Pallas lived until the death of his second wife in 1810. He was then granted permission to leave Russia by Emperor Alexander, and returned to Berlin and his grave is preserved in the Protestant Friedhof I der Jerusalems- und Neuen Kirchengemeinde in Berlin-Kreuzberg, south of Hallesches Tor
32.
Mongolian wild ass
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The Mongolian wild ass, also known as Mongolian khulan is the nominate subspecies of the onager. It is found in Mongolia and northern China, and was found in eastern Kazakhstan. Previously in 2003, there were a population of less than 20,000 individuals. In 2009, the Mongolian wild ass population had decreased to 14,000 total individuals, in 2015, the Mongolian wild ass are listed as Near Threatened by the IUCN and it is also estimated that there are 23,000 mature individuals in total. The Mongolian wild ass may be synonymous with the possible proposed separate subspecies, the Mongolian wild ass, like most Asiatic wild asses, lives in deserts, cold deserts, steppes, shrublands and mountains. The Gobi Desert is the main stronghold. It may also be found at the Altai Mountains, the Mongolian wild ass is the most widespread subspecies, although despite that, the subspecies lost about 50% of its former distribution range in Mongolia in the past 70 years. Its distribution range then dramatically reduced during the 1990s, a 1994–1997 survey estimated its population size at 33,000 to 63,000 individuals over a continuous distribution range encompassing all of southern Mongolia. In 2003, a new survey found approximately 20,000 individuals over an area of 177,563 square kilometres in southern Mongolia, the species had decreased to 14,000 individuals in 2009. The population estimates of the Mongolian population should be treated with due to a lack of proven survey protocols. The Mongolian wild ass are herbivorous mammals and they feed on grasses, herbs and vegetation. They also feed on shrubs and trees in drier habitats, during spring and summer in Mongolia, the succulent plants of the Zygophyllaceae family form an important component of the diet of the Mongolian wild ass. Mongolian khulans are known to dig holes at dry river beds, watering holes dug by khulans are also used by other species as well as by humans to access to water. The Mongolian wild ass population is declining due to poaching and competition from grazing livestock, the conservation status of the species is evaluated as endangered. The Mongolian khulan are threatened by predators such as gray wolves, dholes. Poaching for meat appears to be a problem in Mongolia. For some parts of the population, wild ass and other wildlife meat seems to provide a substitute or even a cheap alternative to meat from domestic animals. In 2005, a survey based on questionnaires, suggested that as many as 4,500 wild asses, about 20% of the whole population
33.
Turkmenian kulan
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The Turkmenian kulan, also called Transcaspian wild ass, Turkmenistani onager or simply the kulan, is a subspecies of onager native to Central Asia. It was declared Endangered in 2016, the speciess population had recently been in decline in the country while it slowly increases in reintroduction sites. Previously in 2005, the population was estimated at 1, 295-1,345 in Turkmenistan, however, certain fragmented populations of Transcaspian wild ass are currently on the rise to even more than 2,000 individuals in the wild. It is also estimated that over 6,000 kulans live in Central Asia, the Turkmenian kulan is one of the largest subspecies of the onagers. It is 200–250 cm long, 100–140 cm tall at the shoulder, male onagers are larger than the females. The Transcaspian wild ass is characterized by a brown coat. It also has shaggy black mane and a tuft at the tail end, during the summer, the Turkmenian kulans coat is leaner and brown, then it turns into thick grayish-brown winter coat during the cold season. The Turkmenian kulan lives in Central Asian deltas, hot and cold deserts or semi-deserts, steppes, the Turkmen specimen used to be the most widespread of the onagers, ranging from northernmost Iran and northern Afghanistan, the Transcaspian Oblast and western China to southern Siberia. It used to live in Saryesik-Atyrau Desert of Kazakhstan, however, it has gone extinct from that location. The Turkmenian kulan are herbivorous mammals and they feed on herbs, shrubbery and plants. Most of their liquid comes from food, although often traveling at water sources, the mare lives with foals in small herds. The dominant stallion defends the surrounding areas to the sources and tries to mate with any females that come close to drink. The mare, after a period of about a year. The scorching heat of the Central Asian deserts makes the Transcaspian wild ass mostly active at dawn, the kulans, like most onagers, are one of the fastest land mammals and can run at high speed of 70 km/h. The now-extinct Central Asian predators, the Caspian tigers and Asiatic cheetahs also preyed on the kulans, however, like other onagers, they have anti-predatory protections. Group of stallions can cooperate and chase off the predators, the Asiatic wild ass of Central Asia is mostly threatened by illegal hunting. In early 1919, the Soviet Union placed the Turkmenian kulan under protection, however, the measures enacted in their favor have not saved them from local extinctions, such as in Kazakhstan, where the subspecies disappeared in 1935. The main kulan population has suffered a decline in recent years
34.
Colin Groves
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Colin Peter Groves is Professor of Biological Anthropology at the Australian National University in Canberra, Australia. Born in England on 24 June 1942, Colin Groves completed a Bachelor of Science at University College London in 1963, and a Doctor of Philosophy at the Royal Free Hospital School of Medicine in 1966. From 1966 to 1973, Dr. Groves was a Postdoctoral Researcher and Teaching Fellow at the University of California, Berkeley, Queen Elizabeth College and he emigrated to Australia in 1974, and has been at the Australian National University since, being promoted to full Professor in 2000. Professor Groves research interests are human evolution, primates, mammalian taxonomy, skeletal analysis, biological anthropology, ethnobiology and he has conducted extensive fieldwork in Kenya, Tanzania, Rwanda, India, Iran, China, Indonesia, Sri Lanka and the Democratic Republic of Congo. Along with the Czech biologist Professor Vratislav Mazák, Groves was the describer of Homo ergaster, Groves also wrote Primate Taxonomy published by the Smithsonian Institution Press in 2001, and Ungulate Taxonomy. He is a member of the Australian Skeptics and has many published skeptical papers. He has also conducted regular debates with creationists and anti-evolutionists, a theory of human and primate evolution. Laycock, D, ed. Skeptical, a handbook on pseudoscience, from Ussher to Slusher, from Archbish to Gish, or, not in a million years. A Personal Look at the History of Primatology, the Colin Groves Pages The Groves Collection at No Answers in Genesis ANU Faculty Homepage ANU Researcher Profile page
35.
Indian wild ass
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The Indian wild ass or Baluchi wild ass also called the ghudkhur in the local Gujarati language, is a subspecies of the onager native to Southern Asia. As of 2016, it is listed as Near Threatened by IUCN, previously in 2009, the previous census estimated 4,038 Indian wild ass. However, the population was still growing, in December 2014, the population was estimated at 4,451 individuals. It has increased from a jump of 454, however, as of 2015, the current census of the Indian wild ass population has increased to more than 4,800 individuals in and outside of the Wild Ass Wildlife Sanctuary of India. The Indian wild ass, as with most other Asian wild ass subspecies, is different from the African wild ass species. The coat is sandy, but varies from reddish grey, fawn. The animal possesses an erect, dark mane which runs from the back of the head, the mane is then followed by a dark brown stripe running along the back, to the root of the tail. The Indian wild asss range once extended from western India, southern Pakistan, Afghanistan, today, its last refuge lies in the Indian Wild Ass Sanctuary, Little Rann of Kutch and its surrounding areas of the Great Rann of Kutch in the Gujarat province of India. The animal, however, is seen in the districts of Surendranagar, Banaskantha, Mehsana. Saline deserts, arid grasslands and shrublands are its preferred environments, at this place Rebaris live in the Prosopis juliflora jungles in the company of chinkaras, striped hyenas, red foxes, desert cats and Indian wolves, etc. Indian wild asses graze between dawn and dusk, the animal feeds on grass, leaves and fruits of plant, crop, Prosopis pods, and saline vegetation. It is one of the fastest of Indian animals, with speeds clocked at about 70 –80 km. per hour, stallions live either solitarily, or in small groups of twos and threes while family herds remain large. Mating season is in rainy season, when a mare comes into heat, she separates from the herd with a stallion who battles against rivals for her possession. After few days, the returns to the herd. The mare gives birth to one foal, the male foal weans away by 1–2 years of age, while the female continues to stay with the family herd. However, Indias Mughal Emperors and noblemen from the time took great pleasure in hunting it with Emperor Jahangir in his book Tuzk-e-Jahangiri. From 1958-1960, the wild ass became a victim of a known as surra, caused by Trypanosoma evansi and transmitted by flies. In November and December 1961, the wild ass population was reduced to just 870 after to the outbreak of South African Horse Sickness
36.
Syrian wild ass
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The Syrian wild ass, less commonly known as a hemippe or achdari, is an extinct subspecies of onager native to the Arabian peninsula. It ranged across present-day Syria, the Palestinian territories, Israel, Turkey, Jordan, Saudi Arabia, the Syrian wild ass, only one metre high at its shoulder, was the smallest form of Equidae and could not be domesticated. Its coloring changed with the seasons—a tawny olive coat for the summer months and it was known, like other onagers, to be untameable, and was compared to a thoroughbred horse for its beauty and strength. The Syrian wild ass lived in deserts, semideserts, arid grasslands, native to West Asia, they were found in Palestine, Jordan, Turkey, Syria, Saudi Arabia and Iraq. The Syrian wild ass was a grazer and it fed on grass, herbs, leaves, shrubs and tree branches. Syrian wild asses were preyed upon by Asiatic lions, Arabian leopards, striped hyenas, grey wolves, Asiatic cheetahs may have preyed on onager foals. European travelers in the Middle East during the 15th and 16th centuries reported seeing large herds, however, its numbers began to drop precipitously during the 18th and 19th centuries due to overhunting, and its existence was further imperiled by the regional upheaval of World War I. The last known specimen was fatally shot in 1927 et al. Ghams near the Azraq oasis in Jordan, and the last captive specimen died the year at the Tiergarten Schönbrunn. After the extinction of the Syrian wild ass, the Persian onager from Iran was chosen as the subspecies to replace the extinct onagers in the Middle East. The Persian onager was then introduced to the areas of Saudi Arabia. It was also reintroduced, along with the Turkmenian kulan, to Israel and it is believed this may be the wild ass which Ishmael was prophesied to be in Genesis in the Old Testament. References also appear in the Old Testament books of Job, Psalms, Jeremiah, the Qur’an, the main book of Islam, in Surat al-Muddaththir, refers to a scene of humur fleeing from a qaswarah. Mongolian wild ass, Equus hemionus hemionus Turkmenian kulan, Equus hemionus kulan Indian wild ass, Equus hemionus khur Persian onager, Equus hemionus onager Equidae Onager
37.
Synonym (taxonomy)
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For example, Linnaeus was the first to give a scientific name to the Norway spruce, which he called Pinus abies. This name is no longer in use, it is now a synonym of the current scientific name which is Picea abies, unlike synonyms in other contexts, in taxonomy a synonym is not interchangeable with the name of which it is a synonym. In taxonomy, synonyms are not equals, but have a different status, for any taxon with a particular circumscription, position, and rank, only one scientific name is considered to be the correct one at any given time. A synonym cannot exist in isolation, it is always an alternative to a different scientific name, given that the correct name of a taxon depends on the taxonomic viewpoint used a name that is one taxonomists synonym may be another taxonomists correct name. Synonyms may arise whenever the same taxon is described and named more than once, independently. They may also arise when existing taxa are changed, as when two taxa are joined to one, a species is moved to a different genus. To the general user of scientific names, in such as agriculture, horticulture, ecology, general science. A synonym is a name that was used as the correct scientific name but which has been displaced by another scientific name. Thus Oxford Dictionaries Online defines the term as a name which has the same application as another. In handbooks and general texts, it is useful to have mentioned as such after the current scientific name. Synonyms used in this way may not always meet the strict definitions of the synonym in the formal rules of nomenclature which govern scientific names. Changes of scientific name have two causes, they may be taxonomic or nomenclatural, a name change may be caused by changes in the circumscription, position or rank of a taxon, representing a change in taxonomic, scientific insight. A name change may be due to purely nomenclatural reasons, that is, based on the rules of nomenclature, the earliest such name is called the senior synonym, while the later name is the junior synonym. One basic principle of zoological nomenclature is that the earliest correctly published name, synonyms are important because if the earliest name cannot be used, then the next available junior synonym must be used for the taxon. Objective synonyms refer to taxa with the type and same rank. For example, John Edward Gray published the name Antilocapra anteflexa in 1855 for a species of pronghorn, however, it is now commonly accepted that his specimen was an unusual individual of the species Antilocapra americana published by George Ord in 1815. Ords name thus takes precedence, with Antilocapra anteflexa being a subjective synonym. Objective synonyms are common at the level of genera, because for various reasons two genera may contain the type species, these are objective synonyms
38.
Horse
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The horse is one of two extant subspecies of Equus ferus. It is an ungulate mammal belonging to the taxonomic family Equidae. The horse has evolved over the past 45 to 55 million years from a small multi-toed creature, Eohippus, into the large, humans began to domesticate horses around 4000 BC, and their domestication is believed to have been widespread by 3000 BC. Horses in the subspecies caballus are domesticated, although some domesticated populations live in the wild as feral horses. There is an extensive, specialized vocabulary used to describe equine-related concepts, covering everything from anatomy to life stages, size, colors, markings, breeds, locomotion, and behavior. Horses anatomy enables them to use of speed to escape predators and they have a well-developed sense of balance. Related to this need to flee from predators in the wild is an unusual trait, female horses, called mares, carry their young for approximately 11 months, and a young horse, called a foal, can stand and run shortly following birth. Most domesticated horses begin training under saddle or in harness between the ages of two and four and they reach full adult development by age five, and have an average lifespan of between 25 and 30 years. There are more than 300 breeds of horse in the world today, horses were historically used in warfare, from which a wide variety of riding and driving techniques developed, using many different styles of equipment and methods of control. Many products are derived from horses, including meat, milk, hide, hair, bone, humans provide domesticated horses with food, water and shelter, as well as attention from specialists such as veterinarians and farriers. Specific terms and specialized language are used to describe equine anatomy, different life stages, colors, depending on breed, management and environment, the modern domestic horse has a life expectancy of 25 to 30 years. Uncommonly, a few animals live into their 40s and, occasionally, the oldest verifiable record was Old Billy, a 19th-century horse that lived to the age of 62. In modern times, Sugar Puff, who had listed in Guinness World Records as the worlds oldest living pony. The exception is in endurance riding, where the age to compete is based on the animals actual calendar age. The following terminology is used to describe horses of various ages, Colt, a common terminology error is to call any young horse a colt, when the term actually only refers to young male horses. Filly, A female horse under the age of four, foal, A horse of either sex less than one year old. A nursing foal is sometimes called a suckling and a foal that has been weaned is called a weanling, most domesticated foals are weaned at five to seven months of age, although foals can be weaned at four months with no adverse physical effects. Gelding, A castrated male horse of any age, mare, A female horse four years old and older
39.
Asia
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Asia covers an area of 44,579,000 square kilometres, about 30% of Earths total land area and 8. 7% of the Earths total surface area. The continent, which has long been home to the majority of the population, was the site of many of the first civilizations. Asia is notable for not only its large size and population. In general terms, Asia is bounded on the east by the Pacific Ocean, on the south by the Indian Ocean, the western boundary with Europe is a historical and cultural construct, as there is no clear physical and geographical separation between them. The most commonly accepted boundaries place Asia to the east of the Suez Canal, the Ural River, and the Ural Mountains, and south of the Caucasus Mountains, China and India alternated in being the largest economies in the world from 1 to 1800 A. D. The accidental discovery of America by Columbus in search for India demonstrates this deep fascination, the Silk Road became the main East-West trading route in the Asian hitherland while the Straits of Malacca stood as a major sea route. Asia has exhibited economic dynamism as well as robust population growth during the 20th century, given its size and diversity, the concept of Asia—a name dating back to classical antiquity—may actually have more to do with human geography than physical geography. Asia varies greatly across and within its regions with regard to ethnic groups, cultures, environments, economics, historical ties, the boundary between Asia and Africa is the Red Sea, the Gulf of Suez, and the Suez Canal. This makes Egypt a transcontinental country, with the Sinai peninsula in Asia, the border between Asia and Europe was historically defined by European academics. In Sweden, five years after Peters death, in 1730 Philip Johan von Strahlenberg published a new atlas proposing the Urals as the border of Asia, the Russians were enthusiastic about the concept, which allowed them to keep their European identity in geography. Tatishchev announced that he had proposed the idea to von Strahlenberg, the latter had suggested the Emba River as the lower boundary. Over the next century various proposals were made until the Ural River prevailed in the mid-19th century, the border had been moved perforce from the Black Sea to the Caspian Sea into which the Ural River projects. The border between the Black Sea and the Caspian is usually placed along the crest of the Caucasus Mountains, the border between Asia and the loosely defined region of Oceania is usually placed somewhere in the Malay Archipelago. The terms Southeast Asia and Oceania, devised in the 19th century, have had several different geographic meanings since their inception. The chief factor in determining which islands of the Malay Archipelago are Asian has been the location of the possessions of the various empires there. Lewis and Wigen assert, The narrowing of Southeast Asia to its present boundaries was thus a gradual process, Asia is larger and more culturally diverse than Europe. It does not exactly correspond to the borders of its various types of constituents. From the time of Herodotus a minority of geographers have rejected the three-continent system on the grounds there is no or is no substantial physical separation between them
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Species description
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A species description is a formal description of a newly discovered species, usually in the form of a scientific paper. Its purpose is to give a description of a new species of organism and explain how it differs from species which have been described previously. The species description often contains photographs or other illustrations of the type material, the publication in which the species is described gives the new species a formal scientific name. Today, some 1.9 million species have been described and named and it is customary for scientists to introduce all relevant new findings and research in a scientific manuscript, which is sent to other scientists for peer review. If accepted, it is published in a journal of the appropriate discipline or in a book. Descriptions and naming of a new species or other taxa follow the same process, a name of a new species becomes valid with the date of publication of its formal scientific description. A scientific species description must fulfill several formal criteria specified by the nomenclature codes, Species names are written in the 26 letters of the Latin alphabet, but many species names are based on words from other languages, Latinized. Once the manuscript has been accepted for publication, the new name is officially created. Once a species name has been assigned and approved, it can not be changed except in the case of error. For example, a species of beetle was named by a German collector after Adolf Hitler in 1933 when he had become chancellor of Germany. It is not clear whether such a dedication would be considered acceptable or appropriate today, Species names have been chosen on many different bases. Most common is a naming for the external appearance, its origin. Examples would include a bat species named for the two stripes on its back, a named for its Bolivian origin, and an ant species dedicated to the actor Harrison Ford. A scientific name in honor of a person or persons is a known as a taxonomic patronym or patronymic, a number of humorous species names also exist. Literary examples include the genus name Borogovia, which is named after the borogove, a second example, Macrocarpaea apparata was named after the magical spell to apparate from the Harry Potter novels by J. K. Rowling, as it seemed to appear out of nowhere. For example, the genus Victoria was named in honour of Queen Victoria of Great Britain, more recently, a species of lemur was named after the actor John Cleese in recognition of his work to publicize the plight of lemurs in Madagascar. Non-profit ecological organizations may also allow benefactors to name new species in exchange for support for taxonomic research. A German non-profit organisation, BIOPAT - Patrons for Biodiversity has raised more than $450,000 for research, the International Code of Nomenclature for algae, fungi, and plants discourages this practice somewhat, Recommendation 20A
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African wild ass
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The African wild ass or African wild donkey is a wild member of the horse family, Equidae. This species is believed to be the ancestor of the domestic donkey and they live in the deserts and other arid areas of the Horn of Africa, in Eritrea, Ethiopia and Somalia. It formerly had a range north and west into Sudan, Egypt. About 570 individuals exist in the wild, different authors consider the African wild ass and the domesticated donkey one or two species, either view is technically legitimate, though the former is phylogenetically more accurate. The species name for the African wild ass is given as asinus, from the domestic donkey, whose specific name is older. But this usage is erroneous since the International Commission on Zoological Nomenclature has conserved the name Equus africanus in Opinion 2027 and this was done to prevent the confusing situation of the phylogenetic ancestor being taxonomically included in its descendant. Thus, if one species is recognized, the scientific name of the donkey is E. africanus asinus. The first published name for the African wild ass, Asinus africanus, the first available name thus becomes Asinus africanus von Heuglin & Fitzinger,1866. A lectotype is designated, a skull of a female collected by von Heuglin near Atbarah River, Sudan. The two subspecies recognized are the Nubian wild ass Equus africanus africanus, and the Somali wild ass Equus africanus somaliensis, the African wild ass is 2 metres long and 1.25 to 1.45 metres tall at the shoulders, with a tail 30–50 centimetres long. The short, smooth coat is a grey to fawn colour, fading quickly to white on the undersides. There is a slender, dark stripe in all subspecies, while in the Nubian wild ass, as well as the domestic donkey. The legs of the Somali wild ass are horizontally striped with black, on the nape of the neck, there is a stiff, upright mane, the hairs of which are tipped with black. The ears are large with black margins, the tail terminates with a black brush. The hooves are slender and approximately of the diameter as the legs, African wild asses are well suited to life in a desert or semidesert environment. They have tough digestive systems, which can break down desert vegetation and they can also go without water for a fairly long time. Their large ears give them an excellent sense of hearing and help in cooling, because of the sparse vegetation in their environment wild asses live somewhat separated from each other, unlike the tightly grouped herds of wild horses. They have very loud voices, which can be heard for over 3 km, the African wild ass is primarily active in the cooler hours between late afternoon and early morning, seeking shade and shelter amongst the rocky hills during the day
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Donkey
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The donkey or ass is a domesticated member of the horse family, Equidae. The wild ancestor of the donkey is the African wild ass, the donkey has been used as a working animal for at least 5000 years. There are more than 40 million donkeys in the world, mostly in underdeveloped countries, working donkeys are often associated with those living at or below subsistence levels. Small numbers of donkeys are kept for breeding or as pets in developed countries, a male donkey or ass is called a jack, a female a jenny or jennet, a young donkey is a foal. Jack donkeys are used to mate with female horses to produce mules. Asses were first domesticated around 3000 BC, probably in Egypt or Mesopotamia and they continue to fill important roles in many places today. While domesticated species are increasing in numbers, the African wild ass is an endangered species, as beasts of burden and companions, asses and donkeys have worked together with humans for millennia. Traditionally, the name for the donkey is Equus asinus asinus based on the principle of priority used for scientific names of animals. This means that the scientific name for the donkey is Equus africanus asinus when it is considered a subspecies. At one time, the ass was the more common term for the donkey. The first recorded use of donkey was in either 1784 or 1785, while the word ass has cognates in most other Indo-European languages, donkey is an etymologically obscure word for which no credible cognate has been identified. From the 18th century, donkey gradually replaced ass, and jenny replaced she-ass, by the end of the 17th century, changes in pronunciation of both ass and arse had caused them to become homophones. Donkeys vary considerably in size, depending on breed and management, the height at the withers ranges from 7.3 to 15.3 hands, and the weight from 80 to 480 kg. Working donkeys in the poorest countries have an expectancy of 12 to 15 years, in more prosperous countries. Donkeys are adapted to desert lands. Unlike wild and feral horses, wild donkeys in dry areas are solitary, each adult donkey establishes a home range, breeding over a large area may be dominated by one jack. Donkeys have large ears, which may pick up more distant sounds, donkeys can defend themselves by biting, striking with the front hooves or kicking with the hind legs. A jenny is normally pregnant for about 12 months, though the period varies from 11 to 14 months
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Kiang
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The kiang is the largest of the wild asses. It is native to the Tibetan Plateau, where it inhabits montane and alpine grasslands and its current range is restricted to Ladakh in Jammu and Kashmir, plains of the Tibetan plateau and northern Nepal along the Tibetan border. Other common names for species include Tibetan wild ass, khyang. The kiang is closely related to the onager, and in some classifications it is considered a subspecies, molecular studies, however, indicate that it is a distinct species. Kiangs can crossbreed with onagers, horses, donkeys, and Burchells zebras in captivity, although, like mules, the kiang is the largest of the wild asses, with an average shoulder height of 13.3 hands. They range from 132 to 142 cm high at the shoulder, with a body 182 to 214 cm long, kiangs have only slight sexual dimorphism, with the males weighing from 350 to 400 kg, while females weigh 250 to 300 kg. They have a head, with a blunt muzzle and a convex nose. The mane is upright and relatively short, the coat is a rich chestnut colour, darker brown in winter and a sleek reddish brown in late summer, when the animal moults its woolly fur. The summer coat is 1.5 cm long and the coat is double that length. The legs, underparts, end of the muzzle, and the inside of the ears are all white, a broad, dark chocolate-coloured dorsal stripe extends from the mane to the end of the tail, which ends in a tuft of blackish brown hairs. Kiangs are found on the Tibetan Plateau, between the Himalayas in the south and the Kunlun Mountains in the north. This restricts them almost entirely to China, but small numbers are found across the borders in the Ladakh and Sikkim regions of India, the western kiang is slightly smaller than the eastern and also has a darker coat. However, no genetic information confirms the validity of the three subspecies, which may represent a cline, with gradual variation between the three forms. Kiangs inhabit alpine meadows and steppe country between 2,700 and 5,300 m elevation and they prefer relatively flat plateaus, wide valleys, and low hills, dominated by grasses, sedges, and smaller amounts of other low-lying vegetation. This open terrain, in addition to supplying them with suitable forage absent in the arid regions of central Asia, may make it easier for them to detect. Like all equids, kiangs are herbivores, feeding on grasses and sedges, especially Stipa, but also including other plants such as bog sedges, true sedges. When little grass is available, such as during winter or in the more arid margins of their habitat, they have been observed eating shrubs, herbs. Their only real predator other than humans is the wolf, kiangs defend themselves by forming a circle, and with heads down, kick out violently
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Israel
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Israel, officially the State of Israel, is a country in the Middle East, on the southeastern shore of the Mediterranean Sea and the northern shore of the Red Sea. The country contains geographically diverse features within its small area. Israels economy and technology center is Tel Aviv, while its seat of government and proclaimed capital is Jerusalem, in 1947, the United Nations adopted a Partition Plan for Mandatory Palestine recommending the creation of independent Arab and Jewish states and an internationalized Jerusalem. The plan was accepted by the Jewish Agency for Palestine, next year, the Jewish Agency declared the establishment of a Jewish state in Eretz Israel, to be known as the State of Israel. Israel has since fought several wars with neighboring Arab states, in the course of which it has occupied territories including the West Bank, Golan Heights and it extended its laws to the Golan Heights and East Jerusalem, but not the West Bank. Israels occupation of the Palestinian territories is the worlds longest military occupation in modern times, efforts to resolve the Israeli–Palestinian conflict have not resulted in peace. However, peace treaties between Israel and both Egypt and Jordan have successfully been signed, the population of Israel, as defined by the Israel Central Bureau of Statistics, was estimated in 2017 to be 8,671,100 people. It is the worlds only Jewish-majority state, with 74. 8% being designated as Jewish, the countrys second largest group of citizens are Arabs, at 20. 8%. The great majority of Israeli Arabs are Sunni Muslims, including significant numbers of semi-settled Negev Bedouins, other minorities include Arameans, Armenians, Assyrians, Black Hebrew Israelites, Circassians, Maronites and Samaritans. Israel also hosts a significant population of foreign workers and asylum seekers from Africa and Asia, including illegal migrants from Sudan, Eritrea. In its Basic Laws, Israel defines itself as a Jewish, Israel is a representative democracy with a parliamentary system, proportional representation and universal suffrage. The prime minister is head of government and the Knesset is the legislature, Israel is a developed country and an OECD member, with the 35th-largest economy in the world by nominal gross domestic product as of 2016. The country benefits from a skilled workforce and is among the most educated countries in the world with one of the highest percentage of its citizens holding a tertiary education degree. The country has the highest standard of living in the Middle East and the third highest in Asia, in the early weeks of independence, the government chose the term Israeli to denote a citizen of Israel, with the formal announcement made by Minister of Foreign Affairs Moshe Sharett. The names Land of Israel and Children of Israel have historically used to refer to the biblical Kingdom of Israel. The name Israel in these phrases refers to the patriarch Jacob who, jacobs twelve sons became the ancestors of the Israelites, also known as the Twelve Tribes of Israel or Children of Israel. The earliest known artifact to mention the word Israel as a collective is the Merneptah Stele of ancient Egypt. The area is known as the Holy Land, being holy for all Abrahamic religions including Judaism, Christianity, Islam
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Saudi Arabia
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Saudi Arabia is bordered by Jordan and Iraq to the north, Kuwait to the northeast, Qatar, Bahrain and the United Arab Emirates to the east, Oman to the southeast and Yemen to the south. It is separated from Israel and Egypt by the Gulf of Aqaba and it is the only nation with both a Red Sea coast and a Persian Gulf coast and most of its terrain consists of arid desert and mountains. The area of modern-day Saudi Arabia formerly consisted of four regions, Hejaz, Najd and parts of Eastern Arabia. The Kingdom of Saudi Arabia was founded in 1932 by Ibn Saud and he united the four regions into a single state through a series of conquests beginning in 1902 with the capture of Riyadh, the ancestral home of his family, the House of Saud. Saudi Arabia has since been a monarchy, effectively a hereditary dictatorship governed along Islamic lines. The ultraconservative Wahhabi religious movement within Sunni Islam has been called the predominant feature of Saudi culture, with its global spread largely financed by the oil and gas trade. Saudi Arabia is sometimes called the Land of the Two Holy Mosques in reference to Al-Masjid al-Haram and Al-Masjid an-Nabawi, the state has a total population of 28.7 million, of which 20 million are Saudi nationals and 8 million are foreigners. The states official language is Arabic, petroleum was discovered on 3 March 1938 and followed up by several other finds in the Eastern Province. Saudi Arabia has since become the worlds largest oil producer and exporter, controlling the second largest oil reserves. The kingdom is categorized as a World Bank high-income economy with a high Human Development Index and is the only Arab country to be part of the G-20 major economies. However, the economy of Saudi Arabia is the least diversified in the Gulf Cooperation Council, the state has attracted criticism for its treatment of women and use of capital punishment. Saudi Arabia is an autocracy, has the fourth highest military expenditure in the world. Saudi Arabia is considered a regional and middle power, in addition to the GCC, it is an active member of the Organisation of Islamic Cooperation and OPEC. Following the unification of the Hejaz and Nejd kingdoms, the new state was named al-Mamlakah al-ʻArabīyah as-Suʻūdīyah by royal decree on 23 September 1932 by its founder, Abdulaziz Al Saud. Although this is translated as the Kingdom of Saudi Arabia in English it literally means the Saudi Arab kingdom. Its inclusion expresses the view that the country is the possession of the royal family. Al Saud is an Arabic name formed by adding the word Al, meaning family of or House of, in the case of the Al Saud, this is the father of the dynastys 18th century founder, Muhammad bin Saud. There is evidence that human habitation in the Arabian Peninsula dates back to about 125,000 years ago
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