1.
Taxonomy (biology)
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Taxonomy is the science of defining groups of biological organisms on the basis of shared characteristics and giving names to those groups. The exact definition of taxonomy varies from source to source, but the core of the remains, the conception, naming. There is some disagreement as to whether biological nomenclature is considered a part of taxonomy, the broadest meaning of taxonomy is used here. The word taxonomy was introduced in 1813 by Candolle, in his Théorie élémentaire de la botanique, the term alpha taxonomy is primarily used today to refer to the discipline of finding, describing, and naming taxa, particularly species. In earlier literature, the term had a different meaning, referring to morphological taxonomy, ideals can, it may be said, never be completely realized. They have, however, a value of acting as permanent stimulants. Some of us please ourselves by thinking we are now groping in a beta taxonomy, turrill thus explicitly excludes from alpha taxonomy various areas of study that he includes within taxonomy as a whole, such as ecology, physiology, genetics, and cytology. He further excludes phylogenetic reconstruction from alpha taxonomy, thus, Ernst Mayr in 1968 defined beta taxonomy as the classification of ranks higher than species. This activity is what the term denotes, it is also referred to as beta taxonomy. How species should be defined in a group of organisms gives rise to practical and theoretical problems that are referred to as the species problem. The scientific work of deciding how to define species has been called microtaxonomy, by extension, macrotaxonomy is the study of groups at higher taxonomic ranks, from subgenus and above only, than species. While some descriptions of taxonomic history attempt to date taxonomy to ancient civilizations, earlier works were primarily descriptive, and focused on plants that were useful in agriculture or medicine. There are a number of stages in scientific thinking. Early taxonomy was based on criteria, the so-called artificial systems. Later came systems based on a complete consideration of the characteristics of taxa, referred to as natural systems, such as those of de Jussieu, de Candolle and Bentham. The publication of Charles Darwins Origin of Species led to new ways of thinking about classification based on evolutionary relationships and this was the concept of phyletic systems, from 1883 onwards. This approach was typified by those of Eichler and Engler, the advent of molecular genetics and statistical methodology allowed the creation of the modern era of phylogenetic systems based on cladistics, rather than morphology alone. Taxonomy has been called the worlds oldest profession, and naming and classifying our surroundings has likely been taking place as long as mankind has been able to communicate
2.
Plant
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Plants are mainly multicellular, predominantly photosynthetic eukaryotes of the kingdom Plantae. The term is generally limited to the green plants, which form an unranked clade Viridiplantae. This includes the plants, conifers and other gymnosperms, ferns, clubmosses, hornworts, liverworts, mosses and the green algae. Green plants have cell walls containing cellulose and obtain most of their energy from sunlight via photosynthesis by primary chloroplasts and their chloroplasts contain chlorophylls a and b, which gives them their green color. Some plants are parasitic and have lost the ability to produce amounts of chlorophyll or to photosynthesize. Plants are characterized by sexual reproduction and alternation of generations, although reproduction is also common. There are about 300–315 thousand species of plants, of which the great majority, green plants provide most of the worlds molecular oxygen and are the basis of most of Earths ecologies, especially on land. Plants that produce grains, fruits and vegetables form humankinds basic foodstuffs, Plants play many roles in culture. They are used as ornaments and, until recently and in variety, they have served as the source of most medicines. The scientific study of plants is known as botany, a branch of biology, Plants are one of the two groups into which all living things were traditionally divided, the other is animals. The division goes back at least as far as Aristotle, who distinguished between plants, which generally do not move, and animals, which often are mobile to catch their food. Much later, when Linnaeus created the basis of the system of scientific classification. Since then, it has become clear that the plant kingdom as originally defined included several unrelated groups, however, these organisms are still often considered plants, particularly in popular contexts. When the name Plantae or plant is applied to a group of organisms or taxon. The evolutionary history of plants is not yet settled. Those which have been called plants are in bold, the way in which the groups of green algae are combined and named varies considerably between authors. Algae comprise several different groups of organisms which produce energy through photosynthesis, most conspicuous among the algae are the seaweeds, multicellular algae that may roughly resemble land plants, but are classified among the brown, red and green algae. Each of these groups also includes various microscopic and single-celled organisms
3.
Flowering plant
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The flowering plants, also known as Angiospermae or Magnoliophyta, are the most diverse group of land plants, with 416 families, approx. 13,164 known genera and a total of c.295,383 known species, etymologically, angiosperm means a plant that produces seeds within an enclosure, in other words, a fruiting plant. The term angiosperm comes from the Greek composite word meaning enclosed seeds, the ancestors of flowering plants diverged from gymnosperms in the Triassic Period, during the range 245 to 202 million years ago, and the first flowering plants are known from 160 mya. They diversified extensively during the Lower Cretaceous, became widespread by 120 mya, angiosperms differ from other seed plants in several ways, described in the table. These distinguishing characteristics taken together have made the angiosperms the most diverse and numerous land plants, the amount and complexity of tissue-formation in flowering plants exceeds that of gymnosperms. The vascular bundles of the stem are arranged such that the xylem and phloem form concentric rings, in the dicotyledons, the bundles in the very young stem are arranged in an open ring, separating a central pith from an outer cortex. In each bundle, separating the xylem and phloem, is a layer of meristem or active formative tissue known as cambium, the soft phloem becomes crushed, but the hard wood persists and forms the bulk of the stem and branches of the woody perennial. Among the monocotyledons, the bundles are more numerous in the stem and are scattered through the ground tissue. They contain no cambium and once formed the stem increases in diameter only in exceptional cases, the characteristic feature of angiosperms is the flower. Flowers show remarkable variation in form and elaboration, and provide the most trustworthy external characteristics for establishing relationships among angiosperm species, the function of the flower is to ensure fertilization of the ovule and development of fruit containing seeds. The floral apparatus may arise terminally on a shoot or from the axil of a leaf, occasionally, as in violets, a flower arises singly in the axil of an ordinary foliage-leaf. There are two kinds of cells produced by flowers. Microspores, which divide to become pollen grains, are the male cells and are borne in the stamens. The female cells called megaspores, which divide to become the egg cell, are contained in the ovule. The flower may consist only of parts, as in willow. Usually, other structures are present and serve to protect the sporophylls, the individual members of these surrounding structures are known as sepals and petals. The outer series is usually green and leaf-like, and functions to protect the rest of the flower, the inner series is, in general, white or brightly colored, and is more delicate in structure. It functions to attract insect or bird pollinators, attraction is effected by color, scent, and nectar, which may be secreted in some part of the flower
4.
Monocotyledon
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Monocotyledons, commonly referred to as monocots, are flowering plants whose seeds typically contain only one embryonic leaf, or cotyledon. However, molecular research has shown that while the monocots form a monophyletic group or clade. Monocots have almost always recognized as a group, but with various taxonomic ranks. The APG III system of 2009 recognises a clade called monocots, the monocots include about 60,000 species. The largest family in this group by number of species are the orchids, about half as many species belong to the true grasses, which are economically the most important family of monocots. In agriculture the majority of the biomass produced comes from monocots and these include not only major grains, but also forage grasses, sugar cane, and the bamboos. Other economically important monocot crops include various palms, bananas, gingers and their relatives, turmeric and cardamom, asparagus and the onions and garlic family. Additionally most of the bulbs, plants cultivated for their blooms, such as lilies, daffodils, irises, amaryllis, cannas, bluebells. The monocots or monocotyledons have, as the name implies, a single cotyledon, or embryonic leaf, from a diagnostic point of view the number of cotyledons is neither a particularly useful characteristic, nor is it completely reliable. Nevertheless, monocots are sufficiently distinctive that there has rarely been disagreement as to membership of this group, however, morphological features that reliably characterise major clades are rare. Thus monocots are distinguishable from other angiosperms both in terms of their uniformity and diversity, although largely herbaceous, some arboraceous monocots reach great height, length and mass. The latter include agaves, palms, pandans, and bamboos and this creates challenges in water transport that monocots deal with in various ways. Some such as species of Yucca develop anomalous secondary growth, while palm trees, the axis undergoes primary thickening, that progresses from internode to internode, resulting in a typical inverted conical shape of the basal primary axis. The limited conductivity also contributes to limited branching of the stems, despite these limitations a wide variety of adaptive growth forms has resulted from epiphytic orchids and bromeliads to submarine Alismatales and mycotrophic Burmanniaceae and Triuridaceae. Other monocots, particularly Poales have adopted a life form. Leaves The cotyledon, the primordial Angiosperm leaf consists of a proximal leaf base or hypophyll, in moncots the hypophyll tends to be the dominant part in contrast to other angiosperms. Mature monocot leaves are narrow and linear, forming a sheathing around the stem at its base. There is usually only one leaf per node because the leaf base encompasses more than half the circumference, the evolution of this monocot characteristic has been attributed to developmental differences in early zonal differentiation rather than meristem activity
5.
Asparagales
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Asparagales is an order of plants in modern classification systems such as the Angiosperm Phylogeny Group and the Angiosperm Phylogeny Web. The order takes its name from the type family Asparagaceae and is placed in the monocots amongst the lilioid monocots, the order has only recently been recognized in classification systems. It was first put forward by Huber in 1977 and later taken up in the Dahlgren system of 1985 and then the APG in 1998,2003 and 2009. Before this, many of its families were assigned to the old order Liliales, DNA sequence analysis indicated that many of the taxa previously included in Liliales should actually be redistributed over three orders, Liliales, Asparagales and Dioscoreales. The boundaries of the Asparagales and of its families have undergone a series of changes in recent years, future research may lead to further changes and ultimately greater stability. In the APG circumscription, Asparagales is the largest order of monocots with 14 families,1,122 genera, the order is clearly circumscribed on the basis of molecular phylogenetics, but is difficult to define morphologically, since its members are structurally diverse. Most species of Asparagales are herbaceous perennials, although some are climbers, the order also contains many geophytes. According to telomere sequence, at least two evolutionary switch-points happened within the order, basal sequence is formed by TTTAGGG like in majority of higher plants. Basal motif was changed to vertebrate-like TTAGGG and finally the most divergent motif CTCGGTTATGGG appears in Allium, one of the defining characteristics of the order is the presence of phytomelanin, a black pigment present in the seed coat, creating a dark crust. Phytomelanin is found in most families of the Asparagales, the leaves of almost all species form a tight rosette, either at the base of the plant or at the end of the stem, but occasionally along the stem. The flowers are not particularly distinctive, being lily type, with six tepals, from an economic point of view, the order Asparagales is second in importance within the monocots to the order Poales. Species are used as food and flavourings, as cut flowers, and as garden ornamentals. Thus although most species in the order are herbaceous, some no more than 15 cm high, there are a number of climbers, as well as several genera forming trees, succulent genera occur in several families. Almost all species have a cluster of leaves, either at the base of the plant or at the end of a more-or-less woody stem as with Yucca. In some cases the leaves are produced along the stem, the flowers are in the main not particularly distinctive, being of a general lily type, with six tepals, either free or fused from the base and up to six stamina. They are frequently clustered at the end of the plant stem and they are generally geophytes, but with linear leaves, and a lack of fine reticular venation. The seeds characteristically have the external epidermis either obliterated, or if present, have a layer of black carbonaceous phytomelanin in species with dry fruits. The inner part of the coat is generally collapsed, in contrast to Liliales whose seeds have a well developed outer epidermis, lack phytomelanin
6.
Orchidaceae
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The Orchidaceae are a diverse and widespread family of flowering plants, with blooms that are often colourful and often fragrant, commonly known as the orchid family. Along with the Asteraceae, they are one of the two largest families of flowering plants, the Orchidaceae have about 28,000 currently accepted species, distributed in about 763 genera. The determination of which family is larger is still under debate, regardless, the number of orchid species nearly equals the number of bony fishes and is more than twice the number of bird species, and about four times the number of mammal species. The family also encompasses about 6–11% of all seed plants, the largest genera are Bulbophyllum, Epidendrum, Dendrobium and Pleurothallis. The family also includes Vanilla, Orchis, and many cultivated plants such as Phalaenopsis. Moreover, since the introduction of species into cultivation in the 19th century, horticulturists have produced more than 100,000 hybrids. Orchids are easily distinguished from plants, as they share some very evident, shared derived characteristics. Among these are, bilateral symmetry of the flower, many flowers, a nearly always highly modified petal, fused stamens and carpels. All orchids are perennial herbs that lack any permanent woody structure and they can grow according to two patterns, Monopodial, The stem grows from a single bud, leaves are added from the apex each year and the stem grows longer accordingly. The stem of orchids with a monopodial growth can reach several metres in length, as in Vanda, Sympodial, Sympodial orchids have a front and a back. The plant produces a series of adjacent shoots which grow to a size, bloom. Sympodial orchids grow laterally rather than vertically, following the surface of their support, the growth continues by development of new leads, with their own leaves and roots, sprouting from or next to those of the previous year, as in Cattleya. While a new lead is developing, the rhizome may start its growth again from a so-called eye, Sympodial orchids may have visible pseudobulbs joined by a rhizome, which creeps along the top or just beneath the soil. Terrestrial orchids may be rhizomatous or form corms or tubers, the root caps of terrestrial orchids are smooth and white. Some sympodial terrestrial orchids, such as Orchis and Ophrys, have two subterranean tuberous roots, one is used as a food reserve for wintry periods, and provides for the development of the other one, from which visible growth develops. In warm and constantly humid climates, many terrestrial orchids do not need pseudobulbs, epiphytic orchids, those that grow upon a support, have modified aerial roots that can sometimes be a few meters long. In the older parts of the roots, a modified spongy epidermis and it is made of dead cells and can have a silvery-grey, white or brown appearance. In some orchids, the velamen includes spongy and fibrous bodies near the passage cells, the cells of the root epidermis grow at a right angle to the axis of the root to allow them to get a firm grasp on their support
7.
Orchidoideae
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The Orchidoideae, or the orchidoid orchids, are a subfamily of the orchid family. They typically contain the orchids with a single, fertile anther which is erect and it is sister to the tribe Diurideae. l. BATEMAN, PETER M. HOLLINGSWORTH, JILLIAN PRESTON, LUO YI-BO, ALEC M. PRIDGEON, molecular phylogenetics and evolution of Orchidinae and selected Habenariinae. Botanical Journal of the Linnean Society, Oxford University Press, Oxford, U. K. Salazar, G. A. Chase, M. A. Soto Arenas and M. Ingrouille, phylogenetics of Cranichideae with emphasis on Spiranthinae, evidence from plastid and nuclear DNA sequences. American Journal of Botany 90, 777-795, Phylogenetic relationships of Cranichidinae and Prescottiinae inferred from plastid and nuclear DNA sequences. Chemisquy, M. A. Morrone, O. Phylogenetic analysis of the subtribe Chloraeinae, a preliminary approach based on three chloroplast markers
8.
Carl Linnaeus
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Carl Linnaeus, also known after his ennoblement as Carl von Linné, was a Swedish botanist, physician, and zoologist, who formalised the modern system of naming organisms called binomial nomenclature. He is known by the father of modern taxonomy. Many of his writings were in Latin, and his name is rendered in Latin as Carolus Linnæus, Linnaeus was born in the countryside of Småland, in southern Sweden. He received most of his education at Uppsala University. He lived abroad between 1735 and 1738, where he studied and also published a first edition of his Systema Naturae in the Netherlands and he then returned to Sweden, where he became professor of medicine and botany at Uppsala. In the 1740s, he was sent on journeys through Sweden to find and classify plants. In the 1750s and 1760s, he continued to collect and classify animals, plants, and minerals, at the time of his death, he was one of the most acclaimed scientists in Europe. The philosopher Jean-Jacques Rousseau sent him the message, Tell him I know no man on earth. The German writer Johann Wolfgang von Goethe wrote, With the exception of Shakespeare and Spinoza, Swedish author August Strindberg wrote, Linnaeus was in reality a poet who happened to become a naturalist. Among other compliments, Linnaeus has been called Princeps botanicorum, The Pliny of the North and he is also considered as one of the founders of modern ecology. In botany, the abbreviation used to indicate Linnaeus as the authority for species names is L. In older publications, sometimes the abbreviation Linn. is found, Linnæus was born in the village of Råshult in Småland, Sweden, on 23 May 1707. He was the first child of Nicolaus Ingemarsson and Christina Brodersonia and his siblings were Anna Maria Linnæa, Sofia Juliana Linnæa, Samuel Linnæus, and Emerentia Linnæa. One of a line of peasants and priests, Nils was an amateur botanist, a Lutheran minister. Christina was the daughter of the rector of Stenbrohult, Samuel Brodersonius, a year after Linnæus birth, his grandfather Samuel Brodersonius died, and his father Nils became the rector of Stenbrohult. The family moved into the rectory from the curates house, even in his early years, Linnæus seemed to have a liking for plants, flowers in particular. Whenever he was upset, he was given a flower, which calmed him. Nils spent much time in his garden and often showed flowers to Linnaeus, soon Linnæus was given his own patch of earth where he could grow plants
9.
Type species
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A similar concept is used for suprageneric groups called a type genus. In botanical nomenclature, these terms have no standing under the code of nomenclature. In botany, the type of a name is a specimen which is also the type of a species name. The species name that has that type can also be referred to as the type of the genus name, names of genus and family ranks, the various subdivisions of those ranks, and some higher-rank names based on genus names, have such types. In bacteriology, a species is assigned for each genus. Every named genus or subgenus in zoology, whether or not currently recognized as valid, is associated with a type species. In practice, however, there is a backlog of untypified names defined in older publications when it was not required to specify a type, a type species is both a concept and a practical system that is used in the classification and nomenclature of animals. The type species represents the species and thus definition for a particular genus name. In the Glossary, type species is defined as The nominal species that is the type of a nominal genus or subgenus. The type species permanently attaches a formal name to a genus by providing just one species within that genus to which the name is permanently linked. The species name in turn is fixed, in theory, to a type specimen, for example, the type species for the land snail genus Monacha is Monacha cartusiana. That genus is placed within the family Hygromiidae. The type genus for that family is the genus Hygromia, the concept of the type species in zoology was introduced by Pierre André Latreille. The International Code of Zoological Nomenclature states that the name of the type species should always be cited. It gives an example in Article 67.1, Astacus marinus Fabricius,1775 was later designated as the type species of the genus Homarus, thus giving it the name Homarus marinus. However, the species of Homarus should always be cited using its original name. 3, the type of the genus name Elodes is quoted as the type of the species name Hypericum aegypticum, Glossary of scientific naming Genetypes – genetic sequence data from type specimens. Holotype Paratype Principle of Typification Type Type genus
10.
Ophrys insectifera
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Ophrys insectifera, the fly orchid, is a species of orchid and the type species of the genus Ophrys. It is native to Europe and favors sites with alkaline soil, the name arises because its inflorescence resembles a fly, although it is dependent on wasps and bees for pollination. The plants use scent to attract male wasps and bees which pollinate the flowers as they attempt to mate with the flower, the scent released by the flowers mimic female sexual pheromones. Claessens, J. Kleynen, European orchids - Argogorytes mystaceus pollinator of Ophrys insectifera IUCN Red List description
11.
Synonym (taxonomy)
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For example, Linnaeus was the first to give a scientific name to the Norway spruce, which he called Pinus abies. This name is no longer in use, it is now a synonym of the current scientific name which is Picea abies, unlike synonyms in other contexts, in taxonomy a synonym is not interchangeable with the name of which it is a synonym. In taxonomy, synonyms are not equals, but have a different status, for any taxon with a particular circumscription, position, and rank, only one scientific name is considered to be the correct one at any given time. A synonym cannot exist in isolation, it is always an alternative to a different scientific name, given that the correct name of a taxon depends on the taxonomic viewpoint used a name that is one taxonomists synonym may be another taxonomists correct name. Synonyms may arise whenever the same taxon is described and named more than once, independently. They may also arise when existing taxa are changed, as when two taxa are joined to one, a species is moved to a different genus. To the general user of scientific names, in such as agriculture, horticulture, ecology, general science. A synonym is a name that was used as the correct scientific name but which has been displaced by another scientific name. Thus Oxford Dictionaries Online defines the term as a name which has the same application as another. In handbooks and general texts, it is useful to have mentioned as such after the current scientific name. Synonyms used in this way may not always meet the strict definitions of the synonym in the formal rules of nomenclature which govern scientific names. Changes of scientific name have two causes, they may be taxonomic or nomenclatural, a name change may be caused by changes in the circumscription, position or rank of a taxon, representing a change in taxonomic, scientific insight. A name change may be due to purely nomenclatural reasons, that is, based on the rules of nomenclature, the earliest such name is called the senior synonym, while the later name is the junior synonym. One basic principle of zoological nomenclature is that the earliest correctly published name, synonyms are important because if the earliest name cannot be used, then the next available junior synonym must be used for the taxon. Objective synonyms refer to taxa with the type and same rank. For example, John Edward Gray published the name Antilocapra anteflexa in 1855 for a species of pronghorn, however, it is now commonly accepted that his specimen was an unusual individual of the species Antilocapra americana published by George Ord in 1815. Ords name thus takes precedence, with Antilocapra anteflexa being a subjective synonym. Objective synonyms are common at the level of genera, because for various reasons two genera may contain the type species, these are objective synonyms
12.
Orchis
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Orchis is a genus in the orchid family, occurring mainly in Europe and Northwest Africa, and ranging as far as Tibet, Mongolia, and Xinjiang. The name is from the Ancient Greek ὄρχις orchis, meaning testicle and these terrestrial orchids have root tubers instead of pseudobulbs. They are extremely diverse in appearance, the inflorescence is a cylindrical to globular spike, 5–15 cm long, with yellow, red to purple flowers. They start flowering at the base, slowly progressing upwards, except for the Monkey orchid, the original genus Orchis used to contain more than 1,300 names. Since it was polyphyletic, it has been divided by Pridgeon et al. into several new genera, Ponerorchis, Schizodium, as of June 2014, the World Checklist of Selected Plant Families accepts 21 species, along with a number of subspecies, Orchis adenocheila Czerniak. – Anatolian Orchid Orchis anthropophora All, – Man Orchid, Italian Orchid Orchis laeta Steinh. – Early Purple Orchid, Orchis mascula subsp, – Military Orchid Orchis militaris subsp. Orchis pallens L. – Pale-flowered Orchid Orchis patens Desf, – Small-dotted Orchid Orchis purpurea Huds. – Lady orchid, Purple Orchid Orchis purpurea subsp, purpurea Orchis quadripunctata Cirillo ex Ten. – Four-spotted Orchid Orchis simia Lam, – Monkey Orchid Orchis simia subsp. Taubertiana Kreutz Orchis sitiaca P. Delforge Orchis spitzelii Saut, ex W. D. J. Koch – Spitzels Orchid Orchis spitzelii subsp. Cazorlensis D. Rivera & Lopez Velez Orchis spitzelii subsp, Orchis × apollinaris W. Rossi & al. Orchis × aurunca W. Rossi & Minut, Orchis × bergonii Nanteuil Orchis × beyrichii A. Kern. Orchis × bispurium H. Kretzschmar, Eccarius & H. Dietr, Orchis × caesii De Angelis & Fumanti Orchis × calliantha Renz & Taubenheim Orchis × chabalensis B. Baumann & al. Orchis × colemanii Cortesi Orchis × fallax Willk, Orchis × klopfensteiniae P. Delforge Orchis × kretzschmariorum B. Baumann & H. Baumann Orchis × ligustica Ruppert Orchis × loreziana Brügger Orchis × loreziana nothosubsp. Kisslingii Potucek Orchis × loreziana nothosubsp, loreziana Orchis × lucensis Antonetti & Bertolini Orchis × macra Lindl. Orchis × orphanidesii B. Bock Orchis × palanchonii G. Foelsche & W. Foelsche Orchis × penzigiana A. Camus Orchis × penzigiana nothosubsp, sardoa Scrugli & M. P. Grasso Orchis × permixta Soó Orchis × petterssonii G. Keller ex Pett. Petterssonii Orchis × plessidiaca Renz Orchis × pseudoanatolica H. Fleischm, Orchis × schebestae Griebl Orchis × serraniana P. Delforge Orchis × sezikiana B. Baumann & H. Baumann Orchis × spuria Rchb. f
13.
Europe
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Europe is a continent that comprises the westernmost part of Eurasia. Europe is bordered by the Arctic Ocean to the north, the Atlantic Ocean to the west, yet the non-oceanic borders of Europe—a concept dating back to classical antiquity—are arbitrary. Europe covers about 10,180,000 square kilometres, or 2% of the Earths surface, politically, Europe is divided into about fifty sovereign states of which the Russian Federation is the largest and most populous, spanning 39% of the continent and comprising 15% of its population. Europe had a population of about 740 million as of 2015. Further from the sea, seasonal differences are more noticeable than close to the coast, Europe, in particular ancient Greece, was the birthplace of Western civilization. The fall of the Western Roman Empire, during the period, marked the end of ancient history. Renaissance humanism, exploration, art, and science led to the modern era, from the Age of Discovery onwards, Europe played a predominant role in global affairs. Between the 16th and 20th centuries, European powers controlled at times the Americas, most of Africa, Oceania. The Industrial Revolution, which began in Great Britain at the end of the 18th century, gave rise to economic, cultural, and social change in Western Europe. During the Cold War, Europe was divided along the Iron Curtain between NATO in the west and the Warsaw Pact in the east, until the revolutions of 1989 and fall of the Berlin Wall. In 1955, the Council of Europe was formed following a speech by Sir Winston Churchill and it includes all states except for Belarus, Kazakhstan and Vatican City. Further European integration by some states led to the formation of the European Union, the EU originated in Western Europe but has been expanding eastward since the fall of the Soviet Union in 1991. The European Anthem is Ode to Joy and states celebrate peace, in classical Greek mythology, Europa is the name of either a Phoenician princess or of a queen of Crete. The name contains the elements εὐρύς, wide, broad and ὤψ eye, broad has been an epithet of Earth herself in the reconstructed Proto-Indo-European religion and the poetry devoted to it. For the second part also the divine attributes of grey-eyed Athena or ox-eyed Hera. The same naming motive according to cartographic convention appears in Greek Ανατολή, Martin Litchfield West stated that phonologically, the match between Europas name and any form of the Semitic word is very poor. Next to these there is also a Proto-Indo-European root *h1regʷos, meaning darkness. Most major world languages use words derived from Eurṓpē or Europa to refer to the continent, in some Turkic languages the originally Persian name Frangistan is used casually in referring to much of Europe, besides official names such as Avrupa or Evropa
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North Africa
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North Africa or Northern Africa is the northernmost region of Africa. The United Nationss definition of Northern Africa is, Algeria, Egypt, Libya, Morocco, Sudan, Tunisia, the countries of Algeria, Morocco, Tunisia, and Libya are often collectively referred to as the Maghreb, which is the Arabic word for sunset. Egypt lies to the northeast and encompasses part of West Asia, while Sudan is situated on the edge of the Sahel, Egypt is a transcontinental country because of the Sinai Peninsula, which geographically lies in Western Asia. North Africa also includes a number of Spanish possessions, the Canary Islands and Madeira in the North Atlantic Ocean northwest of the African mainland are included in considerations of the region. From 3500 BC, following the abrupt desertification of the Sahara due to changes in the Earths orbit. The Islamic influence in the area is significant, and North Africa is a major part of the Muslim world. Some researchers have postulated that North Africa rather than East Africa served as the point for the modern humans who first trekked out of the continent in the Out of Africa migration. The Atlas Mountains extend across much of Morocco, northern Algeria and Tunisia, are part of the mountain system that also runs through much of Southern Europe. They recede to the south and east, becoming a steppe landscape before meeting the Sahara desert, the sediments of the Sahara overlie an ancient plateau of crystalline rock, some of which is more than four billion years old. Sheltered valleys in the Atlas Mountains, the Nile Valley and Delta, a wide variety of valuable crops including cereals, rice and cotton, and woods such as cedar and cork, are grown. Typical Mediterranean crops, such as olives, figs, dates and citrus fruits, the Nile Valley is particularly fertile, and most of the population in Egypt and Sudan live close to the river. Elsewhere, irrigation is essential to improve yields on the desert margins. The inhabitants of Saharan Africa are generally divided in a manner corresponding to the principal geographic regions of North Africa, the Maghreb, the Nile valley. The edge of the Sahel, to the south of Egypt has mainly been inhabited by Nubians, Ancient Egyptians record extensive contact in their Western desert with people that appear to have been Berber or proto-Berber, as well as Nubians from the south. They have contributed to the Arabized Berber populations, the official language or one of the official languages in all of the countries in North Africa is Arabic. The people of the Maghreb and the Sahara regions speak Berber languages and several varieties of Arabic, the Arabic and Berber languages are distantly related, both being members of the Afroasiatic language family. The Tuareg Berber languages are more conservative than those of the coastal cities. Over the years, Berbers have been influenced by contact with cultures, Greeks, Phoenicians, Egyptians, Romans, Vandals, Arabs, Europeans
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Canary Islands
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The Canary Islands, also known as the Canaries, are an archipelago and autonomous community of Spain located on the Atlantic Ocean,100 kilometres west of Morocco. The Canaries are among the outermost regions of the European Union proper and it is also one of the eight regions with special consideration of historical nationality recognized as such by the Spanish Government. The main islands are Tenerife, Fuerteventura, Gran Canaria, Lanzarote, La Palma, La Gomera, the archipelago also includes a number of islands and islets, La Graciosa, Alegranza, Isla de Lobos, Montaña Clara, Roque del Oeste and Roque del Este. In ancient times, the chain was often referred to as the Fortunate Isles. The Canary Islands is the most southerly region of Spain and the largest and most populated archipelago of the Macaronesia region, the islands have a subtropical climate, with long hot summers and moderately warm winters. The precipitation levels and the level of maritime moderation varies depending on location and elevation, green areas as well as desert exist on the archipelago. Due to their location above the inversion layer, the high mountains of these islands are ideal for astronomical observation. For this reason, two professional observatories, Teide Observatory on the island of Tenerife and Roque de los Muchachos Observatory on the island of La Palma, have built on the islands. Las Palmas de Gran Canaria has been the largest city in the Canaries since 1768, between the 1833 territorial division of Spain and 1927 Santa Cruz de Tenerife was the sole capital of the Canary Islands. In 1927 a decree ordered that the capital of the Canary Islands be shared, the third largest city of the Canary Islands is San Cristóbal de La Laguna on Tenerife. This city is home to the Consejo Consultivo de Canarias. During the time of the Spanish Empire, the Canaries were the main stopover for Spanish galleons on their way to the Americas, who came south to catch the prevailing northeasterly trade winds. The name Islas Canarias is likely derived from the Latin name Canariae Insulae, meaning Islands of the Dogs, according to the historian Pliny the Elder, the Mauretanian king Juba II named the island Canaria because it contained vast multitudes of dogs of very large size. Another speculation is that the dogs were actually a species of monk seal, critically endangered. The dense population of seals may have been the characteristic that most struck the few ancient Romans who established contact with these islands by sea. Alternatively, it is said that the inhabitants of the island, Guanches, used to worship dogs, mummified them. The ancient Greeks also knew about a people, living far to the west, who are the dog-headed ones, who worshipped dogs on an island. Some hypothesize that the Canary Islands dog-worship and the ancient Egyptian cult of the god, Anubis are closely connected
16.
Middle East
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The Middle East is a transcontinental region centered on Western Asia and Egypt. The corresponding adjective is Middle-Eastern and the noun is Middle-Easterner. The term has come into usage as a replacement of the term Near East beginning in the early 20th century. Arabs, Turks, Persians, Kurds, and Azeris constitute the largest ethnic groups in the region by population. Indigenous minorities of the Middle East include Jews, Assyrians and other Arameans, Baloch, Berbers, Copts, Druze, Lurs, Mandaeans, Samaritans, Shabaks, Tats, in the Middle East, there is also a Romani community. European ethnic groups form a diaspora in the region include Albanians, Bosniaks, Circassians, Crimean Tatars, Franco-Levantines. Among other migrant populations are Bengalis as well as other Indians, Chinese, Filipinos, Indonesians, Pakistanis, the history of the Middle East dates back to ancient times, with the importance of the region being recognized for millennia. Most of the countries border the Persian Gulf have vast reserves of crude oil. The term Middle East may have originated in the 1850s in the British India Office, however, it became more widely known when American naval strategist Alfred Thayer Mahan used the term in 1902 to designate the area between Arabia and India. During this time the British and Russian Empires were vying for influence in Central Asia, Mahan realized not only the strategic importance of the region, but also of its center, the Persian Gulf. Mahan first used the term in his article The Persian Gulf and International Relations, published in September 1902 in the National Review, a British journal. The Middle East, if I may adopt a term which I have not seen, will some day need its Malta, as well as its Gibraltar, it does not follow that either will be in the Persian Gulf. The British Navy should have the facility to concentrate in force if occasion arise, about Aden, India, mahans article was reprinted in The Times and followed in October by a 20-article series entitled The Middle Eastern Question, written by Sir Ignatius Valentine Chirol. During this series, Sir Ignatius expanded the definition of Middle East to include regions of Asia which extend to the borders of India or command the approaches to India. After the series ended in 1903, The Times removed quotation marks from subsequent uses of the term, in the late 1930s, the British established the Middle East Command, which was based in Cairo, for its military forces in the region. After that time, the term Middle East gained broader usage in Europe, the description Middle has also led to some confusion over changing definitions. Before the First World War, Near East was used in English to refer to the Balkans and the Ottoman Empire, while Middle East referred to Iran, the Caucasus, Afghanistan, Central Asia, and Turkestan. The first official use of the term Middle East by the United States government was in the 1957 Eisenhower Doctrine, the Associated Press Stylebook says that Near East formerly referred to the farther west countries while Middle East referred to the eastern ones, but that now they are synonymous
17.
Turkmenistan
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Turkmenistan has been at the crossroads of civilizations for centuries. In medieval times, Merv was one of the cities of the Islamic world and an important stop on the Silk Road. Annexed by the Russian Empire in 1881, Turkmenistan later figured prominently in the movement in Central Asia. In 1924, Turkmenistan became a constituent republic of the Soviet Union, Turkmen Soviet Socialist Republic, Turkmenistan possesses the worlds fourth largest reserves of natural gas resources. Most of the country is covered by the Karakum Desert, since 1993, citizens have received government-provided electricity, water and natural gas free of charge. Turkmenistan was ruled by President for Life Saparmurat Niyazov until his death in 2006, Gurbanguly Berdimuhamedow was elected president in 2007. According to Human Rights Watch, Turkmenistan remains one of the world’s most repressive countries, after suspending the death penalty, the use of capital punishment was formally abolished in the 2008 constitution. Historically inhabited by the Indo-Iranians, the history of Turkmenistan begins with its annexation by the Achaemenid Empire of Ancient Iran. In the 8th century AD, Turkic-speaking Oghuz tribes moved from Mongolia into present-day Central Asia, part of a powerful confederation of tribes, these Oghuz formed the ethnic basis of the modern Turkmen population. In the 10th century, the name Turkmen was first applied to Oghuz groups that accepted Islam, There they were under the dominion of the Seljuk Empire, which was composed of Oghuz groups living in present-day Iran and Turkmenistan. Turkmen soldiers in the service of the played a important role in the spreading of Turkic culture when they migrated westward into present-day Azerbaijan. In the 12th century, Turkmen and other tribes overthrew the Seljuk Empire, in the next century, the Mongols took over the more northern lands where the Turkmens had settled, scattering the Turkmens southward and contributing to the formation of new tribal groups. The sixteenth and eighteenth centuries saw a series of splits and confederations among the nomadic Turkmen tribes, by the 16th century, most of those tribes were under the nominal control of two sedentary Uzbek khanates, Khiva and Bukhoro. Turkmen soldiers were an important element of the Uzbek militaries of this period, in the 19th century, raids and rebellions by the Yomud Turkmen group resulted in that groups dispersal by the Uzbek rulers. According to Paul R. Spickard, Prior to the Russian conquest, Russian forces began occupying Turkmen territory late in the 19th century. From their Caspian Sea base at Krasnovodsk, the Russians eventually overcame the Uzbek khanates, in 1916 the Russian Empires participation in World War I resonated in Turkmenistan, as an anticonscription revolt swept most of Russian Central Asia. In 1924 the Turkmen Soviet Socialist Republic was formed from the tsarist province of Transcaspia, by the late 1930s, Soviet reorganization of agriculture had destroyed what remained of the nomadic lifestyle in Turkmenistan, and Moscow controlled political life. The Ashgabat earthquake of 1948 killed over 110,000 people, during the next half-century, Turkmenistan played its designated economic role within the Soviet Union and remained outside the course of major world events
18.
Pseudocopulation
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Pseudocopulation describes behaviors similar to copulation that serve a reproductive function for one or both participants but do not involve actual sexual union between the individuals. It is most generally applied to a pollinator attempting to copulate with a flower, some flowers mimic a potential female mate visually, but the key stimuli are often chemical and tactile. This form of mimicry in plants is called Pouyannian mimicry, orchids commonly achieve reproduction in this manner, secreting chemicals from glands in the sepals, petals, or labellum, that are indistinguishable from the insects natural pheromones. The pollinator then has an attached to its body, which it transfers to the stigma of another flower when it attempts another copulation. Pollinators are often bees and wasps of the order Hymenoptera, the cost to the pollinating insects might be seen as negligible, but study of Cryptostylis pollinators shows that they may waste large amounts of sperm by ejaculating onto the flower. Thus there could be antagonistic coevolution such that pollinators become better at identifying their own species correctly, pseudocopulation is also used to describe close physical contact between mating animals which have their eggs externally fertilized. Frogs provide one such case, with the male releasing sperm as the female discharges her eggs, in some species of starfish, a male and female may come together and form a pair. The male climbs on top, placing his arms between those of the female, when she releases eggs into the water, he is induced to release his sperm. Pseudocopulation is also used as a term to describe behaviours of birds appear to be copulating but may merely involve mounting
19.
Bee
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Bees are flying insects closely related to wasps and ants, known for their role in pollination and, in the case of the best-known bee species, the European honey bee, for producing honey and beeswax. Bees are a lineage within the superfamily Apoidea, presently considered as a clade Anthophila. There are nearly 20,000 known species of bees in seven to nine recognized families, though many are undescribed and they are found on every continent except Antarctica, in every habitat on the planet that contains insect-pollinated flowering plants. Some species including honey bees, bumblebees, and stingless bees live socially in colonies, Bees are adapted for feeding on nectar and pollen, the former primarily as an energy source and the latter primarily for protein and other nutrients. Most pollen is used as food for larvae, Bee pollination is important both ecologically and commercially, the decline in wild bees has increased the value of pollination by commercially managed hives of honey bees. The most common bees in the Northern Hemisphere are the Halictidae, or sweat bees, vertebrate predators of bees include birds such as bee-eaters, insect predators include beewolves and dragonflies. Human beekeeping or apiculture has been practised for millennia, since at least the times of Ancient Egypt, apart from honey and pollination, honey bees produce beeswax, royal jelly and propolis. Bees have appeared in mythology and folklore, again since ancient times, and they feature in works of literature as varied as Virgils Georgics, Beatrix Potters The Tale of Mrs Tittlemouse, yeatss poem The Lake Isle of Innisfree. Bee larvae are included in the Javanese dish botok tawon, where they are steamed with shredded coconut. The ancestors of bees were wasps in the family Crabronidae, which were predators of other insects. The switch from insect prey to pollen may have resulted from the consumption of insects which were flower visitors and were partially covered with pollen when they were fed to the wasp larvae. This same evolutionary scenario may have occurred within the vespoid wasps, until recently, the oldest non-compression bee fossil had been found in New Jersey amber, Cretotrigona prisca of Cretaceous age, a corbiculate bee. A bee fossil from the early Cretaceous, Melittosphex burmensis, is considered an extinct lineage of pollen-collecting Apoidea sister to the modern bees. Derived features of its place it clearly within the bees. By the Eocene there was considerable diversity among eusocial bee lineages. The highly eusocial corbiculate Apidae appeared roughly 87 Mya, and the Allodapini around 53 Mya, the Colletidae appear as fossils only from the late Oligocene to early Miocene. The Melittidae are known from Palaeomacropis eocenicus in the Early Eocene, the Megachilidae are known from trace fossils from the Middle Eocene. The Andrenidae are known from the Eocene-Oligocene boundary, around 34 Mya, the Halictidae first appear in the Early Eocene with species found in amber
20.
Insect
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Insects are a class of hexapod invertebrates within the arthropod phylum that have a chitinous exoskeleton, a three-part body, three pairs of jointed legs, compound eyes and one pair of antennae. They are the most diverse group of animals on the planet, including more than a million described species, the number of extant species is estimated at between six and ten million, and potentially represent over 90% of the differing animal life forms on Earth. Insects may be found in all environments, although only a small number of species reside in the oceans. The life cycles of insects vary but most hatch from eggs, insect growth is constrained by the inelastic exoskeleton and development involves a series of molts. The immature stages can differ from the adults in structure, habit and habitat, Insects that undergo 3-stage metamorphosis lack a pupal stage and adults develop through a series of nymphal stages. The higher level relationship of the Hexapoda is unclear, fossilized insects of enormous size have been found from the Paleozoic Era, including giant dragonflies with wingspans of 55 to 70 cm. The most diverse insect groups appear to have coevolved with flowering plants, adult insects typically move about by walking, flying or sometimes swimming. As it allows for rapid yet stable movement, many insects adopt a tripedal gait in which they walk with their legs touching the ground in alternating triangles, Insects are the only invertebrates to have evolved flight. Many insects spend at least part of their lives under water, with adaptations that include gills. Some species, such as water striders, are capable of walking on the surface of water, Insects are mostly solitary, but some, such as certain bees, ants and termites, are social and live in large, well-organized colonies. Some insects, such as earwigs, show maternal care, guarding their eggs, Insects can communicate with each other in a variety of ways. Male moths can sense the pheromones of female moths over great distances, other species communicate with sounds, crickets stridulate, or rub their wings together, to attract a mate and repel other males. Lampyridae in the beetle order communicate with light, humans regard certain insects as pests, and attempt to control them using insecticides and a host of other techniques. Some insects damage crops by feeding on sap, leaves or fruits, a few parasitic species are pathogenic. Some insects perform complex ecological roles, blow-flies, for example, help consume carrion, Many other insects are considered ecologically beneficial as predators and a few provide direct economic benefit. Silkworms and bees have been used extensively by humans for the production of silk and honey, in some cultures, people eat the larvae or adults of certain insects. Insect first appears documented in English in 1601 in Hollands translation of Pliny, translations of Aristotles term also form the usual word for insect in Welsh, Serbo-Croatian, Russian, etc. The evolutionary relationship of insects to other animal groups remains unclear, in the Pancrustacea theory, insects, together with Entognatha, Remipedia, and Cephalocarida, make up a natural clade labeled Miracrustacea
21.
Pliny the Elder
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In the latter number will be my uncle, by virtue of his own and of your compositions. Pliny is referring to the fact that Tacitus relied on his uncles now missing work on the History of the German Wars. The wind caused by the sixth and largest pyroclastic surge of the eruption would not allow his ship to leave the shore, and Pliny probably died during this event. Plinys dates are pinned to the eruption of Mount Vesuvius in AD79 and a statement of his nephew that he died in his 56th year, Pliny was the son of an equestrian, Gaius Plinius Celer, and his wife, Marcella. Neither the younger nor the elder Pliny mention the names and their ultimate source is a fragmentary inscription found in a field in Verona and recorded by the 16th century Augustinian monk Onofrio Panvinio at Verona. The reading of the inscription depends on the reconstruction, but in all cases the names come through, whether he was an augur and whether she was named Grania Marcella are less certain. Jean Hardouin presents a statement from a source that he claims was ancient, that Pliny was from Verona. Hardouin also cites the conterraneity of Catullus, additional efforts to connect Celer and Marcella with other gentes are highly speculative. Hardouin is the scholar to use his unknown source. He kept statues of his ancestors there, a statue of Pliny on the facade of the Duomo of Como celebrates him as a native son. He had a sister, Plinia, who married into the Caecilii and was the mother of his nephew, Pliny the Younger, whose letters describe his work and study regimen in detail. In one of his letters to Tacitus, Pliny the Younger details how his uncles breakfasts would be light and simple following the customs of our forefathers. This shows that Pliny the Younger wanted it to be conveyed that Pliny the Elder was a good Roman and this statement would have pleased Tacitus. Two inscriptions identifying the hometown of Pliny the Younger as Como take precedence over the Verona theory, one commemorates the youngers career as imperial magistrate and details his considerable charitable and municipal expenses on behalf of the people of Como. Another identifies his father Lucius village as Fecchio near Como and it is likely therefore that Plinia was a local girl and Pliny the Elder, her brother, was from Como. Gaius was a member of the Plinii gens and he did not take his fathers cognomen, Celer, but assumed his own, Secundus. As his adopted son took the same cognomen, Pliny founded a branch, no earlier instances of the Plinii are known. In 59 BC, only about 82 years before Plinys birth, Julius Caesar founded Novum Comum as a colonia to secure the region against the Alpine tribes, whom he had been unable to defeat
22.
Terrestrial plant
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A terrestrial plant is a plant that grows on or in or from land. Other types of plants are aquatic, epiphytic and lithophytic, there are relatively few obligate submersed aquatic plants, but some examples include members of Hydrocharitaceae and Cabombaceae, Ceratophyllum, and Aldrovanda and most macroalgae. Most aquatic plants can, or prefer to, grow in the emersed form, many terrestrial plants can tolerate extended periods of inundation, and this is often part of the natural habitat of the plant where flooding is common. These plants tolerate extended periods of waterlogging around the roots and even complete submersion under flood waters, growth rates of helophytes decrease significantly during these periods of complete submersion and if water levels do not recede the plant will ultimately decline and perish. An epiphyte is a plant that grows non-parasitically upon another plant
23.
Africa
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Africa is the worlds second-largest and second-most-populous continent. At about 30.3 million km² including adjacent islands, it covers 6% of Earths total surface area and 20.4 % of its land area. With 1.2 billion people as of 2016, it accounts for about 16% of the human population. The continent includes Madagascar and various archipelagos and it contains 54 fully recognized sovereign states, nine territories and two de facto independent states with limited or no recognition. Africas population is the youngest amongst all the continents, the age in 2012 was 19.7. Algeria is Africas largest country by area, and Nigeria by population, afarensis, Homo erectus, H. habilis and H. ergaster – with the earliest Homo sapiens found in Ethiopia being dated to circa 200,000 years ago. Africa straddles the equator and encompasses numerous climate areas, it is the continent to stretch from the northern temperate to southern temperate zones. Africa hosts a diversity of ethnicities, cultures and languages. In the late 19th century European countries colonized most of Africa, Africa also varies greatly with regard to environments, economics, historical ties and government systems. However, most present states in Africa originate from a process of decolonization in the 20th century, afri was a Latin name used to refer to the inhabitants of Africa, which in its widest sense referred to all lands south of the Mediterranean. This name seems to have referred to a native Libyan tribe. The name is connected with Hebrew or Phoenician ʿafar dust. The same word may be found in the name of the Banu Ifran from Algeria and Tripolitania, under Roman rule, Carthage became the capital of the province of Africa Proconsularis, which also included the coastal part of modern Libya. The Latin suffix -ica can sometimes be used to denote a land, the later Muslim kingdom of Ifriqiya, modern-day Tunisia, also preserved a form of the name. According to the Romans, Africa lay to the west of Egypt, while Asia was used to refer to Anatolia, as Europeans came to understand the real extent of the continent, the idea of Africa expanded with their knowledge. 25,4, whose descendants, he claimed, had invaded Libya, isidore of Seville in Etymologiae XIV.5.2. Suggests Africa comes from the Latin aprica, meaning sunny, massey, in 1881, stated that Africa is derived from the Egyptian af-rui-ka, meaning to turn toward the opening of the Ka. The Ka is the double of every person and the opening of the Ka refers to a womb or birthplace
24.
Anatolia
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Anatolia, in geography known as Asia Minor, Asian Turkey, Anatolian peninsula, or Anatolian plateau, is the westernmost protrusion of Asia, which makes up the majority of modern-day Turkey. The region is bounded by the Black Sea to the north, the Mediterranean Sea to the south, the Sea of Marmara forms a connection between the Black and Aegean Seas through the Bosphorus and Dardanelles straits and separates Anatolia from Thrace on the European mainland. Traditionally, Anatolia is considered to extend in the east to a line between the Gulf of Alexandretta and the Black Sea to the Armenian Highlands, thus, traditionally Anatolia is the territory that comprises approximately the western two-thirds of the Asian part of Turkey. The Turkification of Anatolia began under the Seljuk Empire in the late 11th century, however, various non-Turkic languages continue to be spoken by minorities in Anatolia today, including Kurdish, Assyrian, Armenian, Arabic, Laz, Georgian, and Greek. Traditionally, Anatolia is considered to extend in the east to a line running from the Gulf of Alexandretta to the Black Sea. This traditional geographical definition is used, for example, in the latest edition of Merriam-Websters Geographical Dictionary, under this definition, Anatolia is bounded to the east by the Armenian Highlands, and the Euphrates before that river bends to the southeast to enter Mesopotamia. To the southeast, it is bounded by the ranges that separate it from the Orontes valley in Syria, the first name the Greeks used for the Anatolian peninsula was Ἀσία, presumably after the name of the Assuwa league in western Anatolia. As the name of Asia came to be extended to areas east of the Mediterranean. The name Anatolia derives from the Greek ἀνατολή meaning “the East” or more literally “sunrise”, the precise reference of this term has varied over time, perhaps originally referring to the Aeolian, Ionian and Dorian colonies on the west coast of Asia Minor. In the Byzantine Empire, the Anatolic Theme was a theme covering the western, the modern Turkish form of Anatolia is Anadolu, which again derives from the Greek name Aνατολή. The Russian male name Anatoly and the French Anatole share the same linguistic origin, in English the name of Turkey for ancient Anatolia first appeared c. It is derived from the Medieval Latin Turchia, which was used by the Europeans to define the Seljuk controlled parts of Anatolia after the Battle of Manzikert. Human habitation in Anatolia dates back to the Paleolithic, neolithic Anatolia has been proposed as the homeland of the Indo-European language family, although linguists tend to favour a later origin in the steppes north of the Black Sea. However, it is clear that the Anatolian languages, the oldest branch of Indo-European, have spoken in Anatolia since at least the 19th century BC. The earliest historical records of Anatolia stem from the southeast of the region and are from the Mesopotamian-based Akkadian Empire during the reign of Sargon of Akkad in the 24th century BC, scholars generally believe the earliest indigenous populations of Anatolia were the Hattians and Hurrians. The region was famous for exporting raw materials, and areas of Hattian-, one of the numerous cuneiform records dated circa 20th century BC, found in Anatolia at the Assyrian colony of Kanesh, uses an advanced system of trading computations and credit lines. They were speakers of an Indo-European language, the Hittite language, originating from Nesa, they conquered Hattusa in the 18th century BC, imposing themselves over Hattian- and Hurrian-speaking populations. According to the most widely accepted Kurgan theory on the Proto-Indo-European homeland, however, the Hittites adopted the cuneiform script, invented in Mesopotamia
25.
Caucasus Mountains
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The Caucasus Mountains are a mountain system in Eurasia between the Black Sea and the Caspian Sea in the Caucasus region. The Caucasus Mountains include the Greater Caucasus in the north and Lesser Caucasus in the south, the Greater Caucasus runs west-northwest to east-southeast, from the Caucasian Natural Reserve in the vicinity of Sochi on the northeastern shore of the Black Sea nearly to Baku on the Caspian Sea. The Lesser Caucasus runs parallel to the Greater about 100 km south, the Greater and Lesser Caucasus ranges are connected by the Likhi Range, and to the west and east of the Likhi Range lie the Colchis Plain and the Kur-Araz Lowland. The Meskheti Range is a part of the Lesser Caucasus system, in the southeast the Aras River separates the Lesser Caucasus from the Talysh Mountains which straddle the border of southeastern Azerbaijan and Iran. The highest peak in the Caucasus range is Mount Elbrus in the Greater Caucasus, Mountains near Sochi hosted part of the 2014 Winter Olympics. Geologically, the Caucasus Mountains belong to a system that extends from southeastern Europe into Asia, the Greater Caucasus Mountains are mainly composed of Cretaceous and Jurassic rocks with the Paleozoic and Precambrian rocks in the higher regions. Some volcanic formations are found throughout the range, on the other hand, the Lesser Caucasus Mountains are formed predominantly of the Paleogene rocks with a much smaller portion of the Jurassic and Cretaceous rocks. The Caucasus Mountains formed largely as the result of a plate collision between the Arabian plate moving northwards with respect to the Eurasian plate. As this happened, the rocks that had been deposited in this basin from the Jurassic to the Miocene were folded to form the Greater Caucasus Mountains. This collision also caused the uplift and the Cenozoic volcanic activity in the Lesser Caucasus Mountains, the entire region is regularly subjected to strong earthquakes from this activity. While the Greater Caucasus Mountains have a mainly folded sedimentary structure, the Javakheti Volcanic Plateau in Georgia and the surrounding volcanic ranges which extend well into central Armenia are some of the youngest features of the region. The Kazbek is no active, but the Elbrus erupted in postglacial times. Contemporary seismic activity is a prominent feature of the region, reflecting active faulting, clusters of seismicity occur in Dagestan and in northern Armenia. Many devastating earthquakes have been documented in historical times, including the Spitak earthquake in December 1988 which destroyed the Gyumri-Vanadzor region of Armenia, europes highest mountain is Mount Elbrus 5,642 m in the Caucasus Mountains. Elbrus is 832 m higher than Mont Blanc, the highest peak in the Alps at 4,810 m, the Caucasus Mountains are defined as the continental divide between Asia and Europe for the region between the Black and Caspian Seas. The table below lists some of the highest peaks of the Caucasus, with the exception of Shkhara, the heights are taken from Soviet 1,50,000 mapping. There are higher and more prominent, but nameless, peaks than some of the peaks included below, the climate of the Caucasus varies both vertically and horizontally. Temperature generally decreases as elevation rises, average annual temperature in Sukhumi, Abkhazia at sea level is 15 °C while on the slopes of Mt. Kazbek at an elevation of 3,700 metres, average annual temperature falls to−6.1 °C
26.
Mediterranean region
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As a rule of thumb, the Mediterranean Basin is the Old World region where olive trees grow. The Mediterranean basin covers portions of three continents Africa, Asia, and Europe and it has a varied and contrasting topography. Contrary to the sandy beach images portrayed in most tourist brochures. Mountains can be seen from almost anywhere, the Mediterranean Basin extends into Western Asia, covering the western and southern portions of the peninsula of Turkey, excluding the temperate-climate mountains of central Turkey. It includes the Mediterranean climate Levant at the end of the Mediterranean, bounded on the east and south by the Syrian. The northern portion of the Maghreb region of northwestern Africa has a Mediterranean climate, separated from the Sahara Desert, Europe lies to the north, and three large Southern European peninsulas, the Iberian Peninsula, Italian Peninsula, and the Balkan Peninsula, extend into the Mediterranean-climate zone. The Mediterranean Basin was shaped by the ancient collision of the northward-moving African-Arabian continent with the stable Eurasian continent, as Africa-Arabia moved north, it closed the former Tethys Sea, which formerly separated Eurasia from the ancient super continent of Gondwana, of which Africa was part. At about the time,170 mya in the Jurassic period. The collision pushed up a vast system of mountains, extending from the Pyrenees in Spain to the Zagros Mountains in Iran and this episode of mountain building, known as the Alpine orogeny, occurred mostly during the Oligocene and Miocene epochs. The Neotethys became larger during these collisions and associated folding and subduction, about 6 mya during the late Miocene, the Mediterranean was closed at its western end by drifting Africa, which caused the entire sea to evaporate. There followed several episodes of sea drawdown and re-flooding known as the Messinian Salinity Crisis, recent studies, however, show that repeated desiccation and re-flooding is unlikely from a geodynamic point of view. The end of the Miocene also marked a change in the Mediterranean Basins climate, fossil evidence shows that the Mediterranean Basin had a relatively humid subtropical climate with summer rainfall during the Miocene, which supported laurel forests. The shift to a Mediterranean climate occurred within the last 3. 2–2.8 million years, during the Pliocene epoch, much of these forests and shrublands have been altered beyond recognition by thousands of years of human habitation. There are now very few relatively intact natural areas in what was once a wooded region. The Mediterranean Region was first proposed by German botanist August Grisebach in the late 19th century, drosophyllaceae, recently segregated from Droseraceae, is the only plant family endemic to the region. Among the endemic plant genera are, The genera Aubrieta, Sesamoides, Cynara, Dracunculus, moreover, many plant taxa are shared with one of the four neighboring floristic regions only. The Mediterranean Basin is the largest of the worlds five Mediterranean forests, woodlands and it is home to a number of plant communities, which vary with rainfall, elevation, latitude, and soils. Scrublands occur in the driest areas, especially areas near the seacoast where wind, low, soft-leaved scrublands around the Mediterranean are known as garrigar in Catalan, garrigue in French, phrygana in Greek, tomillares in Spanish, and batha in Hebrew
27.
Tuber
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Tubers are enlarged structures in some plant species used as storage organs for nutrients. They are used for the plants perennation, to energy and nutrients for regrowth during the next growing season. Stem tubers form from thickened rhizomes or stolons, common plant species with stem tubers include potato and yam. Some sources also treat modified lateral roots under the definition, these are encountered in sweet potato, cassava, the term originates from Latin tuber, meaning lump, bump, swelling. Some sources define the term tuber to mean only structures derived from stems, a stem tuber forms from thickened rhizomes or stolons. The top sides of the tuber produce shoots that grow into stems and leaves. They tend to form at the sides of the parent plant and are most often located near the soil surface, the underground stem tuber is normally a short-lived storage and regenerative organ developing from a shoot that branches off a mature plant. The offsprings or new tubers are attached to a parent tuber or form at the end of a hypogeogenous rhizome, some plants also form smaller tubers and/or tubercules which act like seeds, producing small plants that resemble seedlings. Stem tubers generally start off as enlargements of the section of a seedling. Tuberous begonia, yams, and Cyclamen are commonly grown stem tubers, mignonette vine produces aerial stem tubers on 12-to-25-foot-tall vines, the tubers fall to the ground and grow. Enlarged stolons thicken to develop into storage organs, the tuber has all the parts of a normal stem, including nodes and internodes. The nodes are the eyes and each has a leaf scar, the nodes or eyes are arranged around the tuber in a spiral fashion beginning on the end opposite the attachment point to the stolon. The terminal bud is produced at the farthest point away from the stolon attachment, internally, a tuber is filled with starch stored in enlarged parenchyma like cells. The inside of a tuber has the cell structures of any stem, including a pith, vascular zones. The tuber is produced in one growing season and used to perennate the plant, as the main shoot develops from the tuber, the base of the shoot close to the tuber produces adventitious roots and lateral buds on the shoot. The shoot also produces stolons that are long etiolated stems, the stolon elongates during long days with the presence of high auxins levels that prevent root growth off of the stolon. Before new tuber formation begins, the stolon must be a certain age, the enzyme lipoxygenase makes a hormone, jasmonic acid, which is involved in the control of potato tuber development. The stolons are easily recognized when potato plants are grown from seeds, as the plants grow, stolons are produced around the soil surface from the nodes
28.
Symbiosis
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Symbiosis is any type of a close and long-term biological interaction between two different species, be it mutualistic, commensalistic, or parasitic. In 1879, Heinrich Anton de Bary defined it as the living together of unlike organisms, Symbiosis can be obligatory, which means that one or both of the symbionts entirely depend on each other for survival, or facultative when they can generally live independently. When one organism lives on another such as mistletoe, it is called ectosymbiosis, or endosymbiosis when one partner lives inside the tissues of another, as in Symbiodinium in corals. In 1877, Albert Bernhard Frank used the term symbiosis which previously had used to depict people living together in community to describe the mutualistic relationship in lichens. In 1879, the German mycologist Heinrich Anton de Bary defined it as the living together of unlike organisms, symbiotic relationships can be obligate, meaning that one or both of the symbionts entirely depend on each other for survival. For example, in lichens, which consist of fungal and photosynthetic symbionts, the algal or cyanobacterial symbionts in lichens, such as Trentepohlia, can generally live independently, and their symbiosis is, therefore, facultative. Endosymbiosis is any relationship in which one symbiont lives within the tissues of the other, either within the cells or extracellularly. Mutualism or interspecies reciprocal altruism is a relationship between individuals of different species where both individuals benefit, in general, only lifelong interactions involving close physical and biochemical contact can properly be considered symbiotic. Mutualistic relationships may be either obligate for both species, obligate for one but facultative for the other, or facultative for both, a large percentage of herbivores have mutualistic gut flora to help them digest plant matter, which is more difficult to digest than animal prey. This gut flora is made up of cellulose-digesting protozoans or bacteria living in the herbivores intestines, coral reefs are the result of mutualisms between coral organisms and various types of algae which live inside them. Most land plants and land ecosystems rely on mutualisms between the plants, which fix carbon from the air, and mycorrhyzal fungi, which help in extracting water, an example of mutual symbiosis is the relationship between the ocellaris clownfish that dwell among the tentacles of Ritteri sea anemones. The territorial fish protects the anemone from anemone-eating fish, and in turn the stinging tentacles of the anemone protect the clownfish from its predators, a special mucus on the clownfish protects it from the stinging tentacles. A further example is the fish, which sometimes lives together with a shrimp. The shrimp digs and cleans up a burrow in the sand in which both the shrimp and the fish live. The shrimp is almost blind, leaving it vulnerable to predators when outside its burrow, in case of danger the goby fish touches the shrimp with its tail to warn it. When that happens both the shrimp and goby fish quickly retreat into the burrow, different species of gobies also exhibit mutualistic behavior through cleaning up ectoparasites in other fish. Another non-obligate symbiosis is known from encrusting bryozoans and hermit crabs, the bryozoan colony develops a cirumrotatory growth and offers the crab a helicospiral-tubular extension of its living chamber that initially was situated within a gastropod shell. A spectacular examples of mutualism is between the siboglinid tube worms and symbiotic bacteria that live at hydrothermal vents and cold seeps
29.
Fungus
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A fungus is any member of the group of eukaryotic organisms that includes microorganisms such as yeasts and molds, as well as the more familiar mushrooms. These organisms are classified as a kingdom, Fungi, which is separate from the other eukaryotic life kingdoms of plants, a characteristic that places fungi in a different kingdom from plants, bacteria and some protists, is chitin in their cell walls. Similar to animals, fungi are heterotrophs, they acquire their food by absorbing dissolved molecules, growth is their means of mobility, except for spores, which may travel through the air or water. Fungi are the principal decomposers in ecological systems and this fungal group is distinct from the structurally similar myxomycetes and oomycetes. The discipline of biology devoted to the study of fungi is known as mycology, in the past, mycology was regarded as a branch of botany, although it is now known fungi are genetically more closely related to animals than to plants. Abundant worldwide, most fungi are inconspicuous because of the size of their structures. Fungi include symbionts of plants, animals, or other fungi and they may become noticeable when fruiting, either as mushrooms or as molds. Fungi perform a role in the decomposition of organic matter and have fundamental roles in nutrient cycling. Since the 1940s, fungi have been used for the production of antibiotics, Fungi are also used as biological pesticides to control weeds, plant diseases and insect pests. Many species produce bioactive compounds called mycotoxins, such as alkaloids and polyketides, the fruiting structures of a few species contain psychotropic compounds and are consumed recreationally or in traditional spiritual ceremonies. Fungi can break down manufactured materials and buildings, and become significant pathogens of humans, losses of crops due to fungal diseases or food spoilage can have a large impact on human food supplies and local economies. The fungus kingdom encompasses a diversity of taxa with varied ecologies, life cycle strategies. However, little is known of the biodiversity of Kingdom Fungi. Advances in molecular genetics have opened the way for DNA analysis to be incorporated into taxonomy, phylogenetic studies published in the last decade have helped reshape the classification within Kingdom Fungi, which is divided into one subkingdom, seven phyla, and ten subphyla. The English word fungus is directly adopted from the Latin fungus, used in the writings of Horace, a group of all the fungi present in a particular area or geographic region is known as mycobiota, e. g. the mycobiota of Ireland. Like plants, fungi grow in soil and, in the case of mushrooms, form conspicuous fruit bodies. The fungi are now considered a kingdom, distinct from both plants and animals, from which they appear to have diverged around one billion years ago. Fungi have membrane-bound cytoplasmic organelles such as mitochondria, sterol-containing membranes and they have a characteristic range of soluble carbohydrates and storage compounds, including sugar alcohols, disaccharides, and polysaccharides
30.
Leaf
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A leaf is an organ of a vascular plant and is the principal lateral appendage of the stem. The leaves and stem together form the shoot, Leaves are collectively referred to as foliage, as in autumn foliage. Although leaves can be seen in different shapes, sizes and textures, typically a leaf is a thin, dorsiventrally flattened organ, borne above ground. Most leaves have distinctive upper surface and lower surface that differ in colour, hairiness, broad, flat leaves with complex venation are known as megaphylls and the species that bear them, the majority, as broad-leaved or megaphyllous plants. In others, such as the clubmosses, with different evolutionary origins, some leaves, such as bulb scales are not above ground, and in many aquatic species the leaves are submerged in water. Succulent plants often have thick juicy leaves, but some leaves are without major photosynthetic function and may be dead at maturity, as in some cataphylls, furthermore, several kinds of leaf-like structures found in vascular plants are not totally homologous with them. Examples include flattened plant stems called phylloclades and cladodes, and flattened leaf stems called phyllodes which differ from both in their structure and origin. Many structures of plants, such as the phyllids of mosses and liverworts and even of some foliose lichens. Leaves are the most important organs of most vascular plants and these are then further processed by chemical synthesis into more complex organic molecules such as cellulose, the basic structural material in plant cell walls. The plant must therefore bring these three together in the leaf for photosynthesis to take place. Once sugar has been synthesized, it needs to be transported to areas of growth such as the plant shoots and roots. Vascular plants transport sucrose in a tissue called the phloem. The phloem and xylem are parallel to each other but the transport of materials is usually in opposite directions. Within the leaf these vascular systems branch to form veins which supply as much as the leaf as possible and they are arranged on the plant so as to expose their surfaces to light as efficiently as possible without shading each other, but there are many exceptions and complications. For instance plants adapted to windy conditions may have pendent leaves, such as in many willows, the flat, or laminar, shape also maximises thermal contact with the surrounding air, promoting cooling. Functionally, in addition to photosynthesis the leaf is the site of transpiration and guttation. Many gymnosperms have thin needle-like or scale-like leaves that can be advantageous in cold climates with frequent snow and these are interpreted as reduced from megaphyllous leaves of their Devonian ancestors. For xerophytes the major constraint is not light flux or intensity, some window plants such as Fenestraria species and some Haworthia species such as Haworthia tesselata and Haworthia truncata are examples of xerophytes. and Bulbine mesembryanthemoides
31.
Plant stem
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A stem is one of two main structural axes of a vascular plant, the other being the root. The stem is divided into nodes and internodes, The nodes hold one or more leaves. Adventitious roots may also be produced from the nodes, the internodes distance one node from another. The term shoots is often confused with stems, shoots generally refers to new plant growth including both stems and other structures like leaves or flowers. In most plants stems are located above the surface but some plants have underground stems. Stems have four main functions which are, Support for and the elevation of leaves, flowers, the stems keep the leaves in the light and provide a place for the plant to keep its flowers and fruits. Transport of fluids between the roots and the shoots in the xylem and phloem Storage of nutrients Production of new living tissue, the normal lifespan of plant cells is one to three years. Stems have cells called meristems that annually generate new living tissue, Stems are often specialized for storage, asexual reproduction, protection or photosynthesis, including the following, Acaulescent – used to describe stems in plants that appear to be stemless. Actually these stems are just extremely short, the leaves appearing to rise out of the ground. Arborescent – tree like with woody stems normally with a single trunk, branched – aerial stems are described as being branched or unbranched Bud – an embryonic shoot with immature stem tip. Bulb – a short underground stem with fleshy storage leaves attached, e. g. onion, daffodil. Bulbs often function in reproduction by splitting to form new bulbs or producing small new bulbs termed bulblets, bulbs are a combination of stem and leaves so may better be considered as leaves because the leaves make up the greater part. Caespitose – when stems grow in a mass or clump or in low growing mats. Cladode – a flattened stem that appears more-or-less leaf like and is specialized for photosynthesis, climbing – stems that cling or wrap around other plants or structures. Corm – a short enlarged underground, storage stem, e. g. taro, decumbent – stems that lie flat on the ground and turn upwards at the ends. Fruticose – stems that grow shrublike with woody like habit, herbaceous – non woody, they die at the end of the growing season. Pedicel – stems that serve as the stalk of a flower in an inflorescence or infrutescence. Peduncle – a stem that supports an inflorescence Prickle – a sharpened extension of the outer layers
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Pollinator
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A pollinator is an animal that moves pollen from the male anther of a flower to the female stigma of a flower. This helps to bring about fertilization of the ovules in the flower by the gametes from the pollen grains. Insect pollinators include bees, pollen wasps, ants, flies including bee flies and hoverflies, lepidopterans, vertebrates, mainly bats and birds, but also some non-bat mammals and some lizards pollinate certain plants. Among the pollinating birds are hummingbirds, honeyeaters and sunbirds with long beaks, cycads, which are not flowering plants, are also pollinated by insects. A pollinator is different from a pollenizer, a plant that is a source of pollen for the pollination process, plants fall into pollination syndromes that reflect the type of pollinator being attracted. These are characteristics such as, overall size, the depth and width of the corolla, the color. When these characteristics are experimentally modified, pollinator visitation may decline and it has recently been discovered that cycads, which are not flowering plants, are also pollinated by insects. The most recognized pollinators are the species of bees, which are plainly adapted to pollination. Bees typically are fuzzy and carry an electrostatic charge, Honey bees, bumblebees, and their relatives do not have a scopa, but the hind leg is modified into a structure called the corbicula. Most bees gather nectar, an energy source, and pollen, which is high protein food, to nurture their young. Euglossine bees pollinate orchids, but these are male bees collecting floral scents rather than females gathering nectar or pollen, female orchid bees act as pollinators, but of flowers other than orchids. Eusocial bees such as honey bees need an abundant and steady pollen source to multiply, Honey bees travel from flower to flower, collecting nectar, and pollen grains. The bee collects the pollen by rubbing against the anthers, the pollen collects on the hind legs, in a structure referred to as a pollen basket. As the bee flies from flower to flower, some of the grains are transferred onto the stigma of other flowers. Nectar provides the energy for bee nutrition, pollen provides the protein, when bees are rearing large quantities of brood, bees deliberately gather pollen to meet the nutritional needs of the brood. Good pollination management seeks to have bees in a state during the bloom period of the crop, thus requiring them to gather pollen. Thus, the management techniques of a beekeeper providing pollination services are different from, and to some extent in tension with, many insects other than bees accomplish pollination by visiting flowers for nectar or pollen, or commonly both. Many do so adventitiously, but the most important pollinators are specialists for at least parts of their lifecycles for at least certain functions
33.
Alkene
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In organic chemistry, an alkene is an unsaturated hydrocarbon that contains at least one carbon–carbon double bond. The words alkene, olefin, and olefine are used often interchangeably, acyclic alkenes, with only one double bond and no other functional groups, known as mono-enes, form a homologous series of hydrocarbons with the general formula CnH2n. Alkenes have two hydrogen atoms less than the corresponding alkane, the simplest alkene, ethylene, with the International Union of Pure and Applied Chemistry name ethene, is the organic compound produced on the largest scale industrially. Aromatic compounds are drawn as cyclic alkenes, but their structure and properties are different. This double bond is stronger than a single covalent bond and also shorter, each carbon of the double bond uses its three sp2 hybrid orbitals to form sigma bonds to three atoms. The unhybridized 2p atomic orbitals, which lie perpendicular to the created by the axes of the three sp² hybrid orbitals, combine to form the pi bond. This bond lies outside the main C–C axis, with half of the bond on one side of the molecule, rotation about the carbon–carbon double bond is restricted because it incurs an energetic cost to break the alignment of the p orbitals on the two carbon atoms. As a consequence, substituted alkenes may exist as one of two isomers, called cis or trans isomers, more complex alkenes may be named with the E–Z notation for molecules with three or four different substituents. For example, of the isomers of butene, the two groups of -but-2-ene appear on the same side of the double bond, and in -but-2-ene the methyl groups appear on opposite sides. These two isomers of butene are slightly different in their chemical and physical properties, a 90° twist of the C=C bond requires less energy than the strength of a pi bond, and the bond still holds. This contradicts a common assertion that the p orbitals would be unable sustain such a bond. In truth, the misalignment of the p orbitals is less than expected because pyramidalization takes place, trans-Cyclooctene is a stable strained alkene and the orbital misalignment is only 19° with a dihedral angle of 137° and a degree of pyramidalization of 18°. The trans isomer of cycloheptene is stable only at low temperatures, as predicted by the VSEPR model of electron pair repulsion, the molecular geometry of alkenes includes bond angles about each carbon in a double bond of about 120°. The angle may vary because of steric strain introduced by nonbonded interactions between groups attached to the carbons of the double bond. For example, the C–C–C bond angle in propylene is 123. 9°, for bridged alkenes, Bredts rule states that a double bond cannot occur at the bridgehead of a bridged ring system unless the rings are large enough. The physical properties of alkenes and alkanes are similar and they are colourless, nonpolar, combustable, and almost odorless. The physical state depends on mass, like the corresponding saturated hydrocarbons, the simplest alkenes, ethene, propene. Linear alkenes of approximately five to sixteen carbons are liquids, Alkenes are relatively stable compounds, but are more reactive than alkanes, either because of the reactivity of the carbon–carbon pi-bond or the presence of allylic CH centers
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Alkane
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In organic chemistry, an alkane, or paraffin, is an acyclic saturated hydrocarbon. In other words, an alkane consists of hydrogen and carbon atoms arranged in a structure in which all the carbon-carbon bonds are single. Alkanes have the chemical formula CnH2n+2. The alkanes range in complexity from the simplest case of methane, CH4 where n =1, in an alkane, each carbon atom has 4 bonds, and each hydrogen atom is joined to one of the carbon atoms. The longest series of linked carbon atoms in a molecule is known as its skeleton or carbon backbone. The number of atoms may be thought of as the size of the alkane. One group of the alkanes are waxes, solids at standard ambient temperature and pressure. They can be viewed as molecular trees upon which can be hung the more functional groups of biological molecules. The alkanes have two main sources, petroleum and natural gas. Saturated hydrocarbons are hydrocarbons having only single covalent bonds between their carbons, according to the definition by IUPAC, the former two are alkanes, whereas the third group is called cycloalkanes. Saturated hydrocarbons can also combine any of the linear, cyclic and branching structures, the formula is CnH 2n−2k+2. Alkanes are the ones, corresponding to k =0. Alkanes with more than three carbon atoms can be arranged in different ways, forming structural isomers. The simplest isomer of an alkane is the one in which the atoms are arranged in a single chain with no branches. This isomer is called the n-isomer. However the chain of atoms may also be branched at one or more points. The number of possible isomers increases rapidly with the number of carbon atoms, for example, 3-methylhexane and its higher homologues are chiral due to their stereogenic center at carbon atom number 3. In addition to the alkane isomers, the chain of atoms may form one or more loops
35.
Pheromone
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A pheromone is a secreted or excreted chemical factor that triggers a social response in members of the same species. Pheromones are chemicals capable of acting outside the body of the individual to impact the behavior of the receiving individuals. There are alarm pheromones, food trail pheromones, sex pheromones, Pheromones are used from basic unicellular prokaryotes to complex multicellular eukaryotes. Their use among insects has been well documented. In addition, some vertebrates, plants and ciliates communicate by using pheromones, the portmanteau word pheromone was coined by Peter Karlson and Martin Lüscher in 1959, based on the Greek φερω pheroo and ὁρμων hormon. Pheromones are also classified as ecto-hormones. For this reason, bacteria are too small to use pheromones as sex attractants on an individual basis, however, they do use them to determine the local population density of similar organisms and control behaviors that take more time to execute. Aggregation pheromones function in mate selection, overcoming host resistance by mass attack, a group of individuals at one location is referred to as an aggregation, whether consisting of one sex or both sexes. Most sex pheromones are produced by the females, only a percentage of sex attractants are produced by males. Aggregation pheromones have been found in members of the Coleoptera, Diptera, Hemiptera, Dictyoptera, aggregation pheromones are among the most ecologically selective pest suppression methods. They are nontoxic and effective at low concentrations. Some species release a volatile substance when attacked by a predator that can trigger flight or aggression in members of the same species, for example, Vespula squamosa use alarm pheromones to alert others to a threat. In Polistes exclamans, alarm pheromones are used as an alert to incoming predators. Pheromones also exist in plants, Certain plants emit alarm pheromones when grazed upon and these tannins make the plants less appetizing for the herbivore. Epideictic pheromones are different from territory pheromones, when it comes to insects, releaser pheromones are pheromones that cause an alteration in the behavior of the recipient. For example, some organisms use powerful attractant molecules to attract mates from a distance of two miles or more, in general, this type of pheromone elicits a rapid response, but is quickly degraded. In contrast, a primer pheromone has an onset and a longer duration. For example, rabbit release mammary pheromones that trigger immediate nursing behavior by their babies, signal pheromones cause short-term changes, such as the neurotransmitter release that activates a response
36.
Hybrid (biology)
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In biology, a hybrid, also known as a cross breed, is the result of combining, through sexual reproduction, the qualities of two animals or plants of different breeds, varieties, species or genera. Hybrids are not always intermediate between their parents, but can show heterosis or hybrid vigour, often growing larger or taller than either parent, some act before fertilization, others after it. A few animal species and many plant species, however, are the result of hybrid speciation, doubling the number of chromosomes to create polyploids is important in hybrid speciation, because homoploid hybrids are rarely fertile, the polyploid hybrids are allopolyploids. Human impact on the environment has greatly increased the mixing of species, with introduced species worldwide, the genetic mixing may threaten many species with extinction, while genetic erosion in crop plants may be damaging the gene pools of many species for future breeding. Many commercially useful fruits, flowers, garden herbs and trees have been produced by hybridization, one flower, Oenothera lamarckiana, was central to early genetics research into polyploidy. The term hybrid is derived from Latin hybrida, used for such as of a tame sow and a wild boar, or the child of a freeman. The term came into use in English in the 19th century. Conspicuous hybrids are named with portmanteau words, starting in the 1920s with the breeding of tiger-lion hybrids. The cross between two different homozygous lines produces an F1 hybrid that is heterozygous, having two alleles, one contributed by each parent and typically one is dominant and the other recessive. Typically, the F1 generation is also phenotypically homogeneous, producing offspring that are all similar to each other, double cross hybrids result from the cross between two different F1 hybrids. Three-way cross hybrids result from the cross between an F1 hybrid and an inbred line, triple cross hybrids result from the crossing of two different three-way cross hybrids. Top cross hybrids result from the crossing of a top quality or pure-bred male, population hybrids result from the crossing of plants or animals in a population with those of another population. These include interspecific hybrids or crosses between different breeds. e, from the point of view of genetics, different kinds of hybrid can be distinguished as follows, A genetic hybrid carries two different alleles of the same gene. A structural hybrid results from the fusion of gametes that have differing structure in at least one chromosome, a numerical hybrid results from the fusion of gametes having different haploid numbers of chromosomes. A permanent hybrid results when only the heterozygous genotype occurs, as in Oenothera lamarckiana, from the point of view of taxonomy, hybrids differ according to their parentage, Hybrids between different subspecies are called intra-specific hybrids. Offspring resulting from interspecies mating, are called interspecific hybrids, these result in hybrid speciation. Intergeneric hybrids result from matings between different genera, such as sheep and goats. Interfamilial hybrids such as chickens and guineafowl or pheasants are reliably described but extremely rare
37.
Royal Botanic Gardens, Kew
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Royal Botanic Gardens, Kew is a non-departmental public body in the United Kingdom sponsored by the Department for Environment, Food and Rural Affairs. An internationally important botanical research and education institution, it employs 750 staff and its chief executive is the current Director, Richard Deverell. Its board of trustees is chaired by Marcus Agius, a chairman of Barclays PLC. The Seed Bank is also the site of research projects. Seed stored at the bank fulfils two functions, it provides an ex situ conservation resource and also facilitates research around the globe by acting as a repository for seed scientists, Kew jointly with the Forestry Commission, founded Bedgebury Pinetum in Kent, specialising in growing conifers. The organisation has an average of 1 million paying visitors per year, the 326 acre site has 40 historically important buildings and collections of over 40,000 species of plants. Kew Gardens became a United Nations World Heritage Site on 3 July 2003, Kew is governed by a Board of Trustees which comprises a chairman and 11 members. Ten members and the chairman are appointed by the Secretary of State for Environment, Food and her Majesty the Queen appoints her own trustee on the recommendation of the Secretary of State. The Director of Science is University of Oxford Professor Kathy Willis and her deputy is Professor Monique Simmonds. Professor Mark Chase is Senior Research Professor, the IPNI includes information from the Index Kewensis, a project which began in the 19th century to provide an Index to the Names and Authorities of all known flowering plants and their countries. Kew also cooperates with the Missouri Botanical Garden in a project called The Plant List. The Plant List is an Internet encyclopedia project which was launched in 2010 to compile a comprehensive list of botanical nomenclature, the Plant List has 1,040,426 scientific plant names of species rank of which 298,900 are accepted species names. In addition, the list has 620 plant families and 16,167 plant genera
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Ophrys apifera
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Ophrys apifera, known in Europe as the bee orchid, is a perennial herbaceous plant belonging to the family Orchidaceae. The name Ophrys derives from the Greek word ophrys, meaning eyebrow, while the Latin specific epithet apifera means bee-bearing or bee-bringing, Ophrys apifera grows to a height of 15–50 centimetres. This hardy orchid develops small rosettes of leaves in autumn and they continue to grow slowly during winter. Basal leaves are ovate or oblong-lanceolate, upper leaves and bracts are ovate-lanceolate, the plant blooms from mid-April to July producing a spike composed from one to twelve flowers. The flowers have sepals, with a central green rib and their colour varies from white to pink, while petals are short, pubescent. The labellum is trilobed, with two pronounced humps on the lateral lobes, the median lobe is hairy and similar to the abdomen of a bee. It is quite variable in the pattern of coloration, but usually brownish-red with yellow markings, the gynostegium is at right angles, with an elongated apex. Floral parts display the presence of quercetin and kaempferol glycosides, which are supposed to be acylated, the pink outer tepals show the presence of anthocyanins. Ophrys apifera is the species of the genus Ophrys which preferentially practices self-pollination. The flowers are almost exclusively self-pollinating in the ranges of the plants distribution. In this case the plant attracts these insects by producing a scent that mimics the scent of the female bee, in addition, the lip acts as a decoy as the male bee confuses it with a female. Pollen transfer occurs during the ensuing pseudocopulation, the flowers emit allomones that attract the bee species Tetralonia cressa and Eucera pulveraceae. Eucera longicornis males have been observed attempting to copulate with the flowers and it is also believed that male bees would preferentially select orchids with the most bee-like lips and attempt to mate with them, transferring pollen in the process. Ophrys apifera is a widespread across central and southern Europe, as well as North Africa and its range stretches from Portugal, Ireland and Denmark east to Iran and the Caucasus. It is quite common in the Mediterranean region eastwards to the Black Sea, in the UK, it has a distinct southeastern preference, being more common in England. It is relatively common in the northeast of England and in recent years large numbers have appeared in the grass surrounding the Metro Centre in Gateshead. In Scotland, it was thought to be extinct, but was rediscovered in Ayrshire in 2003, in some countries the plants have protected status. They are unusual in that in years they appear in great numbers
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Iran
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Iran, also known as Persia, officially the Islamic Republic of Iran, is a sovereign state in Western Asia. Comprising a land area of 1,648,195 km2, it is the second-largest country in the Middle East, with 82.8 million inhabitants, Iran is the worlds 17th-most-populous country. It is the country with both a Caspian Sea and an Indian Ocean coastline. The countrys central location in Eurasia and Western Asia, and its proximity to the Strait of Hormuz, Tehran is the countrys capital and largest city, as well as its leading economic and cultural center. Iran is the site of to one of the worlds oldest civilizations, the area was first unified by the Iranian Medes in 625 BC, who became the dominant cultural and political power in the region. The empire collapsed in 330 BC following the conquests of Alexander the Great, under the Sassanid Dynasty, Iran again became one of the leading powers in the world for the next four centuries. Beginning in 633 AD, Arabs conquered Iran and largely displaced the indigenous faiths of Manichaeism and Zoroastrianism by Islam, Iran became a major contributor to the Islamic Golden Age that followed, producing many influential scientists, scholars, artists, and thinkers. During the 18th century, Iran reached its greatest territorial extent since the Sassanid Empire, through the late 18th and 19th centuries, a series of conflicts with Russia led to significant territorial losses and the erosion of sovereignty. Popular unrest culminated in the Persian Constitutional Revolution of 1906, which established a monarchy and the countrys first legislative body. Following a coup instigated by the U. K. Growing dissent against foreign influence and political repression led to the 1979 Revolution, Irans rich cultural legacy is reflected in part by its 21 UNESCO World Heritage Sites, the third-largest number in Asia and 11th-largest in the world. Iran is a member of the UN, ECO, NAM, OIC. Its political system is based on the 1979 Constitution which combines elements of a democracy with a theocracy governed by Islamic jurists under the concept of a Supreme Leadership. A multicultural country comprising numerous ethnic and linguistic groups, most inhabitants are Shia Muslims, the largest ethnic groups in Iran are the Persians, Azeris, Kurds and Lurs. Historically, Iran has been referred to as Persia by the West, due mainly to the writings of Greek historians who called Iran Persis, meaning land of the Persians. As the most extensive interactions the Ancient Greeks had with any outsider was with the Persians, however, Persis was originally referred to a region settled by Persians in the west shore of Lake Urmia, in the 9th century BC. The settlement was then shifted to the end of the Zagros Mountains. In 1935, Reza Shah requested the international community to refer to the country by its native name, opposition to the name change led to the reversal of the decision, and Professor Ehsan Yarshater, editor of Encyclopædia Iranica, propagated a move to use Persia and Iran interchangeably
40.
France
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France, officially the French Republic, is a country with territory in western Europe and several overseas regions and territories. The European, or metropolitan, area of France extends from the Mediterranean Sea to the English Channel and the North Sea, Overseas France include French Guiana on the South American continent and several island territories in the Atlantic, Pacific and Indian oceans. France spans 643,801 square kilometres and had a population of almost 67 million people as of January 2017. It is a unitary republic with the capital in Paris. Other major urban centres include Marseille, Lyon, Lille, Nice, Toulouse, during the Iron Age, what is now metropolitan France was inhabited by the Gauls, a Celtic people. The area was annexed in 51 BC by Rome, which held Gaul until 486, France emerged as a major European power in the Late Middle Ages, with its victory in the Hundred Years War strengthening state-building and political centralisation. During the Renaissance, French culture flourished and a colonial empire was established. The 16th century was dominated by civil wars between Catholics and Protestants. France became Europes dominant cultural, political, and military power under Louis XIV, in the 19th century Napoleon took power and established the First French Empire, whose subsequent Napoleonic Wars shaped the course of continental Europe. Following the collapse of the Empire, France endured a succession of governments culminating with the establishment of the French Third Republic in 1870. Following liberation in 1944, a Fourth Republic was established and later dissolved in the course of the Algerian War, the Fifth Republic, led by Charles de Gaulle, was formed in 1958 and remains to this day. Algeria and nearly all the colonies became independent in the 1960s with minimal controversy and typically retained close economic. France has long been a centre of art, science. It hosts Europes fourth-largest number of cultural UNESCO World Heritage Sites and receives around 83 million foreign tourists annually, France is a developed country with the worlds sixth-largest economy by nominal GDP and ninth-largest by purchasing power parity. In terms of household wealth, it ranks fourth in the world. France performs well in international rankings of education, health care, life expectancy, France remains a great power in the world, being one of the five permanent members of the United Nations Security Council with the power to veto and an official nuclear-weapon state. It is a member state of the European Union and the Eurozone. It is also a member of the Group of 7, North Atlantic Treaty Organization, Organisation for Economic Co-operation and Development, the World Trade Organization, originally applied to the whole Frankish Empire, the name France comes from the Latin Francia, or country of the Franks
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Great Britain
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Great Britain, also known as Britain, is a large island in the north Atlantic Ocean off the northwest coast of continental Europe. With an area of 209,331 km2, Great Britain is the largest European island, in 2011 the island had a population of about 61 million people, making it the worlds third-most populous island after Java in Indonesia and Honshu in Japan. The island of Ireland is situated to the west of it, the island is dominated by a maritime climate with quite narrow temperature differences between seasons. Politically, the island is part of the United Kingdom of Great Britain and Northern Ireland, most of England, Scotland, and Wales are on the island. The term Great Britain often extends to surrounding islands that form part of England, Scotland, and Wales. A single Kingdom of Great Britain resulted from the union of the Kingdom of England, the archipelago has been referred to by a single name for over 2000 years, the term British Isles derives from terms used by classical geographers to describe this island group. By 50 BC Greek geographers were using equivalents of Prettanikē as a name for the British Isles. However, with the Roman conquest of Britain the Latin term Britannia was used for the island of Great Britain, the oldest mention of terms related to Great Britain was by Aristotle, or possibly by Pseudo-Aristotle, in his text On the Universe, Vol. III. To quote his works, There are two large islands in it, called the British Isles, Albion and Ierne. The name Britain descends from the Latin name for Britain, Britannia or Brittānia, Old French Bretaigne and Middle English Bretayne, Breteyne. The French form replaced the Old English Breoton, Breoten, Bryten, Breten, Britannia was used by the Romans from the 1st century BC for the British Isles taken together. It is derived from the writings of the Pytheas around 320 BC. Marcian of Heraclea, in his Periplus maris exteri, described the group as αἱ Πρεττανικαὶ νῆσοι. The peoples of these islands of Prettanike were called the Πρεττανοί, Priteni is the source of the Welsh language term Prydain, Britain, which has the same source as the Goidelic term Cruithne used to refer to the early Brythonic-speaking inhabitants of Ireland. The latter were later called Picts or Caledonians by the Romans, the Greco-Egyptian scientist Ptolemy referred to the larger island as great Britain and to Ireland as little Britain in his work Almagest. The name Albion appears to have out of use sometime after the Roman conquest of Britain. After the Anglo-Saxon period, Britain was used as a term only. It was used again in 1604, when King James VI and I styled himself King of Great Brittaine, France, Great Britain refers geographically to the island of Great Britain, politically to England, Scotland and Wales in combination
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Sicily
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Sicily is the largest island in the Mediterranean Sea. It is an autonomous Region of Italy, along with surrounding minor islands, Sicily is located in the central Mediterranean Sea, south of the Italian Peninsula, from which it is separated by the narrow Strait of Messina. Its most prominent landmark is Mount Etna, the tallest active volcano in Europe, the island has a typical Mediterranean climate. The earliest archaeological evidence of activity on the island dates from as early as 12,000 BC. It became part of Italy in 1860 following the Expedition of the Thousand, a revolt led by Giuseppe Garibaldi during the Italian unification, Sicily was given special status as an autonomous region after the Italian constitutional referendum of 1946. Sicily has a rich and unique culture, especially regard to the arts, music, literature, cuisine. It is also home to important archaeological and ancient sites, such as the Necropolis of Pantalica, the Valley of the Temples, Sicily has a roughly triangular shape, earning it the name Trinacria. To the east, it is separated from the Italian mainland by the Strait of Messina, about 3 km wide in the north, and about 16 km wide in the southern part. The northern and southern coasts are each about 280 km long measured as a line, while the eastern coast measures around 180 km. The total area of the island is 25,711 km2, the terrain of inland Sicily is mostly hilly and is intensively cultivated wherever possible. Along the northern coast, the ranges of Madonie,2,000 m, Nebrodi,1,800 m. The cone of Mount Etna dominates the eastern coast, in the southeast lie the lower Hyblaean Mountains,1,000 m. The mines of the Enna and Caltanissetta districts were part of a leading sulphur-producing area throughout the 19th century, Sicily and its surrounding small islands have some highly active volcanoes. Mount Etna is the largest active volcano in Europe and still casts black ash over the island with its ever-present eruptions and it currently stands 3,329 metres high, though this varies with summit eruptions, the mountain is 21 m lower now than it was in 1981. It is the highest mountain in Italy south of the Alps, Etna covers an area of 1,190 km2 with a basal circumference of 140 km. This makes it by far the largest of the three volcanoes in Italy, being about two and a half times the height of the next largest, Mount Vesuvius. In Greek Mythology, the deadly monster Typhon was trapped under the mountain by Zeus, Mount Etna is widely regarded as a cultural symbol and icon of Sicily. The Aeolian Islands in the Tyrrhenian Sea, to the northeast of mainland Sicily form a volcanic complex, the three volcanoes of Vulcano, Vulcanello and Lipari are also currently active, although the latter is usually dormant
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Hungary
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Hungary is a unitary parliamentary republic in Central Europe. With about 10 million inhabitants, Hungary is a member state of the European Union. The official language is Hungarian, which is the most widely spoken language in Europe. Hungarys capital and largest metropolis is Budapest, a significant economic hub, major urban areas include Debrecen, Szeged, Miskolc, Pécs and Győr. His great-grandson Stephen I ascended to the throne in 1000, converting the country to a Christian kingdom, by the 12th century, Hungary became a middle power within the Western world, reaching a golden age by the 15th century. Hungarys current borders were established in 1920 by the Treaty of Trianon after World War I, when the country lost 71% of its territory, 58% of its population, following the interwar period, Hungary joined the Axis Powers in World War II, suffering significant damage and casualties. Hungary became a state of the Soviet Union, which contributed to the establishment of a four-decade-long communist dictatorship. On 23 October 1989, Hungary became again a democratic parliamentary republic, in the 21st century, Hungary is a middle power and has the worlds 57th largest economy by nominal GDP, as well as the 58th largest by PPP, out of 188 countries measured by the IMF. As a substantial actor in several industrial and technological sectors, it is both the worlds 36th largest exporter and importer of goods, Hungary is a high-income economy with a very high standard of living. It keeps up a security and universal health care system. Hungary joined the European Union in 2004 and part of the Schengen Area since 2007, Hungary is a member of the United Nations, NATO, WTO, World Bank, the AIIB, the Council of Europe and Visegrád Group. Well known for its cultural history, Hungary has been contributed significantly to arts, music, literature, sports and science. Hungary is the 11th most popular country as a tourist destination in Europe and it is home to the largest thermal water cave system, the second largest thermal lake in the world, the largest lake in Central Europe, and the largest natural grasslands in Europe. The H in the name of Hungary is most likely due to historical associations with the Huns. The rest of the word comes from the Latinized form of Medieval Greek Oungroi, according to an explanation the Greek name was borrowed from Proto-Slavic Ǫgǔri, in turn borrowed from Oghur-Turkic Onogur. Onogur was the name for the tribes who later joined the Bulgar tribal confederacy that ruled the eastern parts of Hungary after the Avars. The Hungarians likely belonged to the Onogur tribal alliance and it is possible they became its ethnic majority. The Hungarian endonym is Magyarország, composed of magyar and ország, the word magyar is taken from the name of one of the seven major semi-nomadic Hungarian tribes, magyeri
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Ophrys argolica
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Ophrys argolica, Late Spider Orchid, or Argolian Bee-orchid, is a terrestrial species of orchid native to Greece, Italy, Croatia, Cyprus, Turkey, Lebanon and Syria. The epithet argolica refers to the Argolia region of Greece, southwest of Athens, at present,7 subspecies are recognized, Ophrys argolica subsp. - Greek islands Ophrys argolica subsp, argolica - Greece Ophrys argolica subsp. Biscutella Kreutz - southern Italy, Croatia Ophrys argolica subsp, crabronifera Faurh. - central and southern Italy Ophrys argolica subsp. Elegans E. Nelson - Cyprus Ophrys argolica subsp, - western Turkey and the Greek Islands Ophrys argolica subsp
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