Tragophylloceras is an extinct genus of cephalopod belonging to the Ammonite subclass. Jurassic of Austria, Germany and Montenegro, Switzerland and the United Kingdom
The Triassic is a geologic period and system which spans 50.6 million years from the end of the Permian Period 251.9 million years ago, to the beginning of the Jurassic Period 201.3 Mya. The Triassic is the shortest period of the Mesozoic Era. Both the start and end of the period are marked by major extinction events. Triassic began in the wake of the Permian–Triassic extinction event, which left the Earth's biosphere impoverished. Therapsids and archosaurs were the chief terrestrial vertebrates during this time. A specialized subgroup of archosaurs, called dinosaurs, first appeared in the Late Triassic but did not become dominant until the succeeding Jurassic Period; the first true mammals, themselves a specialized subgroup of therapsids evolved during this period, as well as the first flying vertebrates, the pterosaurs, like the dinosaurs, were a specialized subgroup of archosaurs. The vast supercontinent of Pangaea existed until the mid-Triassic, after which it began to rift into two separate landmasses, Laurasia to the north and Gondwana to the south.
The global climate during the Triassic was hot and dry, with deserts spanning much of Pangaea's interior. However, the climate became more humid as Pangaea began to drift apart; the end of the period was marked by yet another major mass extinction, the Triassic–Jurassic extinction event, that wiped out many groups and allowed dinosaurs to assume dominance in the Jurassic. The Triassic was named in 1834 by Friedrich von Alberti, after the three distinct rock layers that are found throughout Germany and northwestern Europe—red beds, capped by marine limestone, followed by a series of terrestrial mud- and sandstones—called the "Trias"; the Triassic is separated into Early and Late Triassic Epochs, the corresponding rocks are referred to as Lower, Middle, or Upper Triassic. The faunal stages from the youngest to oldest are: During the Triassic all the Earth's land mass was concentrated into a single supercontinent centered more or less on the equator and spanning from pole to pole, called Pangaea.
From the east, along the equator, the Tethys sea penetrated Pangaea, causing the Paleo-Tethys Ocean to be closed. In the mid-Triassic a similar sea penetrated along the equator from the west; the remaining shores were surrounded by the world-ocean known as Panthalassa. All the deep-ocean sediments laid down during the Triassic have disappeared through subduction of oceanic plates; the supercontinent Pangaea was rifting during the Triassic—especially late in that period—but had not yet separated. The first nonmarine sediments in the rift that marks the initial break-up of Pangaea, which separated New Jersey from Morocco, are of Late Triassic age. S. these thick sediments comprise the Newark Group. Because a super-continental mass has less shoreline compared to one broken up, Triassic marine deposits are globally rare, despite their prominence in Western Europe, where the Triassic was first studied. In North America, for example, marine deposits are limited to a few exposures in the west, thus Triassic stratigraphy is based on organisms that lived in lagoons and hypersaline environments, such as Estheria crustaceans.
At the beginning of the Mesozoic Era, Africa was joined with Earth's other continents in Pangaea. Africa shared the supercontinent's uniform fauna, dominated by theropods and primitive ornithischians by the close of the Triassic period. Late Triassic fossils are more common in the south than north; the time boundary separating the Permian and Triassic marks the advent of an extinction event with global impact, although African strata from this time period have not been studied. During the Triassic peneplains are thought to have formed in what is now southern Sweden. Remnants of this peneplain can be traced as a tilted summit accordance in the Swedish West Coast. In northern Norway Triassic peneplains may have been buried in sediments to be re-exposed as coastal plains called strandflats. Dating of illite clay from a strandflat of Bømlo, southern Norway, have shown that landscape there became weathered in Late Triassic times with the landscape also being shaped during that time. At Paleorrota geopark, located in Rio Grande do Sul, the Santa Maria Formation and Caturrita Formations are exposed.
In these formations, one of the earliest dinosaurs, Staurikosaurus, as well as the mammal ancestors Brasilitherium and Brasilodon have been discovered. The Triassic continental interior climate was hot and dry, so that typical deposits are red bed sandstones and evaporites. There is no evidence of glaciation near either pole. Pangaea's large size limited the moderating effect of the global ocean; the strong contrast between the Pangea supercontinent and the global ocean triggered intense cross-equatorial monsoons. The Triassic may have been a dry period, but evidence exists that it was punctuated by several episodes of increased rainfall in tropical and subtropical latitudes of the Tethys Sea and its surrounding land. Sediments and fossils suggestive of a more humid climate are known from the Anisian to Ladinian of the Tethysian domain, from the Carnian and Rhaetian of a larger area that includes the Boreal domain, the North
Global Boundary Stratotype Section and Point
A Global Boundary Stratotype Section and Point, abbreviated GSSP, is an internationally agreed upon reference point on a stratigraphic section which defines the lower boundary of a stage on the geologic time scale. The effort to define GSSPs is conducted by the International Commission on Stratigraphy, a part of the International Union of Geological Sciences. Most, but not all, GSSPs are based on paleontological changes. Hence GSSPs are described in terms of transitions between different faunal stages, though far more faunal stages have been described than GSSPs; the GSSP definition effort commenced in 1977. As of 2012, 64 of the 101 stages that need a GSSP have been formally defined. A geologic section has to fulfill a set of criteria to be adapted as a GSSP by the ICS; the following list summarizes the criteria: A GSSP has to define the lower boundary of a geologic stage. The lower boundary has to be defined using a primary marker. There should be secondary markers; the horizon in which the marker appears should have minerals.
The marker has to have regional and global correlation in outcrops of the same age The marker should be independent of facies. The outcrop has to have an adequate thickness Sedimentation has to be continuous without any changes in facies The outcrop should be unaffected by tectonic and sedimentary movements, metamorphism The outcrop has to be accessible to research and free to access; this includes that the outcrop has to be located where it can be visited has to be kept in good condition, in accessible terrain, extensive enough to allow repeated sampling and open to researchers of all nationalities. The Precambrian-Cambrian boundary GSSP at Fortune Head, Newfoundland is a typical GSSP, it is set aside as a nature preserve. A continuous section is available from beds that are Precambrian into beds that are Cambrian; the boundary is set at the first appearance of a complex trace fossil Treptichnus pedum, found worldwide. The Fortune Head GSSP is unlikely to be built over. Nonetheless, Treptichnus pedum is less than ideal as a marker fossil as it is not found in every Cambrian sequence, it is not assured that it is found at the same level in every exposure.
In fact, further eroding its value as a boundary marker, it has since been identified in strata 4m below the GSSP! However, no other fossil is known. There is no radiometrically datable bed at the boundary at Fortune Head, but there is one above the boundary in similar beds nearby; these factors have led some geologists to suggest. Once a GSSP boundary has been agreed upon, a "golden spike" is driven into the geologic section to mark the precise boundary for future geologists; the first stratigraphic boundary was defined in 1977 by identifying the Silurian-Devonian boundary with a bronze plaque at a locality called Klonk, northeast of the village of Suchomasty in the Czech Republic. GSSPs are sometimes referred to as Golden Spikes; because defining a GSSP depends on finding well-preserved geologic sections and identifying key events, this task becomes more difficult as one goes farther back in time. Before 630 million years ago, boundaries on the geologic timescale are defined by reference to fixed dates, known as "Global Standard Stratigraphic Ages".
Body form European Mammal Neogene Fauna Geologic time scale New Zealand geologic time scale List of GSSPs North American Land Mammal Age Type locality Hedberg, H. D. International stratigraphic guide: A guide to stratigraphic classification and procedure, New York, John Wiley and Sons, 1976 International Stratigraphic Chart from the International Commission on Stratigraphy GSSP table with pages on each ratified GSSP from the ICS Subcommission for Stratigraphic Information USA National Park Service Washington State University Web Geological Time Machine Eon or Aeon, Math Words - An alphabetical index The Global Boundary Stratotype Section and Point: overview Chart of The Global Boundary Stratotype Sections and Points: chart Table of Global Boundary Stratotype Sections and Points with links to summary pages for each one: chart GSSPs and Continental drift 3D views Geotime chart displaying geologic time periods compared to the fossil record - Deals with chronology and classifications for laymen
In the geological timescale, the Berriasian is an age or stage of the Early Cretaceous. It is the oldest, or lowest, subdivision in the entire Cretaceous, it spanned the time between 145.0 139.8 ± 3.0 Ma. The Berriasian precedes the Valanginian; the Berriasian Stage was introduced in scientific literature by Henri Coquand in 1869. It is named after the village of Berrias in the Ardèche department of France; the non-marine English Purbeck Formation is in part of Berriasian age. In fact, the first rocks to be described of this age were the beds of the English Purbeck Formation, named as the Purbeckian by Alexandre Brongniart in 1829 following description by Henry De la Beche, William Buckland, Thomas Webster and William Henry Fitton; the base of the Berriasian, the base of the Cretaceous system, has traditionally been placed at the first appearance of fossils of the ammonite species Berriasella jacobi. But this is a species that has a stratigraphically problematic and geographically limited distribution.
A global reference profile for the Berriasian has been under active consideration by the International Subcommission on Cretaceous Stratigraphy of IUGS. A range of contender GSSP localities has been studied in detail by the ISCS's Berriasian Working Group including localities as far apart as Mexico, Tunisia and the Russian Far East. Several markers have been employed to refine correlations and to work towards defining a base for the Berriasian Stage; these include calcareous nannofossils, such as Nannoconus, ammonites, palynological data and magnetostratigraphy, notably magnetozone M19n. The calibration of these markers Nannoconus steinmannii minor, N. kamptneri minor, Calpionella alpina, within fixed magnetozones give greater precision in trying to identify the best position for a boundary. In June 2016, the Berriasian Working Group voted to adopt Calpionella alpina as the primary marker for the base of the Berriasian Stage. In the western part of the ocean of Tethys, the Berriasian consists of four ammonite biozones, from top to bottom: Thurmanniceras otopeta Subthurmannia boissieri Tirnovella occitanica Berriasella jacobi/Pseudosubplanites grandisThe top of the Berriasian stage is defined by the base of the Valanginian, fixed at the first appearance of calpionellid species Calpionellites darderi.
This is just a little below the first appearance of the ammonite species Thurmanniceras pertransiens. Gradstein, F. M.. G. & Smith, A. G.. GeoWhen Database - Berriasian Jurassic-Cretaceous timescale, at the website of the subcommission for stratigraphic information of the ICS Stratigraphic chart of the Lower Cretaceous, at the website of Norges Network of offshore records of geology and stratigraphy
England is a country, part of the United Kingdom. It shares land borders with Wales to Scotland to the north-northwest; the Irish Sea lies west of England and the Celtic Sea lies to the southwest. England is separated from continental Europe by the North Sea to the east and the English Channel to the south; the country covers five-eighths of the island of Great Britain, which lies in the North Atlantic, includes over 100 smaller islands, such as the Isles of Scilly and the Isle of Wight. The area now called England was first inhabited by modern humans during the Upper Palaeolithic period, but takes its name from the Angles, a Germanic tribe deriving its name from the Anglia peninsula, who settled during the 5th and 6th centuries. England became a unified state in the 10th century, since the Age of Discovery, which began during the 15th century, has had a significant cultural and legal impact on the wider world; the English language, the Anglican Church, English law – the basis for the common law legal systems of many other countries around the world – developed in England, the country's parliamentary system of government has been adopted by other nations.
The Industrial Revolution began in 18th-century England, transforming its society into the world's first industrialised nation. England's terrain is chiefly low hills and plains in central and southern England. However, there is upland and mountainous terrain in the west; the capital is London, which has the largest metropolitan area in both the United Kingdom and the European Union. England's population of over 55 million comprises 84% of the population of the United Kingdom concentrated around London, the South East, conurbations in the Midlands, the North West, the North East, Yorkshire, which each developed as major industrial regions during the 19th century; the Kingdom of England – which after 1535 included Wales – ceased being a separate sovereign state on 1 May 1707, when the Acts of Union put into effect the terms agreed in the Treaty of Union the previous year, resulting in a political union with the Kingdom of Scotland to create the Kingdom of Great Britain. In 1801, Great Britain was united with the Kingdom of Ireland to become the United Kingdom of Great Britain and Ireland.
In 1922 the Irish Free State seceded from the United Kingdom, leading to the latter being renamed the United Kingdom of Great Britain and Northern Ireland. The name "England" is derived from the Old English name Englaland, which means "land of the Angles"; the Angles were one of the Germanic tribes that settled in Great Britain during the Early Middle Ages. The Angles came from the Anglia peninsula in the Bay of Kiel area of the Baltic Sea; the earliest recorded use of the term, as "Engla londe", is in the late-ninth-century translation into Old English of Bede's Ecclesiastical History of the English People. The term was used in a different sense to the modern one, meaning "the land inhabited by the English", it included English people in what is now south-east Scotland but was part of the English kingdom of Northumbria; the Anglo-Saxon Chronicle recorded that the Domesday Book of 1086 covered the whole of England, meaning the English kingdom, but a few years the Chronicle stated that King Malcolm III went "out of Scotlande into Lothian in Englaland", thus using it in the more ancient sense.
According to the Oxford English Dictionary, its modern spelling was first used in 1538. The earliest attested reference to the Angles occurs in the 1st-century work by Tacitus, Germania, in which the Latin word Anglii is used; the etymology of the tribal name itself is disputed by scholars. How and why a term derived from the name of a tribe, less significant than others, such as the Saxons, came to be used for the entire country and its people is not known, but it seems this is related to the custom of calling the Germanic people in Britain Angli Saxones or English Saxons to distinguish them from continental Saxons of Old Saxony between the Weser and Eider rivers in Northern Germany. In Scottish Gaelic, another language which developed on the island of Great Britain, the Saxon tribe gave their name to the word for England. An alternative name for England is Albion; the name Albion referred to the entire island of Great Britain. The nominally earliest record of the name appears in the Aristotelian Corpus the 4th-century BC De Mundo: "Beyond the Pillars of Hercules is the ocean that flows round the earth.
In it are two large islands called Britannia. But modern scholarly consensus ascribes De Mundo not to Aristotle but to Pseudo-Aristotle, i.e. it was written in the Graeco-Roman period or afterwards. The word Albion or insula Albionum has two possible origins, it either derives from a cognate of the Latin albus meaning white, a reference to the white cliffs of Dover or from the phrase the "island of the Albiones" in the now lost Massaliote Periplus, attested through Avienus' Ora Maritima to which the former served as a source. Albion is now applied to England in a more poetic capacity. Another romantic name for England is Loegria, related to the Welsh word for England and made popular by its use in Arthurian legend; the earliest known evidence of human presence in the area now known as England was that of Homo antecessor, dating to approximate
Amphibians are ectothermic, tetrapod vertebrates of the class Amphibia. Modern amphibians are all Lissamphibia, they inhabit a wide variety of habitats, with most species living within terrestrial, arboreal or freshwater aquatic ecosystems. Thus amphibians start out as larvae living in water, but some species have developed behavioural adaptations to bypass this; the young undergo metamorphosis from larva with gills to an adult air-breathing form with lungs. Amphibians use their skin as a secondary respiratory surface and some small terrestrial salamanders and frogs lack lungs and rely on their skin, they are superficially similar to lizards but, along with mammals and birds, reptiles are amniotes and do not require water bodies in which to breed. With their complex reproductive needs and permeable skins, amphibians are ecological indicators; the earliest amphibians evolved in the Devonian period from sarcopterygian fish with lungs and bony-limbed fins, features that were helpful in adapting to dry land.
They diversified and became dominant during the Carboniferous and Permian periods, but were displaced by reptiles and other vertebrates. Over time, amphibians shrank in size and decreased in diversity, leaving only the modern subclass Lissamphibia; the three modern orders of amphibians are Anura and Apoda. The number of known amphibian species is 8,000, of which nearly 90% are frogs; the smallest amphibian in the world is a frog from New Guinea with a length of just 7.7 mm. The largest living amphibian is the 1.8 m Chinese giant salamander, but this is dwarfed by the extinct 9 m Prionosuchus from the middle Permian of Brazil. The study of amphibians is called batrachology, while the study of both reptiles and amphibians is called herpetology; the word "amphibian" is derived from the Ancient Greek term ἀμφίβιος, which means "both kinds of life", ἀμφί meaning "of both kinds" and βιος meaning "life". The term was used as a general adjective for animals that could live on land or in water, including seals and otters.
Traditionally, the class Amphibia includes all tetrapod vertebrates. Amphibia in its widest sense was divided into three subclasses, two of which are extinct: Subclass Lepospondyli† Subclass Temnospondyli† Subclass Lissamphibia Salientia: Jurassic to present—6,200 current species in 53 families Caudata: Jurassic to present—652 current species in 9 families Gymnophiona: Jurassic to present—192 current species in 10 families The actual number of species in each group depends on the taxonomic classification followed; the two most common systems are the classification adopted by the website AmphibiaWeb, University of California and the classification by herpetologist Darrel Frost and the American Museum of Natural History, available as the online reference database "Amphibian Species of the World". The numbers of species cited above follows Frost and the total number of known amphibian species as of March 31, 2019 is 8,000, of which nearly 90% are frogs. With the phylogenetic classification, the taxon Labyrinthodontia has been discarded as it is a polyparaphyletic group without unique defining features apart from shared primitive characteristics.
Classification varies according to the preferred phylogeny of the author and whether they use a stem-based or a node-based classification. Traditionally, amphibians as a class are defined as all tetrapods with a larval stage, while the group that includes the common ancestors of all living amphibians and all their descendants is called Lissamphibia; the phylogeny of Paleozoic amphibians is uncertain, Lissamphibia may fall within extinct groups, like the Temnospondyli or the Lepospondyli, in some analyses in the amniotes. This means that advocates of phylogenetic nomenclature have removed a large number of basal Devonian and Carboniferous amphibian-type tetrapod groups that were placed in Amphibia in Linnaean taxonomy, included them elsewhere under cladistic taxonomy. If the common ancestor of amphibians and amniotes is included in Amphibia, it becomes a paraphyletic group. All modern amphibians are included in the subclass Lissamphibia, considered a clade, a group of species that have evolved from a common ancestor.
The three modern orders are Anura and Gymnophiona. It has been suggested that salamanders arose separately from a Temnospondyl-like ancestor, that caecilians are the sister group of the advanced reptiliomorph amphibians, thus of amniotes. Although the fossils of several older proto-frogs with primitive characteristics are known, the oldest "true frog" is Prosalirus bitis, from the Early Jurassic Kayenta Formation of Arizona, it is anatomically similar to modern frogs. The oldest known caecilian is another Early Jurassic species, Eocaecilia micropodia from Arizona; the earliest salamander is Beiyanerpeton jianpingensis from the Late Jurassic of northeastern China. Authorities disagree as to whether Salientia is a superorder that includes the order Anura, or whether