1.
Taxonomy (biology)
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Taxonomy is the science of defining groups of biological organisms on the basis of shared characteristics and giving names to those groups. The exact definition of taxonomy varies from source to source, but the core of the remains, the conception, naming. There is some disagreement as to whether biological nomenclature is considered a part of taxonomy, the broadest meaning of taxonomy is used here. The word taxonomy was introduced in 1813 by Candolle, in his Théorie élémentaire de la botanique, the term alpha taxonomy is primarily used today to refer to the discipline of finding, describing, and naming taxa, particularly species. In earlier literature, the term had a different meaning, referring to morphological taxonomy, ideals can, it may be said, never be completely realized. They have, however, a value of acting as permanent stimulants. Some of us please ourselves by thinking we are now groping in a beta taxonomy, turrill thus explicitly excludes from alpha taxonomy various areas of study that he includes within taxonomy as a whole, such as ecology, physiology, genetics, and cytology. He further excludes phylogenetic reconstruction from alpha taxonomy, thus, Ernst Mayr in 1968 defined beta taxonomy as the classification of ranks higher than species. This activity is what the term denotes, it is also referred to as beta taxonomy. How species should be defined in a group of organisms gives rise to practical and theoretical problems that are referred to as the species problem. The scientific work of deciding how to define species has been called microtaxonomy, by extension, macrotaxonomy is the study of groups at higher taxonomic ranks, from subgenus and above only, than species. While some descriptions of taxonomic history attempt to date taxonomy to ancient civilizations, earlier works were primarily descriptive, and focused on plants that were useful in agriculture or medicine. There are a number of stages in scientific thinking. Early taxonomy was based on criteria, the so-called artificial systems. Later came systems based on a complete consideration of the characteristics of taxa, referred to as natural systems, such as those of de Jussieu, de Candolle and Bentham. The publication of Charles Darwins Origin of Species led to new ways of thinking about classification based on evolutionary relationships and this was the concept of phyletic systems, from 1883 onwards. This approach was typified by those of Eichler and Engler, the advent of molecular genetics and statistical methodology allowed the creation of the modern era of phylogenetic systems based on cladistics, rather than morphology alone. Taxonomy has been called the worlds oldest profession, and naming and classifying our surroundings has likely been taking place as long as mankind has been able to communicate
2.
Animal
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Animals are multicellular, eukaryotic organisms of the kingdom Animalia. The animal kingdom emerged as a clade within Apoikozoa as the group to the choanoflagellates. Animals are motile, meaning they can move spontaneously and independently at some point in their lives and their body plan eventually becomes fixed as they develop, although some undergo a process of metamorphosis later in their lives. All animals are heterotrophs, they must ingest other organisms or their products for sustenance, most known animal phyla appeared in the fossil record as marine species during the Cambrian explosion, about 542 million years ago. Animals can be divided broadly into vertebrates and invertebrates, vertebrates have a backbone or spine, and amount to less than five percent of all described animal species. They include fish, amphibians, reptiles, birds and mammals, the remaining animals are the invertebrates, which lack a backbone. These include molluscs, arthropods, annelids, nematodes, flatworms, cnidarians, ctenophores, the study of animals is called zoology. The word animal comes from the Latin animalis, meaning having breath, the biological definition of the word refers to all members of the kingdom Animalia, encompassing creatures as diverse as sponges, jellyfish, insects, and humans. Aristotle divided the world between animals and plants, and this was followed by Carl Linnaeus, in the first hierarchical classification. In Linnaeuss original scheme, the animals were one of three kingdoms, divided into the classes of Vermes, Insecta, Pisces, Amphibia, Aves, and Mammalia. Since then the last four have all been subsumed into a single phylum, in 1874, Ernst Haeckel divided the animal kingdom into two subkingdoms, Metazoa and Protozoa. The protozoa were later moved to the kingdom Protista, leaving only the metazoa, thus Metazoa is now considered a synonym of Animalia. Animals have several characteristics that set apart from other living things. Animals are eukaryotic and multicellular, which separates them from bacteria and they are heterotrophic, generally digesting food in an internal chamber, which separates them from plants and algae. They are also distinguished from plants, algae, and fungi by lacking cell walls. All animals are motile, if only at life stages. In most animals, embryos pass through a stage, which is a characteristic exclusive to animals. With a few exceptions, most notably the sponges and Placozoa and these include muscles, which are able to contract and control locomotion, and nerve tissues, which send and process signals
3.
Chordate
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Chordates are deuterostomes, as during the embryo development stage the anus forms before the mouth. They are also bilaterally symmetric coelomates, in the case of vertebrate chordates, the notochord is usually replaced by a vertebral column during development, and they may have body plans organized via segmentation. There are also additional extinct taxa, the Vertebrata are sometimes considered as a subgroup of the clade Craniata, consisting of chordates with a skull, the Craniata and Tunicata compose the clade Olfactores. Of the more than 65,000 living species of chordates, the worlds largest and fastest animals, the blue whale and peregrine falcon respectively, are chordates, as are humans. Fossil chordates are known from at least as early as the Cambrian explosion, Hemichordata, which includes the acorn worms, has been presented as a fourth chordate subphylum, but it now is usually treated as a separate phylum. The Hemichordata, along with the Echinodermata, form the Ambulacraria, the Chordata and Ambulacraria form the superphylum Deuterostomia, composed of the deuterostomes. Attempts to work out the relationships of the chordates have produced several hypotheses. All of the earliest chordate fossils have found in the Early Cambrian Chengjiang fauna. Because the fossil record of early chordates is poor, only molecular phylogenetics offers a prospect of dating their emergence. However, the use of molecular phylogenetics for dating evolutionary transitions is controversial and it has also proved difficult to produce a detailed classification within the living chordates. Attempts to produce family trees shows that many of the traditional classes are paraphyletic. While this has been known since the 19th century, an insistence on only monophyletic taxa has resulted in vertebrate classification being in a state of flux. Although the name Chordata is attributed to William Bateson, it was already in prevalent use by 1880, ernst Haeckel described a taxon comprising tunicates, cephalochordates, and vertebrates in 1866. Though he used the German vernacular form, it is allowed under the ICZN code because of its subsequent latinization, among the vertebrate sub-group of chordates the notochord develops into the spine, and in wholly aquatic species this helps the animal to swim by flexing its tail. In fish and other vertebrates, this develops into the spinal cord, the pharynx is the part of the throat immediately behind the mouth. In fish, the slits are modified to form gills, a muscular tail that extends backwards behind the anus. This is a groove in the wall of the pharynx. In filter-feeding species it produces mucus to gather food particles, which helps in transporting food to the esophagus and it also stores iodine, and may be a precursor of the vertebrate thyroid gland
4.
Actinopterygii
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Actinopterygii /ˌæktᵻnˌɒptəˈrɪdʒi. aɪ/, or the ray-finned fishes, constitute a class or subclass of the bony fishes. These actinopterygian fin rays attach directly to the proximal or basal skeletal elements, the radials, numerically, actinopterygians are the dominant class of vertebrates, comprising nearly 99% of the over 30,000 species of fish. They are ubiquitous throughout freshwater and marine environments from the sea to the highest mountain streams. Extant species can range in size from Paedocypris, at 8 mm, to the ocean sunfish, at 2,300 kg. Ray-finned fishes occur in variant forms. The main features of a typical ray-finned fish are shown in the diagram at the left. In nearly all ray-finned fish, the sexes are separate, development then proceeds with a free-swimming larval stage. However other patterns of ontogeny exist, with one of the commonest being sequential hermaphroditism, in most cases this involves protogyny, fish starting life as females and converting to males at some stage, triggered by some internal or external factor. This may be advantageous as females become less prolific as they age while male fecundity increases with age, protandry, where a fish converts from male to female, is much less common than protogyny. Most families use external rather than internal fertilization, of the oviparous teleosts, most do not provide parental care. Viviparity is relatively rare and is found in about 6% of teleost species, male territoriality preadapts a species for evolving male parental care. There are a few examples of fish that self-fertilise, the mangrove rivulus is an amphibious, simultaneous hermaphrodite, producing both eggs and spawn and having internal fertilisation. This mode of reproduction may be related to the habit of spending long periods out of water in the mangrove forests it inhabits. Males are occasionally produced at temperatures below 19 °C and can fertilise eggs that are spawned by the female. This maintains genetic variability in a species that is otherwise highly inbred, the earliest known fossil actinopterygiian is Andreolepis hedei, dating back 420 million years. Remains have been found in Russia, Sweden, and Estonia, actinopterygians are divided into the subclasses Chondrostei and Neopterygii. The Neopterygii, in turn, are divided into the infraclasses Holostei and Teleostei, during the Mesozoic and Cenozoic the teleosts in particular diversified widely, and as a result, 96% of all known fish species are teleosts. The cladogram shows the groups of actinopterygians and their relationship to the terrestrial vertebrates that evolved from a related group of fish
5.
Salmonidae
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Salmonidae is a family of ray-finned fish, the only living family currently placed in the order Salmoniformes. It includes salmon, trout, chars, freshwater whitefishes, and graylings, the Atlantic salmon and trout of the genus Salmo give the family and order their names. Salmonids have a primitive appearance among the teleost fish, with the pelvic fins being placed far back. They are slender fish, with rounded scales and forked tails and their mouths contain a single row of sharp teeth. Although the smallest species is just 13 cm long as an adult, most are much larger, all salmonids spawn in fresh water, but in many cases, the fish spend most of their lives at sea, returning to the rivers only to reproduce. This lifecycle is described as anadromous and they are predators, feeding on small crustaceans, aquatic insects, and smaller fish. Current salmonids comprise three lineages, taxonomically treated as subfamilies, whitefish, graylings, and the char, trout, generally, all three lineages are accepted to allocate a suite of derived traits indicating a monophyletic group. This genus shares traits found in the Salmoninae, whitefish, hence, E. driftwoodensis is an archaic salmonid, representing an important stage in salmonid evolution. A gap appears in the fossil record after E. driftwoodensis until about seven million years ago, in the late Miocene. Several of these appear to be Oncorhynchus—the current genus for Pacific salmon. Consequently, the split between Oncorhynchus and Salmo must have occurred well before the Pliocene, suggestions have gone back as far as the early Miocene. Based on the most current evidence, salmonids diverged from the rest of teleost fish no later than 88 million years ago and this divergence was marked by a whole-genome duplication event in the ancestral salmonid, where the diploid ancestor became tetraploid. Extant salmonids all show evidence of partial tetraploidy, as show the genome has undergone selection to regain a diploid state. Work done in the trout has shown that the genome is still partially-tetraploid. This pattern of partial tetraploidy is thought to be reflected in the rest of extant salmonids, the first fossil species representing a true salmonid fish does not appear until the middle Eocene. This fossil is also younger than the proposed salmonid divergence from the rest of the teleost fishes. This means that the salmonids have a ghost lineage of approximately 33 million years and this more precise dating and examination of the salmonid whole-genome duplication event has allowed more speculation on the radiation of species within the group. Historically, the duplication event was thought to be the reason for the variation within Salmonidae
6.
Oncorhynchus
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Oncorhynchus is a genus of fish in the family Salmonidae, it contains the Pacific salmon and Pacific trout. The name of the genus is derived from the Greek onkos and rynchos, Salmon and trout with native ranges in waters draining to the Pacific Ocean are members of the genus. Their range extends from Beringia southwards, roughly to Taiwan in the west, in North America, some subspecies of O. Several species of Oncorhynchus have been introduced into waters around the globe establishing self-sustaining wild populations. The six Pacific salmons of Oncorhynchus are anadromous and semelparous, migration can be affected by parasites. Infected individuals can become weak and probably have shortened lifespans, infection with parasites creates an effect known as culling whereby fish that are infected are less likely to complete the migration. Anadromous forms of Oncorhynchus mykiss known as steelhead are iteroparus, the Coastal cutthroat trout form of Oncorhynchus clarki is considered semi-anadromous as it spends short periods of time in marine environments. Several late Miocene trout-like fossils in Idaho, in the Clarkia Lake beds, consequently, the split between Oncorhynchus and Salmo must have occurred well before the Pliocene. Suggested dates ions have gone back as far as the early Miocene, one fossil species assigned to this genus, O. rastrosus, the sabertooth salmon, is a 9-foot -long species known from Late Miocene to Pleistocene fossils. Speciation among Oncorhynchus has been examined for decades, and a tree is not yet completely developed for the Pacific salmonids. Mitochondrial DNA research has been completed on a variety of Pacific trout and salmonid species, Chum, pink and sockeye salmon lineages are generally agreed to have diverged in the sequence after other species. Montgomery discusses the pattern of the record as compared to tectonic shifts in the plates of the Pacific Northwest of America. The divergence in Onchorhyncus lineages appear to follow the uprising of the Pacific Rim, the climatic and habitat changes that would follow such a geologic event are discussed, in the context of potential stressors leading to adaptation and speciation. One interesting case involving speciation with salmon is that of the kokanee sockeye, kokanee sockeye evolve differently from anadromous sockeye—they reach the level of biological species. Biological species—as opposed to morphological species—are defined by the capacity to maintain themselves in sympatry as independent genetic entities and this definition can be vexing because it apparently applies only to sympatry, and this limitation makes the definition difficult to apply. Examples in Washington State, Canada, and elsewhere have two living in the same lake, but spawning in different substrates at different times. There is no pressure to compete or interbreed and these types of kokanee salmon show the principal attributes of a biological species, they are reproductively isolated and show strong resources partitioning. A general decline in overall Pacific salmon populations began in the mid 19th century, as the result of western expansion and development in the U. S. experts estimate salmon populations in the Columbia River basin had been reduced to less than 20% of their pre-1850 levels by 1933
7.
Binomial nomenclature
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Such a name is called a binomial name, a binomen, binominal name or a scientific name, more informally it is also called a Latin name. The first part of the name identifies the genus to which the species belongs, for example, humans belong to the genus Homo and within this genus to the species Homo sapiens. The formal introduction of system of naming species is credited to Carl Linnaeus. But Gaspard Bauhin, in as early as 1623, had introduced in his book Pinax theatri botanici many names of genera that were adopted by Linnaeus. Although the general principles underlying binomial nomenclature are common to these two codes, there are differences, both in the terminology they use and in their precise rules. Similarly, both parts are italicized when a binomial name occurs in normal text, thus the binomial name of the annual phlox is now written as Phlox drummondii. In scientific works, the authority for a name is usually given, at least when it is first mentioned. In zoology Patella vulgata Linnaeus,1758, the original name given by Linnaeus was Fringilla domestica, the parentheses indicate that the species is now considered to belong in a different genus. The ICZN does not require that the name of the person who changed the genus be given, nor the date on which the change was made, in botany Amaranthus retroflexus L. – L. is the standard abbreviation used in botany for Linnaeus. – Linnaeus first named this bluebell species Scilla italica, Rothmaler transferred it to the genus Hyacinthoides, the ICN does not require that the dates of either publication be specified. Prior to the adoption of the binomial system of naming species. Together they formed a system of polynomial nomenclature and these names had two separate functions. First, to designate or label the species, and second, to be a diagnosis or description, such polynomial names may sometimes look like binomials, but are significantly different. For example, Gerards herbal describes various kinds of spiderwort, The first is called Phalangium ramosum, Branched Spiderwort, is aptly termed Phalangium Ephemerum Virginianum, Soon-Fading Spiderwort of Virginia. The Latin phrases are short descriptions, rather than identifying labels, the Bauhins, in particular Caspar Bauhin, took some important steps towards the binomial system, by pruning the Latin descriptions, in many cases to two words. The adoption by biologists of a system of binomial nomenclature is due to Swedish botanist and physician Carl von Linné. It was in his 1753 Species Plantarum that he first began using a one-word trivial name together with a generic name in a system of binomial nomenclature. This trivial name is what is now known as an epithet or specific name
8.
Synonym (taxonomy)
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For example, Linnaeus was the first to give a scientific name to the Norway spruce, which he called Pinus abies. This name is no longer in use, it is now a synonym of the current scientific name which is Picea abies, unlike synonyms in other contexts, in taxonomy a synonym is not interchangeable with the name of which it is a synonym. In taxonomy, synonyms are not equals, but have a different status, for any taxon with a particular circumscription, position, and rank, only one scientific name is considered to be the correct one at any given time. A synonym cannot exist in isolation, it is always an alternative to a different scientific name, given that the correct name of a taxon depends on the taxonomic viewpoint used a name that is one taxonomists synonym may be another taxonomists correct name. Synonyms may arise whenever the same taxon is described and named more than once, independently. They may also arise when existing taxa are changed, as when two taxa are joined to one, a species is moved to a different genus. To the general user of scientific names, in such as agriculture, horticulture, ecology, general science. A synonym is a name that was used as the correct scientific name but which has been displaced by another scientific name. Thus Oxford Dictionaries Online defines the term as a name which has the same application as another. In handbooks and general texts, it is useful to have mentioned as such after the current scientific name. Synonyms used in this way may not always meet the strict definitions of the synonym in the formal rules of nomenclature which govern scientific names. Changes of scientific name have two causes, they may be taxonomic or nomenclatural, a name change may be caused by changes in the circumscription, position or rank of a taxon, representing a change in taxonomic, scientific insight. A name change may be due to purely nomenclatural reasons, that is, based on the rules of nomenclature, the earliest such name is called the senior synonym, while the later name is the junior synonym. One basic principle of zoological nomenclature is that the earliest correctly published name, synonyms are important because if the earliest name cannot be used, then the next available junior synonym must be used for the taxon. Objective synonyms refer to taxa with the type and same rank. For example, John Edward Gray published the name Antilocapra anteflexa in 1855 for a species of pronghorn, however, it is now commonly accepted that his specimen was an unusual individual of the species Antilocapra americana published by George Ord in 1815. Ords name thus takes precedence, with Antilocapra anteflexa being a subjective synonym. Objective synonyms are common at the level of genera, because for various reasons two genera may contain the type species, these are objective synonyms
9.
Peter Simon Pallas
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Peter Simon Pallas was a German zoologist and botanist who worked in Russia. Pallas was born in Berlin, the son of Professor of Surgery Simon Pallas and he studied with private tutors and took an interest in natural history, later attending the University of Halle and the University of Göttingen. In 1760, he moved to the University of Leiden and passed his doctors degree at the age of 19, Pallas traveled throughout the Netherlands and to London, improving his medical and surgical knowledge. He then settled at The Hague, and his new system of classification was praised by Georges Cuvier. Pallas wrote Miscellanea Zoologica, which included descriptions of several new to science which he had discovered in the Dutch museum collections. A planned voyage to southern Africa and the East Indies fell through when his father recalled him to Berlin, there, he began work on his Spicilegia Zoologica. He explored the Caspian Sea, the Ural and Altai Mountains, the regular reports which Pallas sent to St Petersburg were collected and published as Reise durch verschiedene Provinzen des Russischen Reichs. They covered a range of topics, including geology and mineralogy, reports on the native peoples and their religions. In 1776, Pallas was elected a member of the Royal Swedish Academy of Sciences. Pallas settled in St Petersburg, becoming a favourite of Catherine II and teaching history to the Grand Dukes Alexander. He was provided with the plants collected by other naturalists to compile the Flora Rossica, a Russian flora, and started work on his Zoographica Rosso-Asiatica and he also published an account of Johann Anton Güldenstädts travels in the Caucasus. The Empress bought Pallass large natural history collection for 2,000 rubles,500 more than his asking price, during this period, Pallas helped plan the Mulovsky expedition, which was cancelled in October 1787. Between 1793 and 1794, Pallas led an expedition to southern Russia, visiting the Crimea. He was accompanied by his daughter and his new wife, an artist, servants, in February 1793, they travelled to Saratov and then downriver to Tsaritsyn. They spent the spring exploring the country to the east, and in August travelled along the banks of the Caspian Sea, in September, they travelled to the Crimea, wintering in Simferopol. Pallas spent the spring of 1794 exploring to the southeast, and in July travelled up the valley of the Dnieper, Pallas gave his account of the journey in his P. S. Pallas Bemerkungen auf einer Reise in die Südlichen Statthalterschaften des Russischen Reichs. Catherine II gave him an estate at Simferopol, where Pallas lived until the death of his second wife in 1810. He was then granted permission to leave Russia by Emperor Alexander, and returned to Berlin and his grave is preserved in the Protestant Friedhof I der Jerusalems- und Neuen Kirchengemeinde in Berlin-Kreuzberg, south of Hallesches Tor
10.
John Richardson (naturalist)
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Sir John Richardson was a Scottish naval surgeon, naturalist and arctic explorer. He studied medicine at Edinburgh University, and became a surgeon in the navy in 1807 and he traveled with John Franklin in search of the Northwest Passage on the Coppermine Expedition of 1819–1822. Richardson wrote the sections on geology, botany and ichthyology for the account of the expedition. Franklin and Richardson returned to Canada in 1825 and went overland by fur trade routes to the mouth of the Mackenzie River, Franklin was to go as far west as possible and Richardson was to go east to the mouth of the Coppermine River. These were the known points on the central coast and had been reached in 1793 and 1771 respectively. He had with him two specially-built boats which were more ocean-worthy than the voyageur canoes used by Franklin on his previous expedition and they gave their names to the Dolphin and Union Strait near the end of his route. His journey was successful and he reached his furthest east the same day that Franklin reached his furthest west and he abandoned his boats at Bloody Falls and trekked overland to Fort Franklin which he reached three weeks before Franklin. Together they had surveyed 1,878 miles of previously unmapped coast and he traveled with John Rae on an unsuccessful search for Franklin in 1848–49, describing it in An Arctic Searching Expedition. He retired to the Lake District in 1855, and is buried at St Oswalds Church, Grasmere. and Arctic Ordeal, the National Marine Biological Library at the Marine Biological Association retains some original illustrations used by Richardson in preparation for the second edition of Yarrells book. Richardson is commemorated in the names of four species of reptiles, Eremiascincus richardsonii, Hemidactylus richardsonii, Myron richardsonii. Works by John Richardson at Project Gutenberg Works by John Richardson at Faded Page Works by or about John Richardson at Internet Archive Works by John Richardson at LibriVox Houston, John Richardson Chisholm, Hugh, ed. Richardson, Sir John
11.
David Starr Jordan
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David Starr Jordan was an American ichthyologist, educator, eugenicist, and peace activist. He was president of Indiana University and was the president of Stanford University. Jordan was born in Gainesville, New York, and grew up on a farm in upstate New York and his parents made the unorthodox decision to educate him at a local girls high school. He was inspired by Louis Agassiz to pursue his studies in ichthyology and he was part of the pioneer class of undergraduates at Cornell University, graduating in 1872 with a Masters degree in botany. He obtained his education from Butler University. His first wife Susan Bowen died after 10 years of marriage, Jordan and his second wife had four children. Jordan initially taught natural history courses at several small Midwestern colleges and he was then accepted into the natural history faculty of Indiana University Bloomington as a professor of zoology in 1879. He improved the finances and public image, doubled its enrollment. In March 1891, he was approached by Leland and Jane Stanford, Jordan arrived at Stanford in June 1891 and immediately set about recruiting faculty for the universitys planned September opening. Pressed for time, he drew heavily on his own acquaintance and that first year at Stanford he was instrumental in establishing the universitys Hopkins Marine Station. He served Stanford as president until 1913 and then chancellor until his retirement in 1916, while chancellor, he was elected president of the National Education Association. Jordan was a member in the Bohemian Club and the University Club in San Francisco, Jordan served as a Director of the Sierra Club from 1892 to 1903. In retirement, Jordan served as an expert witness for the defense in the 1925 Scopes Trial. He continued promoting his views on eugenics, in 1928 Jordan served on the initial board of trustees of the Human Betterment Foundation, a eugenics organization that advocated compulsory sterilization legislation in the United States. In 1905, Jordan launched an apparent coverup of the murder by poisoning of Jane Stanford, while vacationing in Oahu, Stanford had suddenly died of strychnine poisoning, according to the local coroner’s jury. His motive for doing this has been a subject of speculation, one possibility is that he was simply acting to protect the reputation of the university, its finances were precarious and a scandal might have damaged fundraising. He had written the president of Stanfords board of trustees offering several alternate explanations for Mrs. Stanfords death, given that Mrs. Jordans version of Mrs. Stanfords demise was largely accepted until the appearance of several publications in 2003 emphasizing the evidence that she was murdered. Jordan promoted the concept of improving human genetics, through removal from the pool of those deemed unworthy to reproduce
12.
Barton Warren Evermann
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Barton Warren Evermann was an American ichthyologist. He was born in Monroe County, Iowa, and graduated from Indiana University in 1886, for 10 years, he served as teacher and superintendent of schools in Indiana and California. He was professor of biology at the Indiana State University in Terre Haute from 1886 to 1891 and he lectured at Stanford University in 1893-94, at Cornell in 1900-03, and at Yale in 1903-06. He died in Berkeley, California, aged 78, in the early 20th century, as director of the California Academy of Sciences in San Francisco, he promoted research on the Revillagigedo Islands off the Pacific coast of Mexico. Mount Evermann on Socorro Island, the highest peak of the archipelago, was named in his honor, Evermann was United States fur seal commissioner in 1892 and became chairman of the fur seal board in 1908. In collaboration with D. S. Jordan, Evermann contributed to American Food and his publications include bulletins and reports of the United States Fish Commission and contributions to the proceedings of various societies. Sabertooth fish This article incorporates text from a now in the public domain, Gilman. Thurston, H. T. Colby, F. M. eds. article name needed
13.
Trout
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Trout is the common name for a number of species of freshwater fish belonging to the genera Oncorhynchus, Salmo and Salvelinus, all of the subfamily Salmoninae of the family Salmonidae. The word trout is also used as part of the name of some fish such as Cynoscion nebulosus. Trout are closely related to salmon and char, species termed salmon, a rainbow trout that spends time in the ocean is called a steelhead. Arctic char and brook trout are part of the char family, Trout are an important food source for humans and wildlife including brown bears, birds of prey such as eagles, and other animals. They are classified as oily fish, mostly, these colors and patterns form as camouflage, based on the surroundings, and will change as the fish moves to different habitats. In general trout that are about to breed have extremely intense coloration and they can look like an entirely different fish outside of spawning season. It is virtually impossible to define a color pattern as belonging to a specific breed, however, in general, wild fish are claimed to have more vivid colors. Trout have fins entirely without spines, and all of them have an adipose fin along the back. The pelvic fins sit well back on the body, on side of the anus. The swim bladder is connected to the esophagus, allowing for gulping or rapid expulsion of air, unlike many other physostome fish, trout do not use their bladder as an auxiliary device for oxygen uptake, relying solely on their gills. There are many species, and even more populations, that are isolated from each other, the trout found in the eastern United States are a good example of this. Lake trout, like brook trout, belong to the char genus, Lake trout inhabit many of the larger lakes in North America, and live much longer than rainbow trout, which have an average maximum lifespan of 7 years. Lake trout can live many decades, and can grow to more than 30 kilograms, Trout are usually found in cool, clear streams and lakes, although many of the species have anadromous strains as well. Young trout are referred to as troutlet, troutling or fry and they are distributed naturally throughout North America, northern Asia and Europe. Several species of trout were introduced to Australia and New Zealand by amateur fishing enthusiasts in the 19th century, the introduced species included brown trout from England and rainbow trout from California. The rainbow trout were a strain, generally accepted as coming from Sonoma Creek. The rainbow trout of New Zealand still show the tendency to run up rivers in winter to spawn. In Australia the rainbow trout was introduced in 1894 from New Zealand and is a popular gamefish in recreational angling
14.
Salmonid
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Salmonidae is a family of ray-finned fish, the only living family currently placed in the order Salmoniformes. It includes salmon, trout, chars, freshwater whitefishes, and graylings, the Atlantic salmon and trout of the genus Salmo give the family and order their names. Salmonids have a primitive appearance among the teleost fish, with the pelvic fins being placed far back. They are slender fish, with rounded scales and forked tails and their mouths contain a single row of sharp teeth. Although the smallest species is just 13 cm long as an adult, most are much larger, all salmonids spawn in fresh water, but in many cases, the fish spend most of their lives at sea, returning to the rivers only to reproduce. This lifecycle is described as anadromous and they are predators, feeding on small crustaceans, aquatic insects, and smaller fish. Current salmonids comprise three lineages, taxonomically treated as subfamilies, whitefish, graylings, and the char, trout, generally, all three lineages are accepted to allocate a suite of derived traits indicating a monophyletic group. This genus shares traits found in the Salmoninae, whitefish, hence, E. driftwoodensis is an archaic salmonid, representing an important stage in salmonid evolution. A gap appears in the fossil record after E. driftwoodensis until about seven million years ago, in the late Miocene. Several of these appear to be Oncorhynchus—the current genus for Pacific salmon. Consequently, the split between Oncorhynchus and Salmo must have occurred well before the Pliocene, suggestions have gone back as far as the early Miocene. Based on the most current evidence, salmonids diverged from the rest of teleost fish no later than 88 million years ago and this divergence was marked by a whole-genome duplication event in the ancestral salmonid, where the diploid ancestor became tetraploid. Extant salmonids all show evidence of partial tetraploidy, as show the genome has undergone selection to regain a diploid state. Work done in the trout has shown that the genome is still partially-tetraploid. This pattern of partial tetraploidy is thought to be reflected in the rest of extant salmonids, the first fossil species representing a true salmonid fish does not appear until the middle Eocene. This fossil is also younger than the proposed salmonid divergence from the rest of the teleost fishes. This means that the salmonids have a ghost lineage of approximately 33 million years and this more precise dating and examination of the salmonid whole-genome duplication event has allowed more speculation on the radiation of species within the group. Historically, the duplication event was thought to be the reason for the variation within Salmonidae
15.
Tributary
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A tributary or affluent is a stream or river that flows into a larger stream or main stem river or a lake. A tributary does not flow directly into a sea or ocean, tributaries and the main stem river drain the surrounding drainage basin of its surface water and groundwater, leading the water out into an ocean. A confluence, where two or more bodies of water together, usually refers to the joining of tributaries. The opposite to a tributary is a distributary, a river or stream that branches off from, distributaries are most often found in river deltas. Right tributary and left tributary are terms stating the orientation of the relative to the flow of the main stem river. These terms are defined from the perspective of looking downstream, where tributaries have the same name as the river into which they feed, they are called forks. These are typically designated by compass direction, for example, the American River receives flow from its North, Middle, and South forks. The Chicago Rivers North Branch has the East, West, and Middle Fork, the South Branch has its South Fork, forks are sometimes designated as right or left. Here, the handedness is from the point of view of an observer facing upstream, for instance, Steer Creek has a left tributary which is called Right Fork Steer Creek. Tributaries are sometimes listed starting with those nearest to the source of the river, the Strahler Stream Order examines the arrangement of tributaries in a hierarchy of first, second, third, and higher orders, with the first-order tributary being typically the least in size. For example, a second-order tributary would be the result of two or more first-order tributaries combining to form the second-order tributary, another method is to list tributaries from mouth to source, in the form of a tree structure, stored as a tree data structure
16.
Fish migration
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Many types of fish migrate on a regular basis, on time scales ranging from daily to annually or longer, and over distances ranging from a few metres to thousands of kilometres. Fish usually migrate to feed or to reproduce, but in cases the reasons are unclear. Migrations involves the fish moving from one part of a body to another on a regular basis. Marine forage fish often make large migrations between their spawning, feeding and nursery grounds, movements are associated with ocean currents and with the availability of food in different areas at different times of year. The migratory movements may partly be linked to the fact that the fish cannot identify their own offspring and moving in this way prevents cannibalism, some species have been described by the United Nations Convention on the Law of the Sea as highly migratory species. These are large fish that move in and out of the exclusive economic zones of different nations. Salmon and striped bass are well-known anadromous fish, and freshwater eels are catadromous fish that make large migrations, some fish such as tuna move to the north and south at different times of year following temperature gradients. The patterns of migration are of great interest to the fishing industry, movements of fish in fresh water also occur, often the fish swim upriver to spawn, and these traditional movements are increasingly being disrupted by the building of dams. As with various aspects of fish life, zoologists have developed empirical classifications for fish migrations. Two terms in particular have been in long-standing wide usage, Anadromous fish migrate from the sea up into water to spawn, examples are salmon. Catadromous fish migrate from fresh water down into the sea to spawn, in a 1949 journal article, George S. Myers coined the inclusive term diadromous to refer to all fishes that migrate between the sea and fresh water. Like the two well known terms, it was formed from classical Greek, diadromous proved a useful word, but terms proposed by Myers for other types of diadromous fishes did not catch on. These included amphidromous, potamodromous, and oceanodromous, although these classifications were originated for fishes, they are, in principle, applicable to any aquatic organism. Forage fish often make great migrations between their spawning, feeding and nursery grounds, schools of a particular stock usually travel in a triangle between these grounds. For example, one stock of herrings have their ground in southern Norway, their feeding ground in Iceland. Wide triangular journeys such as these may be important because forage fish, capelin are a forage fish of the smelt family found in the Atlantic and Arctic oceans. In summer, they graze on dense swarms of plankton at the edge of the ice shelf, larger capelin also eat krill and other crustaceans. The capelin move inshore in large schools to spawn and migrate in spring, the migration is affected by ocean currents
17.
Spawn (biology)
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Spawn is the eggs and sperm released or deposited into water by aquatic animals. As a verb, to refers to the process of releasing the eggs and sperm. Most aquatic animals, except for aquatic mammals and reptiles, reproduce through the process of spawning, spawn consists of the reproductive cells of many aquatic animals, some of which will become fertilized and produce offspring. The process of spawning typically involves females releasing ova into the water, often in large quantities, fungi, such as mushrooms, are also said to spawn a white, fibrous matter that forms the matrix from which they grow. Marine animals, and particularly fish, commonly reproduce by broadcast spawning. This is a method of reproduction where the female releases many unfertilised eggs into the water. At the same time, a male or many males release a lot of sperm into the water which fertilises some of these eggs, the eggs contain a drop of nutrient oil to sustain the embryo as it develops inside the egg case. The oil also provides buoyancy, so the float and drift with the current. The strategy for survival of broadcast spawning is to disperse the fertilised eggs, there the larvae develop as they consume their fat stores, and eventually hatch from the egg capsule into miniature versions of their parents. To survive, they must then become miniature predators themselves, feeding on plankton, fish eventually encounter others of their own kind, where they form aggregations and learn to school. Internally, the sexes of most marine animals can be determined by looking at the gonads, male lampreys, hagfish and salmon discharge their sperm into the body cavity where it is expelled through pores in the abdomen. Male sharks and rays can pass sperm along a duct into a seminal vesicle, externally, many marine animals, even when spawning, show little sexual dimorphism or little difference in colouration. Where species are dimorphic, such as sharks or guppies, the males often have penis-like intromittent organs in the form of a modified fin, a species is semelparous if its individuals spawn only once in their lifetime, and iteroparous if its individuals spawn more than once. The term semelparity comes from the Latin semel, once, and pario, to beget, while iteroparity comes from itero, to repeat, semelparity is sometimes called big bang reproduction, since the single reproductive event of semelparous organisms is usually large and fatal to the spawners. The classic example of an animal is the Pacific salmon, which lives for many years in the ocean before swimming to the freshwater stream of its birth, spawning. Other spawning animals which are semelparous include mayflies, squid, octopus, smelt, capelin, semelparity is often associated with r-strategists. However, most fish and other spawning animals are iteroparous, when the internal ovaries or egg masses of fish and certain marine animals are ripe for spawning they are called roe. Roe from certain species, such as shrimp, scallop, crab, caviar is a name for the processed, salted roe of non-fertilized sturgeon
18.
Introduced species
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Non-native species can have various effects on the local ecosystem. Introduced species that become established and spread beyond the place of introduction are called invasive species, some have a negative effect on a local ecosystem. Some introduced species may have no effect or only minor impact. Some species have been introduced intentionally to combat pests and they are called biocontrols and may be regarded as beneficial as an alternative to pesticides in agriculture for example. In some instances the potential for being beneficial or detrimental in the long run remains unknown, a list of some introduced species is given in a separate article. The effects of introduced species on natural environments have gained much scrutiny from scientists, governments, the terminology associated with introduced species is now in flux for various reasons. The term invasive refers only to those species that become established, for practical purposes, this term is applied only to invasive species that cause damage. With respect to plants, these latter are in this case defined as either ornamental or cultivated plants, also called an exotic or non-native species. Such species might be termed naturalized, established, wild non-native species, if they further spread beyond the place of introduction they are called invasive. The transition from introduction, to establishment and to invasion has been described in the context of plants, introduced species are essentially non-native species. Invasive species are introduced species that spreadwidely or quickly and cause harm, be that to the environment, human health. There have been calls from scientists to consider a species only in terms of their spread. According to a definition, an invasive species is one that has been introduced and become a pest in its new location. The term is used to both a sense of urgency and actual or potential harm. Executive Order 13112 defines invasive species as a species whose introduction does or is likely to cause economic or environmental harm or harm to human health. The biological definition of species, on the other hand, makes no reference to the harm they may cause. From a regulatory perspective, it is neither desirable nor practical to list as undesirable or outright ban all non-native species, regulations require a definitional distinction between non-natives that are deemed especially onerous and all others. Introduced pest species that are listed as invasive, best fit the definition of an invasive species
19.
Great Lakes
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Consisting of Lakes Superior, Michigan, Huron, Erie, and Ontario, they form the largest group of freshwater lakes on Earth, containing 21% of the worlds surface fresh water by volume. The total surface is 94,250 square miles, and the volume is 5,439 cubic miles. Due to their sea-like characteristics the five Great Lakes have also long been referred to as inland seas, Lake Superior is the second largest lake in the world by area, and Lake Michigan is the largest lake that is entirely within one country. The southern half of the Great Lakes is bordered by the Great Lakes Megalopolis, the lakes have been a major highway for transportation, migration and trade, and they are home to a large number of aquatic species. Many invasive species have been introduced due to trade, and some threaten the regions biodiversity, though the five lakes reside in separate basins, they form a single, naturally interconnected body of fresh water, within the Great Lakes Basin. The lakes form a chain connecting the interior of North America to the Atlantic Ocean. From the interior to the outlet at the Saint Lawrence River, water flows from Superior to Huron and Michigan, southward to Erie, the lakes drain a large watershed via many rivers, and are studded with approximately 35,000 islands. There are also several smaller lakes, often called inland lakes. The surface area of the five primary lakes combined is roughly equal to the size of the United Kingdom, while the area of the entire basin is about the size of the UK. Lake Michigan is the one of the Great Lakes that is located entirely within the United States. The lakes are divided among the jurisdictions of the Canadian province of Ontario and the U. S. states of Minnesota, Wisconsin, Michigan, Illinois, Indiana, Ohio, Pennsylvania, and New York. Both Ontario and Michigan include in their boundaries portions of four of the lakes, Ontario does not border Lake Michigan, New York and Wisconsins jurisdictions extend into two lakes, and the remaining states into one of the lakes. This designation, however, is not universal and those living on the shore of Lake Superior often refer to all the other lakes as the lower lakes, because they are farther south. This corresponds to thinking of Lakes Erie and Ontario as down south, vessels sailing north on Lake Michigan are considered upbound even though they are sailing toward its effluent current. The Chicago River and Calumet River systems connect the Great Lakes Basin to the Mississippi River System through man-made alterations, the St. Marys River, including the Soo Locks, connects Lake Superior to Lake Huron. The Straits of Mackinac connect Lake Michigan to Lake Huron, the St. Clair River connects Lake Huron to Lake St. Clair. The Detroit River connects Lake St. Clair to Lake Erie, the Niagara River, including Niagara Falls, connects Lake Erie to Lake Ontario. The Welland Canal, bypassing the Falls, connects Lake Erie to Lake Ontario, the Saint Lawrence River connects Lake Ontario to the Gulf of St. Lawrence, which connects to the Atlantic Ocean
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Subspecies
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In biological classification, subspecies is either a taxonomic rank subordinate to species, or a taxonomic unit in that rank. A subspecies cannot be recognized independently, a species will either be recognized as having no subspecies at all or at least two, in zoology, under the International Code of Zoological Nomenclature, the subspecies is the only taxonomic rank below that of species that can receive a name. In botany and mycology, under the International Code of Nomenclature for algae, fungi, in bacteriology and virology, under standard bacterial nomenclature and virus nomenclature, there are recommendations but not strict requirements for recognizing other important infraspecific ranks. A taxonomist decides whether to recognize a subspecies or not, the differences between subspecies are usually less distinct than the differences between species. In zoology, the International Code of Zoological Nomenclature accepts only one rank below that of species, other groupings, infrasubspecific entities do not have names regulated by the ICZN. Such forms have no official ICZN status, though they may be useful in describing altitudinal or geographical clines, pet breeds, transgenic animals, etc. While the scientific name of a species is a binomen, the name of a subspecies is a trinomen - a binomen followed by a subspecific name. A tigers binomen is Panthera tigris, so for a Sumatran tiger the trinomen is, for example, names published before 1992 in the rank of variety are taken to be names of subspecies. In botany, subspecies is one of many ranks below that of species, such as variety, subvariety, form, the subspecific name is preceded by subsp. or ssp. as Schoenoplectus californicus ssp. tatora. A botanical name consists of at most three parts, an infraspecific name includes the species binomial, and one infraspecific epithet, such as subspecies or variety. For example, Motacilla alba alba is the subspecies of the white wagtail. The subspecies name that repeats the name is referred to in botanical nomenclature as the subspecies autonym. When zoologists disagree over whether a population is a subspecies or a full species. A subspecies is a rank below species – the only recognized rank in the zoological code. Botanists and mycologists have the choice of ranks lower than subspecies, such as variety or form, in biological terms, rather than in relation to nomenclature, a polytypic species has two or more subspecies, races, or more generally speaking, populations that need a separate description. These are separate groups that are distinct from one another and do not generally interbreed. These subspecies, races, or populations, can be named as subspecies by zoologists, a monotypic species has no distinct population or races, or rather one race comprising the whole species. A taxonomist would not name a subspecies within such a species, monotypic species can occur in several ways, All members of the species are very similar and cannot be sensibly divided into biologically significant subcategories
21.
Habitat
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A habitat is an ecological or environmental area that is inhabited by a particular species of animal, plant, or other type of organism. The term typically refers to the zone in which the organism lives and it is the natural environment in which an organism lives, or the physical environment that surrounds a species population. Every organism has certain habitat needs for the conditions in which it will thrive, habitat types include polar, temperate, subtropical and tropical. The terrestrial vegetation type may be forest, steppe, grassland, the word habitat has been in use since about 1755 and derives from the Latin third-person singular present indicative of habitāre, to inhabit, from habēre, to have or to hold. Habitat can be defined as the environment of an organism. It is similar in meaning to a biotope, an area of environmental conditions associated with a particular community of plants. Generally speaking, animal communities are reliant on specific types of plant communities, some plants and animals are generalists, and their habitat requirements are met in a wide range of locations. The small white butterfly for example is found on all the continents of the world apart from Antarctica and its larvae feed on a wide range of Brassicas and various other plant species, and it thrives in any open location with diverse plant associations. Disturbance is important in the creation of biodiverse habitats, in the absence of disturbance, a climax vegetation cover develops that prevents the establishment of other species. Lightning strikes and toppled trees in tropical forests allow species richness to be maintained as pioneering species move in to fill the gaps created. Similarly coastal habitats can become dominated by kelp until the seabed is disturbed by a storm, another cause of disturbance is when an area may be overwhelmed by an invasive introduced species which is not kept under control by natural enemies in its new habitat. Terrestrial habitat types include forests, grasslands, wetlands and deserts, within these broad biomes are more specific habitats with varying climate types, temperature regimes, soils, altitudes and vegetation types. Many of these habitats grade into each other and each one has its own communities of plants. A habitat may suit a particular species well, but its presence or absence at any particular location depends to some extent on chance, on its dispersal abilities, freshwater habitats include rivers, streams, lakes, ponds, marshes and bogs. Although some organisms are found across most of these habitats, the majority have more specific requirements, similarly, aquatic plants can be floating, semi-submerged, submerged or grow in permanently or temporarily saturated soils besides bodies of water. Marine habitats include brackish water, estuaries, bays, the sea, the intertidal zone. Further variations include rock pools, sand banks, mudflats, brackish lagoons, sandy and pebbly beaches, the benthic zone or seabed provides a home for both static organisms, anchored to the substrate, and for a large range of organisms crawling on or burrowing into the surface. A desert is not the kind of habitat that favours the presence of amphibians, with their requirement for water to keep their skins moist, nevertheless, some frogs live in deserts, creating moist habitats underground and hibernating while conditions are adverse
22.
Lateral line
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The lateral line is a system of sense organs found in aquatic vertebrates, used to detect movement, vibration, and pressure gradients in the surrounding water. Lateral lines serve an important role in schooling behavior, predation, for example, fish can use their lateral line system to follow the vortices produced by fleeing prey. Lateral lines are visible as faint lines of pores running lengthwise down each side. Most amphibian larvae and some fully aquatic adult amphibians possess mechanosensitive systems comparable to the lateral line and this plays an essential role in orientation, predatory behavior, defense, and social schooling. The lateral line system is necessary to detect vibrations made by prey, Fish are able to detect movement, produced either by prey or a vibrating metal sphere, and orient themselves toward the source before proceeding to make a predatory strike at it. This behavior persists even in blinded fish, but is diminished when lateral line function was inhibited by CoCl2 application. Cobalt chloride treatment results in the release of cobalt ions, disrupting ionic transport and these behaviors are dependent specifically on mechanoreceptors located within the canals of the lateral line. The role mechanoreception plays in schooling behavior was demonstrated in a 1976 study, a school of Pollachius virens was established in a tank and individual fish were removed and subjected to different procedures before their ability to rejoin the school was observed. Fish that were experimentally blinded were able to reintegrate into the school, therefore, reliance on functional mechanoreception, not vision, is essential for schooling behavior. A study in 2014 suggests that the lateral line system plays an important role in the behavior of Mexican blind cave fish, the major unit of functionality of the lateral line is the neuromast. The neuromast is an organ which allows the sensing of mechanical changes in water. There are two varieties of neuromasts located in animals, canal neuromasts and superficial or freestanding neuromasts. Superficial neuromasts are located externally on the surface of the body, while canal neuromasts are located along the lines in subdermal. Each neuromast consists of hair cells whose tips are covered by a flexible. Hair cells typically possess both glutamatergic afferent connections and cholinergic efferent connections, the receptive hair cells are modified epithelial cells and typically possess bundles of 40-50 microvilli hairs which function as the mechanoreceptors. These bundles are organized in rough staircases of hairs of increasing length order and this use of mechanosensitive hairs is homologous to the functioning of hair cells in the auditory and vestibular systems, indicating a close link between these systems. Hair cells utilize a system of transduction that uses rate coding in order to transmit the directionality of a stimulus, hair cells of the lateral line system produce a constant, tonic rate of firing. As mechanical motion is transmitted through water to the neuromast, the bends and is displaced
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Gill
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A gill is a respiratory organ found in many aquatic organisms that extracts dissolved oxygen from water and excretes carbon dioxide. The gills of species, such as hermit crabs, have adapted to allow respiration on land provided they are kept moist. The microscopic structure of a gill presents a surface area to the external environment. Many microscopic aquatic animals, and some larger but inactive ones, can absorb oxygen through the entire surface of their bodies. However, more complex or more active aquatic organisms usually require a gill or gills, Gills usually consist of thin filaments of tissue, branches, or slender, tufted processes that have a highly folded surface to increase surface area. A high surface area is crucial to the gas exchange of aquatic organisms, a cubic meter of air contains about 250 grams of oxygen at STP. The concentration of oxygen in water is lower than in air, in fresh water, the dissolved oxygen content is approximately 8 cm3/L compared to that of air which is 210 cm3/L. Water is 777 times more dense than air and is 100 times more viscous, oxygen has a diffusion rate in air 10,000 times greater than in water. The use of sac-like lungs to remove oxygen from water would not be efficient enough to sustain life, rather than using lungs, aseous exchange takes place across the surface of highly vascularised gills over which a one-way current of water is kept flowing by a specialised pumping mechanism. The density of the water prevents the gills from collapsing and lying on top of each other, with the exception of some aquatic insects, the filaments and lamellae contain blood or coelomic fluid, from which gases are exchanged through the thin walls. The blood carries oxygen to other parts of the body, carbon dioxide passes from the blood through the thin gill tissue into the water. Gills or gill-like organs, located in different parts of the body, are found in groups of aquatic animals, including mollusks, crustaceans, insects, fish. The gills of vertebrates typically develop in the walls of the pharynx, most species employ a countercurrent exchange system to enhance the diffusion of substances in and out of the gill, with blood and water flowing in opposite directions to each other. The gills are composed of filaments, the gill lamellae. When a fish breathes, it draws in a mouthful of water at regular intervals, then it draws the sides of its throat together, forcing the water through the gill openings, so it passes over the gills to the outside. Fish gill slits may be the ancestors of the tonsils, thymus glands. Adjacent slits are separated by a gill arch from which projects a cartilaginous gill ray. This gill ray is the support for the sheet-like interbranchial septum, the base of the arch may also support gill rakers, projections into the pharyngeal cavity that help to prevent large pieces of debris from damaging the delicate gills
24.
Fish hatchery
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A fish hatchery is a place for artificial breeding, hatching and rearing through the early life stages of animals, finfish and shellfish in particular. Hatcheries produce larval and juvenile fish primarily to support the industry where they are transferred to on-growing systems i. e. fish farms to reach harvest size. Some species that are raised in hatcheries include Pacific oysters, shrimp, Indian prawns, salmon, tilapia. Additional hatchery production for domestic uses, which is particularly prevalent in South-East Asia or for conservation programmes, has also yet to be quantified. There is much interest in supplementing exploited stocks of fish by releasing juveniles that may be caught and reared in nurseries before transplanting. Culture of finfish larvae has been utilised extensively in the United States in stock enhancement efforts to replenish natural populations, the U. S. Fish and Wildlife Service have established a National Fish Hatchery System to support the conservation of native fish species. Hatcheries produce larval and juvenile fish and shellfish for transferral to aquaculture facilities where they are ‘on-grown’ to reach harvest size, Hatchery production confers three main benefits to the industry,1. Out of season production Consistent supply of fish from aquaculture facilities is an important market requirement, broodstock conditioning can extend the natural spawning season and thus the supply of juveniles to farms. Supply can be guaranteed by sourcing from hatcheries in the opposite hemisphere i. e. with opposite seasons.2. Genetic improvement Genetic modification is conducted in some hatcheries to improve the quality, Genetic improvement can be mediated by selective breeding, via hybridization, or other genetic manipulation techniques.3. Reduce dependence on wild-caught juveniles In 2008 aquaculture accounted for 46% of total food fish supply, broodstock conditioning is the process of bringing adults into spawning condition by promoting the development of gonads. Broodstock conditioning can also extend spawning beyond natural spawning periods, or for production of species reared outside their geographic range with different environmental conditions. Some hatcheries collect wild adults and then bring them in for conditioning whilst others maintain a permanent breeding stock, conditioning is achieved by holding broodstock in flow-through tanks at optimal conditions for light, temperature, salinity, flow rate and food availability. Egg quality is determined by the nutritional condition of the mother. High levels of reserves in particular are required to improve larval survival rates. Some of the common methods are, Manual stripping, For shellfish, gonads are generally removed. Alternatively, if environmental cues that stimulate natural spawning are known, prior to fertilisation, eggs can be gently washed to remove wastes and bacteria that may contaminate cultures. Promoting cross-fertilisation between a number of individuals is necessary to retain genetic diversity in hatchery produced stock
25.
Southern Europe
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Southern Europe is the southern region of the European continent. Most definitions of Southern Europe, also known as Mediterranean Europe, include the countries of the Iberian peninsula, different methods can be used to define Southern Europe, including its political, economic, and cultural attributes. Southern Europe can also be defined by its natural features — its geography, climate, geographically, Southern Europe is the southern half of the landmass of Europe. This definition is relative, with no clear limits and those areas of Mediterranean climate present similar vegetations and landscapes throughout, including dry hills, small plains, pine forests and olive trees. Cooler climates can be found in parts of Southern European countries, for example within the mountain ranges of Spain. Additionally, the north coast of Spain experiences a wetter Atlantic climate, Southern Europes flora is that of the Mediterranean Region, one of the phytochoria recognized by Armen Takhtajan. The period known as classical antiquity began with the rise of the city-states of Ancient Greece, Greek influence reached its zenith under the expansive empire of Alexander the Great, spreading throughout Asia. The Roman Empire came to dominate the entire Mediterranean basin in a vast empire based on Roman law and it promoted trade, tolerance, and Greek culture. By 300 AD the Roman Empire was divided into the Western Roman Empire based in Rome, during the Middle Ages, the Eastern Roman Empire survived, though modern historians refer to this state as the Byzantine Empire. In Western Europe, Germanic peoples moved into positions of power in the remnants of the former Western Roman Empire and established kingdoms, the period known as the Crusades, a series of religiously motivated military expeditions originally intended to bring the Levant back into Christian rule, began. Several Crusader states were founded in the eastern Mediterranean, the Crusaders would have a profound impact on many parts of Europe. Their Sack of Constantinople in 1204 brought an end to the Byzantine Empire. Though it would later be re-established, it would never recover its former glory, the Crusaders would establish trade routes that would develop into the Silk Road and open the way for the merchant republics of Genoa and Venice to become major economic powers. The Reconquista, a movement, worked to reconquer Iberia for Christendom. The Late Middle Ages represented a period of upheaval in Europe, the epidemic known as the Black Death and an associated famine caused demographic catastrophe in Europe as the population plummeted. Dynastic struggles and wars of conquest kept many of the states of Europe at war for much of the period, in the Balkans, the Ottoman Empire, a Turkish state originating in Anatolia, encroached steadily on former Byzantine lands, culminating in the Fall of Constantinople in 1453. An unprecedented series of wars and political revolutions took place around Europe. Observers at the time, and many historians since, have argued that wars caused the revolutions, galileo Galilei, invented the telescope and the thermometer which allowed him to observe and describe the solar system
26.
Whirling disease
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Myxobolus cerebralis is a myxosporean parasite of salmonids that causes whirling disease in farmed salmon and trout and also in wild fish populations. It was first described in rainbow trout in Germany a century ago, but its range has spread and it has appeared in most of Europe, in the 1980s, M. cerebralis was found to require a tubificid oligochaete to complete its life cycle. The parasite infects its hosts with its cells after piercing them with polar filaments ejected from nematocyst-like capsules, Whirling disease afflicts juvenile fish and causes skeletal deformation and neurological damage. Fish whirl forward in an awkward, corkscrew-like pattern instead of swimming normally, find feeding difficult, the mortality rate is high for fingerlings, up to 90% of infected populations, and those that do survive are deformed by the parasites residing in their cartilage and bone. They act as a reservoir for the parasite, which is released into water following the fishs death, M. cerebralis is one of the most economically important myxozoans in fish, as well as one of the most pathogenic. It was the first myxosporean whose pathology and symptoms were described scientifically, the parasite is not transmissible to humans. The taxonomy and naming of both M. cerebralis, and of myxozoans in general, have complicated histories and it was originally thought to infect fish brains and nervous systems, though it soon was found to primarily infect cartilage and skeletal tissue. Attempts to change the name to Myxobolus chondrophagus, which would more accurately describe the organism, similarly, other actinosporeans were folded into the life cycles of various myxosporeans. Today, the myxozoans, previously thought to be protozoans, are considered animals by most scientists. Recent molecular studies suggest they are related to Bilateria or Cnidaria, M. cerebralis has many diverse stages ranging from single cells to relatively large spores, not all of which have been studied in detail. The stages that infect fish, called triactinomyxon spores, are made of a style that is about 150 micrometers long. A sporoplasm packet at the end of the style contains 64 germ cells surrounded by a cellular envelope, there are also three polar capsules, each of which contains a coiled polar filament between 170 and 180 µm long. Polar filaments in both stage and in the myxospore stage rapidly shoot into the body of the host. Upon contact with fish hosts and firing of the polar capsules, firstly, this sporoplasm undergoes mitosis to produce more amoeboid cells, which migrate into deeper tissue layers, to reach the cerebral cartilage. Myxospores, which develop from sporogonic cell stages inside fish hosts, are lenticular and they have a diameter of about 10 micrometers and are made of six cells. Two of these cells form polar capsules, two merge to form a binucleate sporoplasm, and two form protective valves, myxospores are infective to oligochaetes, and are found among the remains of digested fish cartilage. They are often difficult to distinguish from related species because of morphological similarities across genera, though M. cerebralis is the only myxosporean ever found in salmonid cartilage, other visually similar species may be present in the skin, nervous system, or muscle. Myxobolus cerebralis has a two-host life cycle involving a salmonid fish, so far, the only worm known to be susceptible to M. cerebralis infection is Tubifex tubifex, though what scientists currently call T. tubifex may in fact be more than one species
27.
Hybrid (biology)
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In biology, a hybrid, also known as a cross breed, is the result of combining, through sexual reproduction, the qualities of two animals or plants of different breeds, varieties, species or genera. Hybrids are not always intermediate between their parents, but can show heterosis or hybrid vigour, often growing larger or taller than either parent, some act before fertilization, others after it. A few animal species and many plant species, however, are the result of hybrid speciation, doubling the number of chromosomes to create polyploids is important in hybrid speciation, because homoploid hybrids are rarely fertile, the polyploid hybrids are allopolyploids. Human impact on the environment has greatly increased the mixing of species, with introduced species worldwide, the genetic mixing may threaten many species with extinction, while genetic erosion in crop plants may be damaging the gene pools of many species for future breeding. Many commercially useful fruits, flowers, garden herbs and trees have been produced by hybridization, one flower, Oenothera lamarckiana, was central to early genetics research into polyploidy. The term hybrid is derived from Latin hybrida, used for such as of a tame sow and a wild boar, or the child of a freeman. The term came into use in English in the 19th century. Conspicuous hybrids are named with portmanteau words, starting in the 1920s with the breeding of tiger-lion hybrids. The cross between two different homozygous lines produces an F1 hybrid that is heterozygous, having two alleles, one contributed by each parent and typically one is dominant and the other recessive. Typically, the F1 generation is also phenotypically homogeneous, producing offspring that are all similar to each other, double cross hybrids result from the cross between two different F1 hybrids. Three-way cross hybrids result from the cross between an F1 hybrid and an inbred line, triple cross hybrids result from the crossing of two different three-way cross hybrids. Top cross hybrids result from the crossing of a top quality or pure-bred male, population hybrids result from the crossing of plants or animals in a population with those of another population. These include interspecific hybrids or crosses between different breeds. e, from the point of view of genetics, different kinds of hybrid can be distinguished as follows, A genetic hybrid carries two different alleles of the same gene. A structural hybrid results from the fusion of gametes that have differing structure in at least one chromosome, a numerical hybrid results from the fusion of gametes having different haploid numbers of chromosomes. A permanent hybrid results when only the heterozygous genotype occurs, as in Oenothera lamarckiana, from the point of view of taxonomy, hybrids differ according to their parentage, Hybrids between different subspecies are called intra-specific hybrids. Offspring resulting from interspecies mating, are called interspecific hybrids, these result in hybrid speciation. Intergeneric hybrids result from matings between different genera, such as sheep and goats. Interfamilial hybrids such as chickens and guineafowl or pheasants are reliably described but extremely rare
28.
Recreational fishing
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Recreational fishing, also called sport fishing, is fishing for pleasure or competition. It can be contrasted with fishing, which is fishing for profit, or subsistence fishing. The most common form of fishing is done with a rod, reel, line, hooks. Other devices, commonly referred to as terminal tackle, are used to affect or complement the presentation of the bait to the targeted fish. Some examples of terminal tackle include weights, floats, and swivels, lures are frequently used in place of bait. Some hobbyists make handmade tackle themselves, including plastic lures and artificial flies, the practice of catching or attempting to catch fish with a hook is known as angling. Big-game fishing is conducted from boats to catch large species such as tuna, sharks. Noodling and trout tickling are also recreational activities, the early evolution of fishing as recreation is not clear. For example, there is evidence for fly fishing in Japan as early as the ninth century BCE. But for the early Japanese and Macedonians, fly fishing was likely to have been a means of survival and it is possible that antecedents of recreational fly fishing arrived in England with the Norman conquest of 1066. Although the point in history where fishing could first be said to be recreational is not clear, the earliest English essay on recreational fishing was published in 1496, shortly after the invention of the printing press. The authorship of this was attributed to Dame Juliana Berners, the prioress of the Benedictine Sopwell Nunnery, the essay was titled Treatyse of Fysshynge wyth an Angle, and was published in the second Boke of Saint Albans, a treatise on hawking, hunting, and heraldry. During the 16th century the work was read, and was reprinted many times. Treatyse includes detailed information on fishing waters, the construction of rods and lines, and it also includes modern concerns about conservation and angler etiquette. Another Civil War veteran to enthusiastically take up fishing, was Richard Franck and he was the first to describe salmon fishing in Scotland, and both in that and trout-fishing with artificial fly he was a practical angler. He was the first angler to name the burbot, and commended the salmon of the River Thames, Compleat Angler was written by Izaak Walton in 1653 and described the fishing in the Derbyshire Wye. It was a celebration of the art and spirit of fishing in prose and verse,6 verses were quoted from John Dennyss earlier work, a second part to the book was added by Waltons friend Charles Cotton. More than 300 editions of The Compleat Angler have been published, the pastoral discourse was enriched with country fishing folklore, songs and poems, recipes and anecdotes, moral meditations, and quotes from classic literature
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Wyoming
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Wyoming /waɪˈoʊmɪŋ/ is a state in the mountain region of the western United States. The state is the tenth largest by area, the least populous, Wyoming is bordered on the north by Montana, on the east by South Dakota and Nebraska, on the south by Colorado, on the southwest by Utah, and on the west by Idaho. Cheyenne is the capital and the most populous city in Wyoming, the state population was estimated at 586,107 in 2015, which is less than the population of 31 of the largest U. S. cities. The Crow, Arapaho, Lakota, and Shoshone were some of the inhabitants of the region. Southwestern Wyoming was included in the Spanish Empire and then Mexican territory until it was ceded to the United States in 1848 at the end of the Mexican–American War. The region acquired the name Wyoming when a bill was introduced to Congress in 1865 to provide a government for the territory of Wyoming. The territory was named after the Wyoming Valley in Pennsylvania, with the name ultimately being derived from the Munsee word xwé, wamənk, the mineral extraction industry—especially coal, oil, natural gas, and trona—along with the travel and tourism sector are the main drivers behind Wyomings economy. Agriculture has historically been an important component of the economy with the main commodities being livestock, hay, sugar beets, grain. The climate is generally semi-arid and continental, being drier and windier in comparison to the rest of the United States, except for the 1964 election, Wyoming has been a politically conservative state since the 1950s, with the Republican party winning every presidential election. Wyoming is one of three states to have borders along only straight latitudinal and longitudinal lines, rather than being defined by natural landmarks. Wyoming is bordered on the north by Montana, on the east by South Dakota and Nebraska, on the south by Colorado, on the southwest by Utah, and on the west by Idaho. It is the tenth largest state in the United States in total area, from the north border to the south border it is 276 miles, and from the east to the west border is 365 miles at its south end and 342 miles at the north end. The Great Plains meet the Rocky Mountains in Wyoming, the state is a great plateau broken by many mountain ranges. Surface elevations range from the summit of Gannett Peak in the Wind River Mountain Range, at 13,804 feet, to the Belle Fourche River valley in the states northeast corner, at 3,125 feet. In the northwest are the Absaroka, Owl Creek, Gros Ventre, Wind River, in the north central are the Big Horn Mountains, in the northeast, the Black Hills, and in the southern region the Laramie, Snowy and Sierra Madre ranges. The Snowy Range in the central part of the state is an extension of the Colorado Rockies in both geology and appearance. The Wind River Range in the west central part of the state is remote and includes more than 40 mountain peaks in excess of 13,000 ft tall in addition to Gannett Peak, the highest peak in the state. The Big Horn Mountains in the central portion are somewhat isolated from the bulk of the Rocky Mountains
30.
Firehole River
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The Firehole River is located in northwestern Wyoming, and is one of the two major tributaries of the Madison River. It flows north approximately 21 miles from its source in Madison Lake on the Continental Divide to join the Gibbon River at Madison Junction in Yellowstone National Park and it is part of the Missouri River system. The Firehole River flows through several significant geyser basins in the park to include the Upper Geyser Basin, the river was named by early trappers for the steam that makes it appear to be smoking as if on fire. The Firehole contains three of Yellowstones major waterfalls, Kepler Cascades south of Old Faithful, Firehole Falls and the Cascades of the Firehole in Firehole Canyon, the river is surrounded by geothermal features which empty water into it. One effect of the input of water is to increase the temperature of the water. Temperatures in the river have been measured as high as 30 °C, water entering the river from geothermal features contains dissolved chemicals and minerals. Levels of boron and arsenic have been found to be above the limits for protection of aquatic organisms. Despite these levels, brown and rainbow trout live and spawn in the river, Firehole River has been a fishing mecca since the late 19th century and is known today for its excellent fly fishing. The river contains an invasive species, the New Zealand mud snail, significant tributaries of the Firehole are the Little Firehole River, Fairy Creek, Iron Spring Creek, Sentinel Creek and Nez Perce Creek. All these tributaries bring cool waters to the Firehole and provide refuge for trout in the mainstem during mid-summer high temperatures caused by geothermal activity, the Firehole River is a famous and storied destination for serious fly fishermen. When it was discovered in the 1830s by American explorers, the Firehole was barren of trout above what is now called Firehole Falls. Brook trout were first introduced to the upper Firehole in 1889, while brown trout, rainbow trout were not introduced until 1923. Mountain whitefish are native to the Firehole below Firehole Falls, by the late 19th century, the Firehole and Yellowstone National Park in general was a popular destination for fishermen. In 1955 all stocking programs in the park were discontinued and todays Firehole trout are completely wild populations, in 1968, based on increasing pressure on the Firehole, the Gibbon and Madison rivers, the National Park Service designated these waters as Fly Fishing Only. Angling in Yellowstone National Park Fishes of Yellowstone National Park Fly fishing Tributaries of the Missouri River Back, the Waters of the Yellowstone with Rod and Fly. The Living River-A Fishermans Intimate Profile of the Madison River Watershed—Its History, Ecology, Lore, garden City, NJ, Nick Lyons Books. The Yellowstone Fly-Fishing Guide-A authoritative guide to the waters of Yellowstone National Park, clinton, NJ, New Win Publishing Inc. Schullery, Paul, Varley, John D. Yellowstone Fishes—Ecology, History, and Angling in the Park
31.
Threatened species
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Threatened species are any species which are vulnerable to endangerment in the near future. Species that are threatened are sometimes characterised by the population dynamics measure of critical depensation and this quantitative metric is one method of evaluating the degree of endangerment. Species which have not been evaluated, or do not have sufficient data also are not considered threatened by the IUCN, Threatened species are also referred to as a red-listed species, as they are listed in the IUCN Red List of Threatened Species. Subspecies, populations and stocks may also be classified as threatened and it is the less protected of the two protected categories. The Bay checkerspot butterfly is an example of a threatened subspecies protected under the ESA, for example, the trumpeter swan is threatened in the state of Minnesota, while large populations still remain in Canada and Alaska. The EPBC Act also recognises and protects threatened ecosystems such as plant communities, some species, such as Lewins rail, are not listed as threatened species under the EPBC Act, but they may be recognised as threatened by individual states or territories. Biodiversity Action Plan IUCN Red List Illegal logging Rare species Red and blue-listed Slash-and-burn Threatened fauna of Australia Sharrock, S. and Jones, conserving Europes threatened plants Botanic Gardens Conservation International
32.
Endangered species
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An endangered species is a species which has been categorized as likely to become extinct. In 2012, the IUCN Red List featured 3079 animal and 2655 plant species as endangered worldwide, the figures for 1998 were, respectively,1102 and 1197. Many nations have laws that protect conservation-reliant species, for example, population numbers, trends and species conservation status can be found in the lists of organisms by population. The conservation status of a species indicates the likelihood that it will become extinct, the IUCN Red List of Threatened Species is the best-known worldwide conservation status listing and ranking system. Over 40% of the species are estimated to be at risk of extinction. Internationally,199 countries have signed an accord to create Biodiversity Action Plans that will protect endangered, in the United States, such plans are usually called Species Recovery Plans. Those species of Near Threatened and Least Concern status have been assessed and found to have relatively robust and healthy populations, though these may be in decline. The IUCN categories, with examples of animals classified by them, include, Extinct Extinct in the wild Captive individuals survive, critically endangered Faces an extremely high risk of extinction in the immediate future. Endangered Faces a high risk of extinction in the near future, vulnerable Faces a high risk of endangerment in the medium term. Near-threatened May be considered threatened in the near future, Least concern No immediate threat to species survival. A population size reduction of ≥ 50%, projected or suspected to be met within the next 10 years or three generations, whichever is the longer, based on any of to under A1. E) Quantitative analysis showing the probability of extinction in the wild is at least 20% within 20 years or five generations, there is data from the United States that shows a correlation between human populations and threatened and endangered species. Under the Endangered Species Act in the United States, species may be listed as endangered or threatened, the Salt Creek tiger beetle is an example of an endangered subspecies protected under the ESA. Some endangered species laws are controversial, also lobbying from hunters and various industries like the petroleum industry, construction industry, and logging, has been an obstacle in establishing endangered species laws. The Bush administration lifted a policy that required federal officials to consult an expert before taking actions that could damage endangered species. Under the Obama administration, this policy has been reinstated, being listed as an endangered species can have negative effect since it could make a species more desirable for collectors and poachers. This effect is potentially reducible, such as in China where commercially farmed turtles may be reducing some of the pressure to poach endangered species. Another problem with the species is its effect of inciting the use of the shoot, shovel
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Endangered Species Act
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The U. S. Supreme Court found that the plain intent of Congress in enacting the ESA was to halt and reverse the trend toward species extinction, whatever the cost. The Act is administered by two agencies, the United States Fish and Wildlife Service and the National Oceanic and Atmospheric Administration. Experimental, nonessential populations of endangered species are treated as threatened species on land, for consultation purposes. The near-extinction of the bison and the disappearance of the passenger pigeon helped drive the call for wildlife conservation starting in the 1900s, the public was introduced to a new concept, extinction. Market hunting for the trade and for the table was one aspect of the problem. The early naturalists also killed birds and other wildlife for study, personal curio collections, while habitat losses continued as communities and farmland grew, the widespread use of pesticides and the introduction of non-native species also affected wildlife. One species in particular received widespread attention—the whooping crane, the species historical range extended from central Canada South to Mexico, and from Utah to the Atlantic coast. It would be eight years before the first national law regulating wildlife commerce was signed. The whooping crane population by 1941 was estimated at about only 16 birds still in the wild, the Lacey Act of 1900 was the first federal law that regulated commercial animal markets. It prohibited interstate commerce of animals killed in violation of state game laws, other legislation followed, including the Migratory Bird Conservation Act of 1929, a 1937 treaty prohibiting the hunting of right and gray whales, and the Bald Eagle Protection Act of 1940. These later laws had a low cost to society–the species were relatively little opposition was raised. Whereas the Lacey Act dealt with game management and market commerce species. The predecessor of the ESA was the Endangered Species Preservation Act of 1966, passed by Congress, this act permitted the listing of native U. S. animal species as endangered and for limited protections upon those animals. It also directed federal agencies to preserve habitat on their lands. The Act also consolidated and even expanded authority for the Secretary of the Interior to manage, other public agencies were encouraged, but not required, to protect species. The act did not address the commerce in endangered species and parts, in March,1967 the first list of endangered species was issued under the act. It included 14 mammals,36 birds,6 reptiles and amphibians and 22 fish and it included only vertebrates because the Department of Interiors definition of fish and wildlife was limited to vertebrates. However, with time, researchers noticed that the animals on the species list still were not getting enough protection
34.
Washington (state)
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It was admitted to the Union as the 42nd state in 1889. Washington is sometimes referred to as Washington State or the State of Washington to distinguish it from Washington, Washington is the 18th largest state with an area of 71,362 square miles, and the 13th most populous state with over 7 million people. Washington is the second most populous state on the West Coast and in the Western United States, Mount Rainier, an active stratovolcano, is the states highest elevation at almost 14,411 feet and is the most topographically prominent mountain in the contiguous United States. Washington is a leading lumber producer and its rugged surface is rich in stands of Douglas fir, hemlock, ponderosa pine, white pine, spruce, larch, and cedar. Manufacturing industries in Washington include aircraft and missiles, shipbuilding and other equipment, lumber, food processing, metals and metal products, chemicals. Washington has over 1,000 dams, including the Grand Coulee Dam, built for a variety of purposes including irrigation, power, flood control, Washington was named after President George Washington by an act of the United States Congress during the creation of Washington Territory in 1853. Washington is the only U. S. state named after a president, confusion over the state of Washington and the city of Washington, D. C. led to renaming proposals during the statehood process for Washington in 1889, which failed to garner support. Washington, D. C. s own statehood movement in the 21st century includes a proposal to use the name State of Washington, Douglass Commonwealth, which would conflict with the current state of Washington. To distinguish it from the capital, Washington is sometimes referred to as Washington state, or, in more formal contexts. Residents of Washington and the Pacific Northwest simply refer to the state as Washington, Washington is the northwestern-most state of the contiguous United States. Washington is bordered by Oregon to the south, with the Columbia River forming the western part, to the west of Washington lies the Pacific Ocean. The high mountains of the Cascade Range run north-south, bisecting the state, from the Cascade Mountains westward, Western Washington has a mostly marine west coast climate, with mild temperatures and wet winters, autumns and springs, and relatively dry summers. The Cascade Range contains several volcanoes, which reach altitudes significantly higher than the rest of the mountains, from the north to the south, these major volcanoes are Mount Baker, Glacier Peak, Mount Rainier, Mount St. Helens, and Mount Adams. Mount Rainier, the tallest mountain in the state, is 50 miles south of the city of Seattle and it is also covered with more glacial ice than any other peak in the contiguous 48 states. Western Washington also is home of the Olympic Mountains, far west on the Olympic Peninsula and these deep forests, such as the Hoh Rainforest, are among the only temperate rainforests in the continental United States. Eastern Washington – the part of the state east of the Cascades – has a dry climate. It includes large areas of steppe and a few truly arid deserts lying in the rain shadow of the Cascades. Farther east, the climate becomes less arid, with annual rainfall increasing as one goes east to 21.2 inches in Pullman, the Okanogan Highlands and the rugged Kettle River Range and Selkirk Mountains cover much of the northeastern quadrant of the state
35.
Natural history
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Natural history is the research and study of organisms including animals, fungi and plants in their environment, leaning more towards observational than experimental methods of study. It encompasses scientific research but is not limited to it, with articles nowadays more often published in magazines than in academic journals. Grouped among the sciences, natural history is the systematic study of any category of natural objects or organisms. That is a broad designation in a world filled with many narrowly focused disciplines. For example, geobiology has a strong multi-disciplinary nature combining scientists, a person who studies natural history is known as a naturalist or natural historian. The English term natural history is a translation of the Latin historia naturalis and its meaning has narrowed progressively with time, while the meaning of the related term nature has widened. In antiquity, it covered essentially anything connected with nature or which used materials drawn from nature. For example, Pliny the Elders encyclopedia of this title, published circa 77 to 79 AD, covers astronomy, geography, man and his technology, medicine and superstition as well as animals and plants. Medieval European academics considered knowledge to have two divisions, the humanities and divinity, with science studied largely through texts rather than observation or experiment. In modern terms, natural philosophy roughly corresponded to modern physics and chemistry, natural history had been encouraged by practical motives, such as Linnaeus aspiration to improve the economic condition of Sweden. Similarly, the Industrial Revolution prompted the development of geology to help find useful mineral deposits, the astronomer, William Herschel was also a natural historian. Instead of working with plants or minerals he worked with the stars and he spent his time building telescopes to see the stars and the rest of the time watching the stars. In the beginning, he believed there to be a known as a nebulae. Herschel can be considered a natural historian because he observed the natural world, in the process he made charts of all the stars and kept records of all that he saw. S. Wilcove and T. Eisner, The close observation of organisms—their origins, their evolution, their behavior and it encompasses changes in internal states insofar as they pertain to what organisms do. Some definitions go further, focusing on observation of organisms in their environment. Bartholomew, A student of history, or a naturalist, studies the world by observing plants. A common thread in many definitions of natural history is the inclusion of a component, as seen in a recent definition by H. W. Greene
36.
Taxonomist
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Taxonomy is the science of defining groups of biological organisms on the basis of shared characteristics and giving names to those groups. The exact definition of taxonomy varies from source to source, but the core of the remains, the conception, naming. There is some disagreement as to whether biological nomenclature is considered a part of taxonomy, the broadest meaning of taxonomy is used here. The word taxonomy was introduced in 1813 by Candolle, in his Théorie élémentaire de la botanique, the term alpha taxonomy is primarily used today to refer to the discipline of finding, describing, and naming taxa, particularly species. In earlier literature, the term had a different meaning, referring to morphological taxonomy, ideals can, it may be said, never be completely realized. They have, however, a value of acting as permanent stimulants. Some of us please ourselves by thinking we are now groping in a beta taxonomy, turrill thus explicitly excludes from alpha taxonomy various areas of study that he includes within taxonomy as a whole, such as ecology, physiology, genetics, and cytology. He further excludes phylogenetic reconstruction from alpha taxonomy, thus, Ernst Mayr in 1968 defined beta taxonomy as the classification of ranks higher than species. This activity is what the term denotes, it is also referred to as beta taxonomy. How species should be defined in a group of organisms gives rise to practical and theoretical problems that are referred to as the species problem. The scientific work of deciding how to define species has been called microtaxonomy, by extension, macrotaxonomy is the study of groups at higher taxonomic ranks, from subgenus and above only, than species. While some descriptions of taxonomic history attempt to date taxonomy to ancient civilizations, earlier works were primarily descriptive, and focused on plants that were useful in agriculture or medicine. There are a number of stages in scientific thinking. Early taxonomy was based on criteria, the so-called artificial systems. Later came systems based on a complete consideration of the characteristics of taxa, referred to as natural systems, such as those of de Jussieu, de Candolle and Bentham. The publication of Charles Darwins Origin of Species led to new ways of thinking about classification based on evolutionary relationships and this was the concept of phyletic systems, from 1883 onwards. This approach was typified by those of Eichler and Engler, the advent of molecular genetics and statistical methodology allowed the creation of the modern era of phylogenetic systems based on cladistics, rather than morphology alone. Taxonomy has been called the worlds oldest profession, and naming and classifying our surroundings has likely been taking place as long as mankind has been able to communicate
37.
Type (biology)
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In biology, a type is a particular specimen of an organism to which the scientific name of that organism is formally attached. In other words, a type is an example that serves to anchor or centralize the defining features of that particular taxon, in older usage, a type was a taxon rather than a specimen. Types are of significance to biologists, especially to taxonomists. Types are usually physical specimens that are kept in a museum or herbarium research collection, but failing that, describing species and appointing type specimens is part of scientific nomenclature and alpha taxonomy. If there is more than one named type that all appear to be the same taxon, then the oldest name takes precedence, and is considered to be the correct name of the material in hand. If on the hand the taxon appears never to have been named at all, then the scientist or another qualified expert picks a type specimen and publishes a new name. This process is crucial to the science of biological taxonomy, peoples ideas of how living things should be grouped change and shift over time. How do we know that what we call Canis lupus is the thing, or approximately the same thing. Depending on the nomenclature applied to the organism in question, a type can be a specimen, a culture. Some codes consider a subordinate taxon to be the type, for example, in the research collection of the Natural History Museum in London, there is a bird specimen numbered 1886.6.24.20. This is a specimen of a kind of commonly known as the spotted harrier. This particular specimen is the holotype for that species, the name Circus assimilis refers, by definition and that species was named and described by Jardine and Selby in 1828, and the holotype was placed in the museum collection so that other scientists might refer to it as necessary. Note that at least for type specimens there is no requirement for an individual to be used. The usage of the type is somewhat complicated by slightly different uses in botany. In the PhyloCode, type-based definitions are replaced by phylogenetic definitions, in some older taxonomic works the word type has sometimes been used differently. … Après avoir étudié ces diverses formes, jen arrivai à les considérer comme appartenant à un seul et même type spécifique, translation, This single character permits distinguish this type from all other species of the section. After studying the diverse forms, I came to them as belonging to the one. In botanical nomenclature, a type, is that element to which the name of a taxon is permanently attached, in botany a type is either a specimen or an illustration
38.
Kamchatka Peninsula
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The Kamchatka Peninsula is a 1, 250-kilometre-long peninsula in the Russian Far East, with an area of about 270,000 km2. It lies between the Pacific Ocean to the east and the Sea of Okhotsk to the west, immediately offshore along the Pacific coast of the peninsula runs the 10, 500-metre deep Kuril–Kamchatka Trench. The Kamchatka Peninsula, the Commander Islands, and Karaginsky Island constitute the Kamchatka Krai of the Russian Federation, the vast majority of the 322,079 inhabitants are ethnic Russians, but there are also about 13,000 Koryaks. More than half of the lives in Petropavlovsk-Kamchatsky and nearby Yelizovo. The Kamchatka peninsula contains the volcanoes of Kamchatka, a UNESCO World Heritage Site, Kamchatka receives up to 2,700 mm of precipitation per year. The summers are cool, and the winters tend to be rather stormy though rarely producing lightning. Politically, the forms part of Kamchatka Krai. The southern tip is called Cape Lopatka, the circular bay to the north of this on the Pacific side is Avacha Bay with the capital, Petropavlovsk-Kamchatsky. North up the Pacific side, the four peninsulas are called Shipunsky Point, Kronotsky Point, Kamchatsky Point, North of Ozernoy is the large Karaginsky Bay, which features Karaginsky Island. Northeast of this lies Korfa Bay with the town of Tilichiki, on the opposite side is the Shelikhov Gulf. The Kamchatka or Central Range forms the spine of the peninsula, along the southeast coast runs the Vostochny or Eastern Range. Between these lies the central valley, the Kamchatka River rises northwest of Avacha and flows north down the central valley, turning east near Klyuchi to enter the Pacific south of Kamchatsky Point at Ust-Kamchatsk. In the nineteenth century a trail led west from near Klychi over the mountains to the Tegil river, North of Tegil is Koryak Okrug. South of the Tegil is the Icha River, just south of the headwaters of the Kamchatka, the Bistraya River curves southwest to enter the Sea of Okhotsk at Bolsheretsk, which once served as a port connecting the peninsula to Okhotsk. South of the Bistraya flows the Golygina River, another highway connects local capital with Bolsheretsk. Bus service is available on both roads, most other roads are gravel-covered or coverless ground, requiring off-road capable vehicles. There is semi-regular passenger transportation with aircraft, the obvious circular area in the central valley is the Klyuchevskaya Sopka, an isolated volcanic group southeast of the curve of the Kamchatka River. West of Kronotsky Point is the Kronotsky Biosphere Reserve with the Valley of Geysers, at the southern tip is the Southern Kamchatka Wildlife Refuge with Kurile Lake
39.
Chukotko-Kamchatkan languages
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The Chukotko-Kamchatkan or Chukchi–Kamchatkan languages are a language family of extreme northeastern Siberia. Its speakers traditionally were indigenous hunter-gatherers and reindeer-herders, the Kamchatkan branch is moribund, represented only by Western Itelmen, with only 80 elderly speakers left. The Chukotkan branch had close to 7,000 speakers left, while the family is sometimes grouped typologically and geographically as Paleo-Siberian, no external genetic relationship has been widely accepted as proven. The most popular such proposals have been for links with Eskimo–Aleut, less commonly encountered names for the family are Chukchian, Chukotian, Chukotan, Kamchukchee and Kamchukotic. Of these, Chukchian and Chukotian are ambiguous, since both terms are used to refer specifically to the familys northern branch. In addition, Luorawetlan has been in use since 1775 as a name for the family, although it is properly the self-designation of one of its constituent languages. The derivative Luorawetlanic may be preferable as a name for the family, the Chukotko-Kamchatkan family consists of two distantly related dialect clusters, Chukotkan and Kamchatkan. Chukotkan is considered anywhere from one to four languages, whereas there is one surviving Kamchatkan language. The relationship of the Chukotkan languages to Itelmen is distant, and has only been demonstrated recently. All the Chukotko-Kamchatkan languages are under pressure from Russian, almost all speakers are bilingual in Russian, and most younger members of the ethnic groups associated with the languages speak Russian only. The Chukotko-Kamchatkan languages have no generally accepted relation to any language family. Joseph Greenberg identifies Chukotko-Kamchatkan as a member of Eurasiatic, a proposed macrofamily that includes Indo-European, Altaic, Greenberg also assigns Nivkh and Yukaghir, sometimes classed as Paleosiberian languages, to the Eurasiatic family. While the Eurasiatic hypothesis has been received by Nostraticists and some Indo-Europeanists. Later, he has argued for Nivkh as the closest relative of Chukotko-Kamchatkan, in a 2015 paper, Gerhard Jäger reported intriguing and controversial findings regarding Chukotko-Kamchatkan. On the whole, the similarities between the two families were greater than either shared with any language family. This was the even though Jäger factored in the possibility of language contact.969 to a still statistically-significant 0.964
40.
Hudson's Bay Company
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The Hudsons Bay Company, commonly referred to as The Bay, is a Canadian retail business group. HBCs head office is in the Simpson Tower in Toronto, Ontario, the company is listed on the Toronto Stock Exchange under the symbol HBC. It was once the worlds largest landowner, with the area of the Hudson Bay watershed, known as Ruperts Land, having 15% of North American acreage. From its long-time headquarters at York Factory on Hudson Bay, the company controlled the fur trade throughout much of the English, undertaking early exploration, its traders and trappers forged relationships with many groups of aboriginal peoples. Its network of trading posts formed the nucleus for later official authority in areas of Western Canada. By the mid-19th century, the company evolved into a business selling everything from furs to fine homeware. They quickly introduced a new type of client to the HBC – one that shopped for pleasure and not with skins, in July 2008, HBC was acquired by NRDC Equity Partners, which also owns the upmarket American department store Lord & Taylor. From 2008 to 2012, the HBC was run through a company of NRDC, Hudsons Bay Trading Company. Since 2012, the HBC directly oversees its Canadian subsidiaries Hudsons Bay and Home Outfitters, on 29 July 2013, the HBC announced its takeover of Saks, Inc. operator of the Saks Fifth Avenue brand. The merger was completed on 3 November 2013, in September 2015, HBC acquired the German department store chain Galeria Kaufhof and its Belgian subsidiary from Metro Group for $3.2 billion U. S. In May 2016, HBC announced it would expand to the Netherlands by taking over up to 20 former Vroom & Dreesmann sites by 2017, v&D was an historic Dutch department store chain that went bankrupt and shut down in early 2016. HBC said the expansion would cost CAD $340 million and create 2,500 jobs in the stores, the Dutch stores would operate under the Hudsons Bay and Saks Off Fifth brands. In January 2016, HBC announced it would also expand deeper in the space with its acquisition of online flash sales site. In the 17th century the French had a de facto monopoly on the Canadian fur trade with their colony of New France. Assuming this was Hudson Bay, they sought French backing for a plan to set up a trading post on the Bay, despite this refusal, in 1659 Radisson and Groseilliers set out for the upper Great Lakes basin. A year later returned with premium furs, evidence of the potential of the Hudson Bay region. Subsequently, they were arrested for trading without a licence and fined, determined to establish trade in the Hudson Bay, Radisson and Groseilliers approached a group of businessmen in Boston, Massachusetts to help finance their explorations. The Bostonians agreed on the plans merits but their speculative voyage in 1663 failed when their ship ran into pack ice in Hudson Strait, boston-based English commissioner Colonel George Cartwright learned of the expedition and brought the two to England to raise financing
41.
Fort Vancouver
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Fort Vancouver was a 19th-century fur trading post that was the headquarters of the Hudsons Bay Companys Columbia Department, located in the Pacific Northwest. Named for Captain George Vancouver, the fort was located on the bank of the Columbia River in present-day Vancouver. The fort was a center of the regional fur trading. Every year trade goods and supplies from London arrived either via ships sailing to the Pacific Ocean or overland from Hudson Bay via the York Factory Express, supplies and trade goods were exchanged with a plethora of Indigenous cultures for fur pelts. Furs from Fort Vancouver were often shipped to the Chinese port of Guangzhou where they were traded for Chinese manufactured goods for sale in the United Kingdom, at its pinnacle, Fort Vancouver watched over 34 outposts,24 ports, six ships, and 600 employees. Today, a replica of the fort, with internal buildings, has been constructed and is open to the public as Fort Vancouver National Historic Site. During the War of 1812, the Pacific Northwest was a distant region of the conflict, two rival fur trading outfits, the Canadian North West Company and the American Pacific Fur Company, had until then both operated in the region peaceably. Funded largely by John Jacob Astor, the PFC operated without many opportunities for defense by the United States Navy. News of the war and of a coming British warship put the American company into a difficult position, in October 1813, management met at Fort Astoria and agreed to liquidate its assets to the NWC. The HMS Racoon arrived the following month and in honor of George III of the United Kingdom, Fort Astoria was renamed to Fort George. Frederick Merk has argued the definition used by the negotiators of the Gulf of Georgia included the entirety of the Puget Sound, in addition to the Straits of Georgia and Juan de Fuca. This would have given the United Kingdom the most favorable location for ports north of Alta California, robinson didnt agree to the proposal and subsequent talks didnt focus on establishing a permanent border west of the Rocky Mountains. The Treaty of 1818 made the resources of the vast region were to be free, the treaty wasnt made to combine American and British interests against other colonial powers in the region. Rather, the document states that the joint occupancy of the Pacific Northwest was intended to prevent disputes between the two nations from arising, in the ensuing years, the North West Company would continue to expand its operations in the Pacific Northwest. Skirmishes with its competitor, the Hudsons Bay Company, had already flared into the Pemmican War. The end of the conflict in 1821 saw the NWC mandated by the British Government to merge into the HBC, throughout 1825 and 1826, British officials would continue to offer Americans partition plans for the Pacific Coast of North America. These largely originated in part from correspondence with the NWC and later HBC, the border would continue to extend west on the 49th parallel to the Rocky Mountains, where the Columbia would be used as the border until it reached the Pacific Ocean. Secretary of State for Foreign Affairs George Canning has been appraised by later historians most supportive British Foreign minister in securing a border along the Columbia
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. PMID 18562654. doi:10.1534/genetics.107.084251.
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