The Elymniini, or true browns, are one of the large tribes of the browns subfamily in the brush-footed butterfly family. Sometimes, they are elevated to subfamily status as Elymniinae, they are subdivided into four subtribes. The largest tribe is called Lethina, with Parargina treated as a junior synonym, but this is not correct as it seems: according to cladistic analysis of mtDNA COI and NADH dehydrogenase subunit 1 sequence data, the Parargina – i.e. the group around Pararge, including at least the similar Lasiommata as well as Tatinga and it is the doubtfully distinct relative Chonala – form a distinct lineage after all basal to the distinctly apomorphic Mycalesina and the more plesiomorphic Lethina proper. In any case, these three constitute the bulk of the tribe's diversity; the Elymniina and Zetherina, by contrast, quite form a less diverse clade that diverged early on in the evolution of the tribe. Subtribes are listed in the presumed phylogenetic sequence, from the most ancient to the most advanced.
Some species are listed. Subtribe Elymniina Elymnias Hübner, 1818 – palmflies Elymniopsis Fruhstorfer, 1907 Hyantis Hewitson, 1862 Morphopsis Oberthür, 1880Subtribe Zetherina Callarge Leech, 1892 Ethope Moore, 1866 Ethope himachala – dusky diadem Neorina Westwood, 1850 Neorina patria – white owl Penthema Doubleday, 1848 Zethera Felder, 1861 Subtribe Lethina Chonala Moore, 1893 Enodia Hübner, – pearly-eyes Hanipha Moore, 1880 Kirinia Moore, 1893 Lasiommata Westwood, 1841 – walls and wallbrowns Lethe – treebrowns, woodbrowns and relatives Lopinga Moore, 1893 Mandarinia Leech, 1892 Neope Moore, 1866 Ninguta Moore, 1892 Nosea Koiwaya, 1993 Orinoma Gray, 1846 Pararge Hübner, 1819 Ptychandra C. & R. Felder, 1861 Ptychandra ohtanii Rhaphicera Butler, 1867 Satyrodes Scudder, 1875 Tatinga Moore, 1893 Subtribe Mycalesina Bicyclus Kirby, 1871 Bletogona C. & R. Felder, 1867 Hallelesis Condamin, 1961 Heteropsis Westwood, 1850 Mycalesis – bushbrowns Nirvanopsis Orsotriaena – smooth-eyed bush-brown Pseudomycalesis Tsukada & Nishiyama, 1979 Several prehistoric genera have been described from fossils: Neorinella Martins Neto et al. 1993 Neorinopsis Pseudoneorina Nel & Descimon, 1986 Satyrites Scudder, 1872They are all assumed to belong to the Lethina.
Fossils from near the time of such divergences yield interesting insight into the patterns of evolution. Savela, Markku: Markku Savela's Lepidoptera and Some Other Life Forms: Satyrinae. Version of 3 September 2008. Retrieved 18 March 2009. Yin, Xian-bing. Zoological Research 28: 477-484. PDF fulltext
The bamboos are evergreen perennial flowering plants in the subfamily Bambusoideae of the grass family Poaceae. The word "bamboo" comes from the Kannada term bambu, introduced to English through Indonesian and Malay. In bamboo, as in other grasses, the internodal regions of the stem are hollow and the vascular bundles in the cross-section are scattered throughout the stem instead of in a cylindrical arrangement; the dicotyledonous woody xylem is absent. The absence of secondary growth wood causes the stems of monocots, including the palms and large bamboos, to be columnar rather than tapering. Bamboos include some of the fastest-growing plants in the world, due to a unique rhizome-dependent system. Certain species of bamboo can grow 91 cm within a 24-hour period, at a rate of 4 cm an hour. Giant bamboos are the largest members of the grass family. Bamboos are of notable economic and cultural significance in South Asia, Southeast Asia and East Asia, being used for building materials, as a food source, as a versatile raw product.
Bamboo has a higher specific compressive strength than wood, brick or concrete, a specific tensile strength that rivals steel. Bamboos have long been considered the most primitive grasses because of the presence of bracteate, indeterminate inflorescences, "pseudospikelets", flowers with three lodicules, six stamens, three stigmata. Following more recent molecular phylogenetic research, many tribes and genera of grasses included in the Bambusoideae are now classified in other subfamilies, e.g. the Anomochlooideae, the Puelioideae, the Ehrhartoideae. The subfamily in its current sense belongs to the BOP clade of grasses, where it is sister to the Pooideae; the bamboos comprise three clades classified as tribes, these correspond with geographic divisions representing the New World herbaceous species, tropical woody bamboos, temperate woody bamboos. The woody bamboos do not form a monophyletic group. Altogether, more than 1,400 species are placed in 115 genera. Most bamboo species are native to moist tropical and warm temperate climates.
However, many species are found in diverse climates, ranging from hot tropical regions to cool mountainous regions and highland cloud forests. In the Asia-Pacific region they occur across East Asia, from north to 50 °N latitude in Sakhalin, to south to northern Australia, west to India and the Himalayas. China, Korea and Australia, all have several endemic populations, they occur in small numbers in sub-Saharan Africa, confined to tropical areas, from southern Senegal in the north to southern Mozambique and Madagascar in the south. In the Americas, bamboo has a native range from 47 °S in southern Argentina and the beech forests of central Chile, through the South American tropical rainforests, to the Andes in Ecuador near 4,300 m. Bamboo is native through Central America and Mexico, northward into the Southeastern United States. Canada and continental Europe are not known to have any native species of bamboo; as garden plants, many species grow outside these ranges, including most of Europe and the United States.
Some attempts have been made to grow bamboo on a commercial basis in the Great Lakes region of east-central Africa in Rwanda. In the United States, several companies are growing and distributing species such as Phyllostachys nigra and Phyllostachys edulis; the two general patterns for the growth of bamboo are "clumping" and "running". Clumping bamboo species tend to spread as the growth pattern of the rhizomes is to expand the root mass similar to ornamental grasses. "Running" bamboos, need to be controlled during cultivation because of their potential for aggressive behavior. They spread through their rhizomes, which can spread underground and send up new culms to break through the surface. Running bamboo species are variable in their tendency to spread; some can send out runners of several metres a year, while others can stay in the same general area for long periods. If neglected, over time, they can cause problems by moving into adjacent areas. Bamboos include some of the fastest-growing plants on Earth, with reported growth rates up to 91 cm in 24 hours.
However, the growth rate is dependent on local soil and climatic conditions, as well as species, a more typical growth rate for many cultivated bamboos in temperate climates is in the range of 3–10 cm per day during the growing period. Growing in regions of warmer climates during the late Cretaceous period, vast fields existed in what is now Asia; some of the largest timber bamboo can grow over 30 m tall, be as large as 25–30 cm in diameter. However, the size range for mature bamboo is species-dependent, with the smallest bamboos reaching only several inches high at maturity. A typical height range that would cover many of the common bamboos grown in the United States is 4.5–12 m, depending on species. Anji County of China, known as the "Town of Bamboo", provides the optimal climate and soil conditions to grow and process some of the most valued bamboo poles available worldwide. Unlike all trees, individual bamboo culms emerge from the ground at their full diameter and grow to their full height in a single growing season of three to four months.
During this time, each new shoot grows
The Satyrini are a huge tribe of the Satyrinae butterflies, containing the graylings and allies. They belong to the Nymphalidae family; the classification used here is based on the new work by al.. Subtribe Coenonymphina Coenonympha Heteronympha Lyela Swinhoe, 1908 Sinonympha Lee, 1974 Triphysa Zeller, 1850Subtribe Dirina Dingana van Son, 1955 Dira Hübner, 1819 Tarsocera Butler, 1899 Torynesis Butler, 1899Subtribe Erebiina Erebia Dalman, 1816 Proterebia Roos & Arnschied, 1980 Subtribe Euptychiina About 44 genera, see article for details. Subtribe Hypocystina 19 genera, see article for details. Subtribe Maniolina Aphantopus Wallengren, 1853 Cercyonis Scudder, 1875 Hyponephele Muschamp, 1915 Maniola Schrank, 1801 Pyronia Hübner, 1819Subtribe Melanargiina Melanargia Meigen, 1828Subtribe Parargina Tutt, 1896 Chonala Kirinia Lasiommata Lopinga Nosea Orinoma Pararge Rhaphicera TatingaSubtribe Pronophilina Subtribe Satyrina Setodocis Billberg, 1820 is a nomen dubium
In scientific nomenclature, a synonym is a scientific name that applies to a taxon that goes by a different scientific name, although the term is used somewhat differently in the zoological code of nomenclature. For example, Linnaeus was the first to give a scientific name to the Norway spruce, which he called Pinus abies; this name is no longer in use: it is now a synonym of the current scientific name, Picea abies. Unlike synonyms in other contexts, in taxonomy a synonym is not interchangeable with the name of which it is a synonym. In taxonomy, synonyms have a different status. For any taxon with a particular circumscription and rank, only one scientific name is considered to be the correct one at any given time. A synonym cannot exist in isolation: it is always an alternative to a different scientific name. Given that the correct name of a taxon depends on the taxonomic viewpoint used a name, one taxonomist's synonym may be another taxonomist's correct name. Synonyms may arise whenever the same taxon is named more than once, independently.
They may arise when existing taxa are changed, as when two taxa are joined to become one, a species is moved to a different genus, a variety is moved to a different species, etc. Synonyms come about when the codes of nomenclature change, so that older names are no longer acceptable. To the general user of scientific names, in fields such as agriculture, ecology, general science, etc. A synonym is a name, used as the correct scientific name but, displaced by another scientific name, now regarded as correct, thus Oxford Dictionaries Online defines the term as "a taxonomic name which has the same application as another one, superseded and is no longer valid." In handbooks and general texts, it is useful to have synonyms mentioned as such after the current scientific name, so as to avoid confusion. For example, if the much advertised name change should go through and the scientific name of the fruit fly were changed to Sophophora melanogaster, it would be helpful if any mention of this name was accompanied by "".
Synonyms used in this way may not always meet the strict definitions of the term "synonym" in the formal rules of nomenclature which govern scientific names. Changes of scientific name have two causes: they may be taxonomic or nomenclatural. A name change may be caused by changes in the circumscription, position or rank of a taxon, representing a change in taxonomic, scientific insight. A name change may be due to purely nomenclatural reasons, that is, based on the rules of nomenclature. Speaking in general, name changes for nomenclatural reasons have become less frequent over time as the rules of nomenclature allow for names to be conserved, so as to promote stability of scientific names. In zoological nomenclature, codified in the International Code of Zoological Nomenclature, synonyms are different scientific names of the same taxonomic rank that pertain to that same taxon. For example, a particular species could, over time, have had two or more species-rank names published for it, while the same is applicable at higher ranks such as genera, orders, etc.
In each case, the earliest published name is called the senior synonym, while the name is the junior synonym. In the case where two names for the same taxon have been published the valid name is selected accorded to the principle of the first reviser such that, for example, of the names Strix scandiaca and Strix noctua, both published by Linnaeus in the same work at the same date for the taxon now determined to be the snowy owl, the epithet scandiaca has been selected as the valid name, with noctua becoming the junior synonym. One basic principle of zoological nomenclature is that the earliest published name, the senior synonym, by default takes precedence in naming rights and therefore, unless other restrictions interfere, must be used for the taxon. However, junior synonyms are still important to document, because if the earliest name cannot be used the next available junior synonym must be used for the taxon. For other purposes, if a researcher is interested in consulting or compiling all known information regarding a taxon, some of this may well have been published under names now regarded as outdated and so it is again useful to know a list of historic synonyms which may have been used for a given current taxon name.
Objective synonyms refer to taxa with same rank. This may be species-group taxa of the same rank with the same type specimen, genus-group taxa of the same rank with the same type species or if their type species are themselves objective synonyms, of family-group taxa with the same type genus, etc. In the case of subjective synonyms, there is no such shared type, so the synonymy is open to taxonomic judgement, meaning that th
Fossilworks is a portal which provides query and analysis tools to facilitate access to the Paleobiology Database, a large relational database assembled by hundreds of paleontologists from around the world. Fossilworks is housed at Macquarie University, it includes many analysis and data visualization tools included in the Paleobiology Database. "Fossilworks". Retrieved 2010-04-08
Animals are multicellular eukaryotic organisms that form the biological kingdom Animalia. With few exceptions, animals consume organic material, breathe oxygen, are able to move, can reproduce sexually, grow from a hollow sphere of cells, the blastula, during embryonic development. Over 1.5 million living animal species have been described—of which around 1 million are insects—but it has been estimated there are over 7 million animal species in total. Animals range in length from 8.5 millionths of a metre to 33.6 metres and have complex interactions with each other and their environments, forming intricate food webs. The category includes humans, but in colloquial use the term animal refers only to non-human animals; the study of non-human animals is known as zoology. Most living animal species are in the Bilateria, a clade whose members have a bilaterally symmetric body plan; the Bilateria include the protostomes—in which many groups of invertebrates are found, such as nematodes and molluscs—and the deuterostomes, containing the echinoderms and chordates.
Life forms interpreted. Many modern animal phyla became established in the fossil record as marine species during the Cambrian explosion which began around 542 million years ago. 6,331 groups of genes common to all living animals have been identified. Aristotle divided animals into those with those without. Carl Linnaeus created the first hierarchical biological classification for animals in 1758 with his Systema Naturae, which Jean-Baptiste Lamarck expanded into 14 phyla by 1809. In 1874, Ernst Haeckel divided the animal kingdom into the multicellular Metazoa and the Protozoa, single-celled organisms no longer considered animals. In modern times, the biological classification of animals relies on advanced techniques, such as molecular phylogenetics, which are effective at demonstrating the evolutionary relationships between animal taxa. Humans make use of many other animal species for food, including meat and eggs. Dogs have been used in hunting, while many aquatic animals are hunted for sport.
Non-human animals have appeared in art from the earliest times and are featured in mythology and religion. The word "animal" comes from the Latin animalis, having soul or living being; the biological definition includes all members of the kingdom Animalia. In colloquial usage, as a consequence of anthropocentrism, the term animal is sometimes used nonscientifically to refer only to non-human animals. Animals have several characteristics. Animals are eukaryotic and multicellular, unlike bacteria, which are prokaryotic, unlike protists, which are eukaryotic but unicellular. Unlike plants and algae, which produce their own nutrients animals are heterotrophic, feeding on organic material and digesting it internally. With few exceptions, animals breathe oxygen and respire aerobically. All animals are motile during at least part of their life cycle, but some animals, such as sponges, corals and barnacles become sessile; the blastula is a stage in embryonic development, unique to most animals, allowing cells to be differentiated into specialised tissues and organs.
All animals are composed of cells, surrounded by a characteristic extracellular matrix composed of collagen and elastic glycoproteins. During development, the animal extracellular matrix forms a flexible framework upon which cells can move about and be reorganised, making the formation of complex structures possible; this may be calcified, forming structures such as shells and spicules. In contrast, the cells of other multicellular organisms are held in place by cell walls, so develop by progressive growth. Animal cells uniquely possess the cell junctions called tight junctions, gap junctions, desmosomes. With few exceptions—in particular, the sponges and placozoans—animal bodies are differentiated into tissues; these include muscles, which enable locomotion, nerve tissues, which transmit signals and coordinate the body. There is an internal digestive chamber with either one opening or two openings. Nearly all animals make use of some form of sexual reproduction, they produce haploid gametes by meiosis.
These fuse to form zygotes, which develop via mitosis into a hollow sphere, called a blastula. In sponges, blastula larvae swim to a new location, attach to the seabed, develop into a new sponge. In most other groups, the blastula undergoes more complicated rearrangement, it first invaginates to form a gastrula with a digestive chamber and two separate germ layers, an external ectoderm and an internal endoderm. In most cases, a third germ layer, the mesoderm develops between them; these germ layers differentiate to form tissues and organs. Repeated instances of mating with a close relative during sexual reproduction leads to inbreeding depression within a population due to the increased prevalence of harmful recessive traits. Animals have evolved numerous mechanisms for avoiding close inbreeding. In some species, such as the splendid fairywren, females benefit by mating with multiple males, thus producing more offspring of higher genetic quality; some animals are capable of asexual reproduction, which results
Lepidoptera is an order of insects that includes butterflies and moths. About 180,000 species of the Lepidoptera are described, in 126 families and 46 superfamilies, 10 per cent of the total described species of living organisms, it is one of the most widespread and recognizable insect orders in the world. The Lepidoptera show many variations of the basic body structure that have evolved to gain advantages in lifestyle and distribution. Recent estimates suggest the order may have more species than earlier thought, is among the four most speciose orders, along with the Hymenoptera and Coleoptera. Lepidopteran species are characterized by more than three derived features; the most apparent is the presence of scales that cover the bodies, a proboscis. The scales are modified, flattened "hairs", give butterflies and moths their wide variety of colors and patterns. All species have some form of membranous wings, except for a few that have reduced wings or are wingless. Mating and the laying of eggs are carried out by adults near or on host plants for the larvae.
Like most other insects and moths are holometabolous, meaning they undergo complete metamorphosis. The larvae are called caterpillars, are different from their adult moth or butterfly forms, having a cylindrical body with a well-developed head, mandible mouth parts, three pairs of thoracic legs and from none up to five pairs of prolegs; as they grow, these larvae change in appearance, going through a series of stages called instars. Once matured, the larva develops into a pupa. A few butterflies and many moth species spin a silk case or cocoon prior to pupating, while others do not, instead going underground. A butterfly pupa, called a chrysalis, has a hard skin with no cocoon. Once the pupa has completed its metamorphosis, a sexually mature adult emerges; the Lepidoptera have, over millions of years, evolved a wide range of wing patterns and coloration ranging from drab moths akin to the related order Trichoptera, to the brightly colored and complex-patterned butterflies. Accordingly, this is the most recognized and popular of insect orders with many people involved in the observation, collection, rearing of, commerce in these insects.
A person who collects or studies this order is referred to as a lepidopterist. Butterflies and moths play an important role in the natural ecosystem as pollinators and as food in the food chain. In many species, the female may produce from 200 to 600 eggs, while in others, the number may approach 30,000 eggs in one day; the caterpillars hatching from these eggs can cause damage to large quantities of crops. Many moth and butterfly species are of economic interest by virtue of their role as pollinators, the silk they produce, or as pest species; the term was coined by Linnaeus in 1735 and is derived from Greek λεπίς, gen. λεπίδος and πτερόν. Sometimes, the term Rhopalocera is used for the clade of all butterfly species, derived from the Ancient Greek ῥόπαλον and κέρας meaning "club" and "horn" coming from the shape of the antennae of butterflies; the origins of the common names "butterfly" and "moth" are varied and obscure. The English word butterfly is with many variations in spelling. Other than that, the origin is unknown, although it could be derived from the pale yellow color of many species' wings suggesting the color of butter.
The species of Heterocera are called moths. The origins of the English word moth are more clear, deriving from the Old English moððe" from Common Germanic, its origins are related to Old English maða meaning "maggot" or from the root of "midge", which until the 16th century was used to indicate the larva in reference to devouring clothes. The etymological origins of the word "caterpillar", the larval form of butterflies and moths, are from the early 16th century, from Middle English catirpel, catirpeller an alteration of Old North French catepelose: cate, cat + pelose, hairy; the Lepidoptera are among the most successful groups of insects. They are found on all continents, except Antarctica, inhabit all terrestrial habitats ranging from desert to rainforest, from lowland grasslands to mountain plateaus, but always associated with higher plants angiosperms. Among the most northern dwelling species of butterflies and moths is the Arctic Apollo, found in the Arctic Circle in northeastern Yakutia, at an altitude of 1500 m above sea level.
In the Himalayas, various Apollo species such as Parnassius epaphus have been recorded to occur up to an altitude of 6,000 m above sea level. Some lepidopteran species exhibit symbiotic, phoretic, or parasitic lifestyles, inhabiting the bodies of organisms rather than the environment. Coprophagous pyralid moth species, called sloth moths, such as Bradipodicola hahneli and Cryptoses choloepi, are unusual in that they are found inhabiting the fur of sloths, mammals found in Central and South America. Two species of Tinea moths have been recorded as feeding on horny tissue and have been bred from the horns of cattle; the larva of Zenodochium coccivorella is an internal parasite of the coccid Kermes species. Many species have been recorded as breeding in natural materials or refuse such as owl pellets, bat caves, honeycombs or diseased fruit; as of 2007, there was 174,250 lepi