Smilacaceae, the greenbriers, is a family of flowering plants. While they were often assigned to a more broadly defined family Liliaceae, most recent botanists have accepted the two as distinct families, diverging around 55 million years ago during the Early Paleogene. One characteristic that distinguishes Smilacaceae from most of the other members of the Liliaceae-like Liliales is that it has true vessels in its conducting tissue. Another is that the veins of the leaves, between major veins, are reticulate (net-shaped), rather than parallel as in most monocots.
Smilacaceae
The lily family, Liliaceae, consists of about 15 genera and 610 species of flowering plants within the order Liliales. They are monocotyledonous, perennial, herbaceous, often bulbous geophytes. Plants in this family have evolved with a fair amount of morphological diversity despite genetic similarity. Common characteristics include large flowers with parts arranged in threes: with six colored or patterned petaloid tepals arranged in two whorls, six stamens and a superior ovary. The leaves are linear in shape, with their veins usually arranged parallel to the edges, single and arranged alternating on the stem, or in a rosette at the base. Most species are grown from bulbs, although some have rhizomes. First described in 1789, the lily family became a paraphyletic "catch-all" (wastebasket) group of lilioid monocots that did not fit into other families and included a great number of genera now included in other families and in some cases in other orders. Consequently, many sources and descriptions labelled "Liliaceae" deal with the broader sense of the family.
Liliaceae
Section through flower of Fritillaria meleagris
Lilium flower with perigonium of six undifferentiated tepals, arranged in two trimerous whorls and side-connected (dorsifixed) anthers.
Sego lily (Calochortus nuttallii) with tepals in two clearly distinguished whorls of three sepals and three petals.